Arachnomorpha

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Arachnomorpha
Temporal range: Cambrian - Recent
Haeckel Arachnida.jpg
"Arachnida" from Ernst Haeckel's Kunstformen der Natur, 1904
Scientific classification e
Kingdom: Animalia
Phylum: Arthropoda
Clade: Arachnomorpha
Lameere, 1890
Synonyms
  • Arachnata Paulus, 1979
  • Palaeopoda Packard, 1903

Arachnomorpha is a subdivision or clade of Arthropoda, comprising the monophyletic group formed by the trilobites, other great appendage arthropods and trilobite-like families (Helmetiidae, , Naraoiidae, Liwiidae, and ), and a diverse sister clade including the chelicerates.[1][2] Great debate is held on the position of the Pycnogonida, which are currently thought not to be placed in the immediate vicinity of the Chelicerata.[3] Arachnomorpha are considered the sister group to the crustaceans, which are increasingly being accepted as members of the mandibulate clade (including insects and myriapods).[2]

The arachnomorph concept has been challenged by suggestions that the trilobites fall in the mandibulata stem-group.[4]

There is no consensus as to assigning Arachnomorpha a formal Linnean rank.

A proposal contraposing many synapomorphies unites them to the Trilobita instead.[5] Consideration of the Olenellinae as sister group to the Chelicerata has been refuted.[6]

Classification[]

Arachnomorpha Lameere, 1890 [= Arachnata Paulus, 1979, = Palaeopoda Packard, 1903]

  • Trilobita Walch, 1771
  • Megacheira Hou & Bergstrom, 1997
  • Strabopida Hou & Bergström, 1997
  • Aglaspida Walcott, 1911
  • Cheloniellida Broili, 1932
  • Chelicerata Heymons 1901
    • Family † Legg & Pates, 2017
    • Class Pycnogonida Latreille, 1810
    • Clade Euchelicerata Weygoldt & Paulus, 1979
      • Genus †Offacolidae Sutton et al., 2002
      • Clade Prosomapoda Lamsdell, 2013

Phylogeny[]

Using fossil data, Bergström & Hou (2003) gave an outline of arthropod relationships emphasizing trilobitomorphs (a group that includes trilobites and trilobite-like animals).[7]

Arthropods
Arachnomorpha
Limulavida

Sidneyia

Aglaspidida

Chelicerates

Cheloniellon

Trilobitomorpha (s.s.)
Marrellomorpha

Vachonisia

Mimetaster

Marrella

Emeraldellida

Emeraldella

Retifacies

Core trilobitomorphs

 Order Helmetiida

Helmetia

Kuamaia

Sapenion

Tegopelte

 (e.g. Nektaspida)

Naraoia


 Order 

Cindarella

Xandarella

Phytophilaspis

Trilobites, subclass Trilobita

Hexapoda

Crustaceans

Yohoia

Canadaspis

Fuxianhuia

References[]

  1. ^ Trevor J. Cotton & Simon J. Braddy (2003). "The phylogeny of arachnomorph arthropods and the origin of the Chelicerata". Transactions of the Royal Society of Edinburgh: Earth Sciences. 94 (3): 169–193. doi:10.1017/S0263593300000596.
  2. ^ a b Jonathan R. Hendricks & Bruce S. Lieberman (2008). "New phylogenetic insights into the Cambrian radiation of arachnomorph arthropods". Journal of Paleontology. 82 (3): 585–594. doi:10.1666/07-017.1.
  3. ^ J. A. Dunlop & C. P. Arango (2005). "Pycnogonid affinities: a review" (PDF). Journal of Zoological Systematics and Evolutionary Research. 43 (1): 8–21. doi:10.1111/j.1439-0469.2004.00284.x.
  4. ^ Gerhard Scholtz & Gregory Edgecombe (2005). "Heads, Hox and the phylogenetic position of trilobites". Crustacea and Arthropod Relationships. CRC Press. pp. 139–165. CiteSeerX 10.1.1.130.2585.
  5. ^ Karl-Ernst Lauterbach (1980). "Schlüsselereignisse in der Evolution des Grundplans der Arachnata (Arthropoda)" [Key events in the evolution of the ground plan of the Arachnata (Arthropoda)]. . NF (in German). 23: 163–327.
  6. ^ R. A. Fortey & H. B. Whittington (1989). "The Trilobita as a natural group" (PDF). Historical Biology. 2 (2): 125–138. doi:10.1080/08912968909386496.
  7. ^ Bergström, J. & Hou, X. G. (2003). "Arthropod origins" (PDF). Bulletin of Geosciences. 78 (4): 323–334. Archived from the original (PDF) on 2016-03-04. Retrieved 2014-03-03.


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