Epitonium

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Epitonium
Epitonium scalare shell.jpg
The sculpture of the shell of Epitonium scalare has raised ribs that are known as costae. Costae are an almost universal feature in shells of Epitonium species.
Scientific classification e
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Caenogastropoda
Superfamily: Epitonioidea
Family: Epitoniidae
Genus: Epitonium
Linnaeus, 1758
Type species
Turbo scalaris Linnaeus, 1758
Synonyms

See text

Epitonium is a genus of small predatory sea snails, marine gastropod mollusks. Epitonium is the type genus of the family Epitoniidae, the wentletraps.[1]

The common name wentletraps is derived from the Dutch word wenteltrap, denoting a spiral staircase. This refers to the striking form and sculpture of the shells of the mollusks in this genus, and to a lesser extent, the whole family.

The genus Epitonium has been divided in the past by some authors into several subgenera, but these subgenera were based only on shell characters and did not reflect the true underlying relationships or phylogeny.

Shell description[]

Epitonium shells are high-spired, and are all-white in most of the species within this genus. A few species are tinted with brown to varying degrees, or have a brown stripe on the shoulder of the whorl. The shells typically have blade-like vertical ribs, known as costae.

Habitat[]

Wentletraps usually live in sand near sea anemones or corals.

Life habits[]

These snails are predators and feed by inserting their proboscis and biting out small pieces of the anemone's tissues. Some species of wentletrap feed on only one species of sea anemone, in other words they are species-specific in terms of their prey.

Synonymised genera[]

Many species that now are placed in the genus Epitonium have, over the years, been classified in other genera. These have become synonyms of Epitonium :[1]

  • Aciona Leach, 1815
  • Acutiscala de Boury, 1909
  • Amiciscala Jousseaume, 1912
  • Anguliscala de Boury, 1909
  • Asperiscala de Boury, 1909
  • Avalitiscala de Boury, 1912
  • Cinctiscala de Boury, 1909
  • Cirratiscala de Boury, 1909)
  • Clathrus Oken, 1815
  • Connexiscala de Boury, 1909
  • Crenuliscala Iredale, 1936
  • Crisposcala de Boury, 1886
  • Cyclostoma Lamarck, 1799
  • Decussiscala de Boury, 1909
  • Delicatiscala de Boury, 1909
  • Dulciscala de Boury, 1909
  • Eburniscala de Boury, 1909
  • Epitonium (Asperiscala) de Boury, 1909
  • Epitonium (Clathrus)
  • Epitonium (Epitonium) Röding, 1798 · accepted, alternate representation
  • Epitonium (Hyaloscala) de Boury, 1889 · accepted, alternate representation
  • Epitonium (Papyriscala) de Boury, 1909 accepted, alternate representation
  • Evolutiscala de Boury, 1909
  • Firmiscala de Boury, 1909
  • Folaceiscala de Boury, 1912
  • Foliaceiscala de Boury, 1912
  • Foraceiscala de Boury, 1911
  • Foveoscala de Boury, 1909
  • Fragiliscala Azuma, 1962
  • Fusicoscala Monterosato, 1890
  • Glabriscala de Boury, 1909
  • Graciliscala de Boury, 1909
  • Gradatiscala de Boury, 1909
  • Gyroscala (Fragiliscala) Azuma, 1962
  • Hirtoscala Monterosato, 1890
  • Hyaloscala de Boury, 1890
  • Innesiscala Jousseaume, 1912
  • Janthoscala Mörch, 1875
  • Labeoscala de Boury, 1912
  • Laeviscala de Boury, 1909
  • Lamelliscala de Boury, 1909
  • Lepidiscala de Boury, 1909
  • Limiscala de Boury, 1909
  • Limniscala Dollfus, 1913
  • Linctoscala Monterosato, 1890
  • Lineoscala de Boury, 1909
  • Mazescala Iredale, 1936
  • Melanopsis Férussac, 1807
  • Minutiscala de Boury, 1909
  • Nipponoscala Masahito & Habe, 1973
  • Nitidiscala de Boury, 1909
  • Nitidoscala Strong, 1930
  • Papyriscala de Boury, 1909
  • Parviscala de Boury, 1887
  • Perlucidiscala de Boury, 1912
  • Pudentiscala Iredale, 1938
  • Pupiscala Masahito, Kuroda & Habe, 1971
  • Resticuliscala Cossmann, 1912
  • Reticuliscala de Boury, 1909
  • Scala Mörch, 1852
  • Scala (Innesiscala) de Boury in Jousseaume, 1912
  • Scala (Perlucidiscala) de Boury in Jousseaume, 1912
  • Scalaria Lamarck, 1801
  • Scalaria (Parviscala) de Boury, 1887
  • Scalarus Montfort, 1810 (Unjustified emendation of Scalaria)
  • Scalatarius Duméril, 1806
  • Sodaliscala de Boury, 1909
  • Sphaeroscala Monterosato, 1890
  • Spiniscala de Boury, 1909
  • Turbiniscala de Boury, 1909
  • Viciniscala de Boury, 1909

Species[]

Epitonium hexagonum
Epitonium irregulare

Species within the genus Epitonium include:

Note: Some malacologists have recently placed many of the more than 600 named species into other genera, which were previously considered to be subgenera of the genus Epitonium. On the other hand, many species that belonged to subgenera of Epitonium are now included in Epitonium. These subgenera were based on details of the shell sculpture and not on molecular analysis.

  • (Schepman, 1909)
  • (Verco, 1906)
  • Dall, 1917
  • (Masahito & Habe 1976)
  • (G. B. Sowerby II, 1844)
  • (Jousseaume, 1912)
  • (Thiele, 1925)
  • (Jousseaume, 1912)
  • Kilburn, 1985
  • (G. B. Sowerby II, 1844)
  • (d'Orbigny, 1842) - bladed wentletrap
  • (G. B. Sowerby II, 1844)
  • (Weinkauff, 1866)
  • (Melvill, 1912)
  • (Melvill & Standen, 1903)
  • (Jousseaume, 1894)
  • Kilburn, 1985
  • Nakayama, 2003
  • Kilburn, 1985
  • Cossignani T. & Cossignani V., 1998
  • (Say, 1830) - angulate wentletrap
  • (de Boury, 1913)
  • (Iredale, 1936)
  • (Dall, 1889) - semismooth wentletrap
  • Bonfitto, 2017
  • DuShane, 1979
  • (Jousseaume, 1912)
  • (E. A. Smith, 1890)
  • (Sowerby, J. de C., 1874)
  • (Jousseaume, 1894)
  • (Masahito & Habe, 1973)
  • (G. B. Sowerby II, 1844)
  • (Montrouzier, 1859)
  • (Jousseaume, 1912)
  • (Dall, 1889) - tower wentletrap
  • (Iredale, 1936)
  • (Cotton & Godfrey, 1938)
  • (Carpenter, 1864)
  • Hedley, 1907
  • (E. A. Smith, 1899)
  • Garcia E., 2001
  • Clench & Turner, 1953
  • (Barnard, 1963)
  • (Jousseaume, 1894)
  • Garcia, 2016
  • Kilburn, 1985
  • (Seguenza, 1876)
  • Epitonium bucknilli Powell, 1924
  • (Sowerby II, 1844)
  • (Sowerby, 1844)
  • Dall & Bartsch, 1910
  • (Melvill & Standen, 1903)
  • Dall, 1917
  • Dall, 1919
  • (d’Orbigny, 1842)
  • (Monterosato, 1877)
  • Dall, 1919
  • (Weinkauff, 1866)
  • (Iredale, 1936)
  • (A. Adams, 1873)
  • Dall, 1908
  • Dall, 1917
  • (G. B. Sowerby II, 1844)
  • (Aradas, 1854)
  • (Melvill & Standen, 1903)
  • Clench & Turner, 1952
  • Bozzetti, 2007
  • Lee & Wu, 1998
  • Bozzetti, 2008
  • (Iredale, 1936)
  • (Kanmacher, 1798)
  • Epitonium clathrum Linnaeus, 1758 - common wentletrap
  • Grateloup, J.P.S. de, 1840
  • Kilburn, 1985
  • Kilburn, 1985
  • Dall, 1917
  • (E. A. Smith, 1890)
  • Monterosato, 1877
  • (Yokoyama, 1922)
  • (G. B. Sowerby II, 1844)
  • (Melvill & Standen, 1903)
  • Dall, 1917
  • (Melvill, 1904)
  • (Iredale, 1936)
  • Y.-C. Lee & C.-W. Huang, 2016
  • Epitonium coronatum Lamarck, 1816: synonym of (Lamarck, 1816)
  • (Jousseaume, 1912)
  • (de Boury, 1912)
  • Gittenberger & Gittenberger, 2005
  • (G. B. Sowerby II, 1847)
  • (Hinds, 1843)
  • Carpenter, 1864
  • (Pease, 1863)
  • Kilburn, 1985
  • (de Boury, 1913)
  • (Carpenter, 1856)
  • (G. B. Sowerby II, 1844)
  • (Verrill & Smith, 1880)
  • (Jousseaume, 1912)
  • (Melvill & Standen, 1903)
  • (Sowerby II, 1844) tooth-rib wentletrap
  • (Watson, 1883)
  • Garcia, 2003
  • (G. B. Sowerby II, 1844)
  • Lee Y.C., 2001
  • Hertlein & Strong, 1951
  • (E. A. Smith, 1906)
  • (Melvill & Standen, 1903)
  • (G. B. Sowerby II, 1844)
  • Bozzetti, 2007
  • (Dell, 1990)
  • (Melvill & Standen, 1903)
  • Dall, 1917
  • (Tapparone-Canefri, 1876)
  • (Dall & Simpson, 1901)
  • (Sowerby III, 1903)
  • Dall, 1917
  • Zelaya & Güller, 2017
  • (Masahito, Kuroda & Habe, 1971)
  • (A. Adams & Reeve, 1850)
  • (de Boury, 1913)
  • Pastorino & Penchaszadeh, 1998
  • Kilburn, 1985
  • (Sowerby, 1844)
  • (Jousseaume, 1912)
  • (Strebel, 1908)
  • L. G. Brown, 2019
  • (Mörch, 1852)
  • (Audouin, 1816)
  • (Watson, 1897)
  • (d’Orbigny, 1842) - wrinkle-rib wentletrap
  • Dall, 1927 - humble wentletrap
  • (Hanley, 1840)
  • (G. B. Sowerby II, 1844)
  • (Dall, 1889)
  • (Pease, 1861)
  • (Dautzenberg, 1890)
  • (Kiener, 1839)
  • (Hinds, 1843)
  • (Melvill & Standen, 1901)
  • Cotton, 1938
  • (DuShane, 1988)
  • (Melvill & Standen, 1903)
  • (G. B. Sowerby II, 1844)
  • Nakayama, 2000
  • (Fenaux, 1938)
  • (G. B. Sowerby II, 1844)
  • (Jousseaume, F.P., 1912)
  • Gittenberger & Gittenberger, 2005
  • (Jousseaume, F.P., 1912)
  • (Perry, 1811) - Greenland wentletrap
  • Dall, 1917
  • Preston, 1915
  • DuShane, 1970
  • Makiyama, 1924
  • (Jousseaume, 1912)
  • Kilburn, 1985
  • Gittenberger, 2003
  • (Habe, 1961)
  • (Iredale, 1936)
  • DuShane, 1988
  • (G. B. Sowerby II, 1844)
  • (Carpenter, 1856)
  • (Monterosato, 1874)
  • (Jousseaume, 1912)
  • Epitonium hoeksemai Gittenberger, A., Goud & Gittenberger, E., 2000: synonym of (A. Gittenberger & Goud, 2000)
  • Bozzetti, 2011
  • DuShane & McLean, 1968
  • (Sowerby III, 1901)
  • Epitonium humphreysii (Kiener, 1838) - Humphrey's wentletrap
  • (Sowerby, G.B. II, 1844)
  • (Melvill, 1912)
  • Bozzetti, 2007
  • (Barnard, 1963)
  • (Sowerby II, 1844)
  • (G. B. Sowerby II, 1844)
  • (Thiele, 1925)
  • (Carpenter, 1865)
  • (G. B. Sowerby II, 1844)
  • L. Brown & Weil, 1999
  • Gittenberger, A., Goud & Gittenberger, E., 2000
  • (Jousseaume, 1912)
  • (G. B. Sowerby, 1905)
  • (Sowerby, G.B. II, 1844)
  • (Masahito & Habe, 1976)
  • Segers, Swinnen & De Prins, 2009
  • (Dunker, 1861)
  • (Clessin, 1897)
  • Kilburn, 1985
  • (Monterosato, 1878)
  • (Audouin, J.-V., 1827)
  • (de Boury, 1886)
  • Garcia, 2003
  • Epitonium jukesianum (Forbes, 1852)
  • Garcia, 2003
  • (Yokoyama, 1922)
  • Drivas & Jay, 1989
  • Nakayama, 2000
  • (Clessin, 1897)
  • (Nakayama, 1991)
  • (Nakayama, 1995)
  • (Nyst, 1871)
  • (Mörch, 1875) - Krebs' wentletrap
  • (Jousseaume, 1912)
  • (Jousseaume, 1912)
  • Kilburn, 1985
  • (Melvill & Standen, 1903)
  • (Lamarck, 1822) - lamellose wentletrap
  • Boury, E.A. de, 1911
  • (Sowerby II, 1874)
  • Bozzetti, 2009
  • Garcia, 2011
  • Nakayama, 1995
  • (A. Adams, 1861)
  • (de Boury & Monterosato, 1890)
  • (G. B. Sowerby II, 1844)
  • Gittenberger, A., Goud & Gittenberger, E., 2000
  • (Dall, 1906)
  • (DuShane, 1988)
  • (Sowerby, 1847)
  • DuShane, 1970
  • E. A. Smith, 1910
  • Garcia E., 2003
  • (Thiele, 1925)
  • (Melvill, 1898)
  • * (Jousseaume, 1894)
  • Kilburn, 1985
  • (Jousseaume, 1912)
  • (Sowerby II, 1844)
  • Clench & Turner, 1952
  • (Melvill & Standen, 1903)
  • (E. A. Smith, 1890)
  • (Schepman, 1909)
  • (Pease, 1861)
  • (G. B. Sowerby II, 1844)
  • Epitonium minorum (Iredale, 1936)
  • (Jousseaume, 1912)
  • (de Boury, 1912)
  • Bozzetti, 2008
  • de Boury, 1913
  • (G. B. Sowerby II, 1844)
  • van Aartsen, 1996
  • Bonfitto, 2017
  • (Fenaux, 1943)
  • (Sowerby, 1844)
  • (Say, 1826)
  • Kilburn, 1985
  • (Jeffreys, 1884)
  • (Mörch, 1874)
  • L. G. Brown, 2019
  • (Dall, 1889)
  • Nakayama, 2000
  • Epitonium novangliae (Couthouy, 1838) - New England wentletrap
  • (Sowerby II, 1847)
  • (Sowerby, 1844)
  • (G. B. Sowerby II, 1844)
  • (Nyst, 1871) - fine-ribbed wentletrap
  • (G. B. Sowerby, 1905)
  • (Habe, 1961)
  • Bonfitto & Sabelli, 2001
  • Kilburn, 1985
  • (de Boury, 1921)
  • (A. Adams, 1873)
  • Nakayama, 2000
  • (G. B. Sowerby II, 1844)
  • (Thiele, 1925)
  • (Kiener, 1838)
  • (Masahito, Kuroda & Habe 1971)
  • De Boury
  • (Iredale, 1929)
  • Kilburn, 1985
  • (Melvill, 1912)
  • (Pease, 1867)
  • (de Boury, 1913)
  • (Thiele, 1925)
  • (Jousseaume, 1912)
  • (Jousseaume, 1943)
  • (Sowerby II, 1844)
  • (Watson, 1886)
  • Robertson, 1994
  • Garcia, 2011
  • (Verco, 1906)
  • (Dall, 1889)
  • (Sowerby II, 1844)
  • (Hinds, 1843)
  • Nakayama, 2000
  • (Thiele, 1925)
  • Bouchet & Warén, 1986
  • Kilburn, 1985
  • Kilburn, 1977
  • (Bivona, 1832)
  • (G. B. Sowerby II, 1844)
  • (Masahito, Kuroda & Habe,1971)
  • (Sowerby II, 1844)
  • Bozzetti, 2007
  • (Lamarck, 1822)
  • (Carpenter, 1856)
  • (Carpenter, 1857)
  • Kilburn, 1985
  • Kilburn, 1985
  • Sowerby, 1844
  • Lee & Wu, 1998
  • (Watson, 1897)
  • (Masahito & Habe, 1976)
  • (Melvill & Standen, 1903)
  • (Sowerby, J. de C., 1894)
  • Sowerby, G.B. II, 1844
  • (Kurtz, 1860) - brown-banded wentletrap
  • (Kuroda & Habe, 1961)
  • Kilburn, 1985
  • (E. A. Smith, 1890)
  • (Nyst, 1871)
  • Bozzetti, 2007
  • Jousseaume, 1912
  • Azuma, 1960
  • (Dall, 1903)
  • Epitonium scalare (Linnaeus, 1758)
  • (Melvill, 1910)
  • (Thiele, 1925)
  • (Jeffreys, 1884)
  • (Dall, 1889) - silky wentletrap
  • (Jousseaume, 1912)
  • DuShane & McLean, 1968
  • (Sowerby, 1894)
  • DuShane, 1974
  • (Tryon, 1887)
  • Bonfitto, 2010
  • Kilburn, 1985
  • Bozzetti, 2008
  • (Nyst, 1871)
  • Clessin, 1884
  • Kilburn, 1985
  • (Pilsbry, 1911)
  • (Nyst, 1871)
  • (Monterosato, 1878) - frosted wentletrap
  • (S. M. Souverbie, 1866)
  • (E. A. Smith, 1899)
  • (G. B. Sowerby II, 1844)
  • (Carpenter, 1857)
  • (Melvill & Standen, 1903)
  • (Pease, 1867)
  • Kilburn, 1985
  • Masahito & Habe, 1976
  • Dall, 1917
  • (Iredale, 1936)
  • Lee & Wu, 1998
  • Kilburn, 1985
  • (Sowerby, 1903)
  • Epitonium tenellum (Hutton, 1885)
  • (H. Adams, 1873)
  • Gray, 1827
  • (G. B. Sowerby II, 1844)
  • (Sowerby II, 1874)
  • Nakayama, 2000
  • (Thiele, 1925)
  • DuShane, 1977
  • (Gould, 1847)
  • (Melvill & Standen, 1903)
  • DuShane, 1988
  • (Iredale, 1936)
  • Kilburn, 1985
  • (de Boury, 1890)
  • Clench & Turner, 1952
  • Dall, 1919
  • (Carpenter, 1865) - tinted wentletrap
  • (Kuroda, 1930)
  • Bartsch, 1938
  • (de Boury, 1913)
  • (Melvill & Standen, 1903)
  • (Thompson W., 1840)
  • von Maltzan, 1885
  • (de Boury, 1912)
  • (de Boury, 1913)
  • (Thiele, 1930)
  • (Mörch, 1875)
  • (Turton, 1819)
  • Gittenberger, A., Goud & Gittenberger, E., 2000
  • (Pease, 1869)
  • (Sowerby II, 1844) - one-band wentletrap
  • (Jousseaume, 1912)
  • (Jousseaume, 1912)
  • (Sowerby II, 1844)
  • Fenaux, 1938
  • (Jeffreys, 1884)
  • Bozzetti, 2009
  • Hertlein & Strong, 1951
  • (d'Orbigny, 1840)
  • Strong & Hertlein, 1937
  • Robertson, 1994
  • (Melvill & Standen, 1903)
  • Yokoyama, 1922
  • L.G. Brown, 2010
  • Bonfitto, 2017
  • Suzuki & Ichikawa, 1936
  • Bonfitto, 2017
  • (Melvill, 1910)
  • Dall, 1917
Subgenera and species brought into synonymy
  • Epitonium (Asperiscala)'' de Boury, 1909: synonym of Epitonium Röding, 1798
  • Epitonium (Boreoscala) Kobelt, 1902: synonym of Kobelt, 1902
  • Epitonium (Clathrus) : synonym of Epitonium Röding, 1798
  • Epitonium (Kiiscala) Nakayama, 1995 : synonym of Cycloscala Dall, 1889
  • Epitonium delicatulum (Adams H., 1869): synonym of (H. Adams, 1873)

See also[]

  • Wentletrap, an article about the family Epitoniidae.

References[]

  1. ^ a b Bouchet, P.; Gofas, S. (2010). Epitonium Röding, 1798. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=137943 on 2010-11-22
  • Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca. in: Costello, M.J. et al. (eds), European Register of Marine Species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Patrimoines Naturels. 50: 180-213.
  • "Epitonium". Integrated Taxonomic Information System.
  • Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1

External links[]

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