Equilabium

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Equilabium
Scientific classification e
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Lamiales
Family: Lamiaceae
Tribe:
Genus: Equilabium
Mwany., A.J.Paton & Culham[1][2]

Equilabium is a genus of flowering plants in the family Lamiaceae. It was split off from the genus Plectranthus in 2018 as the result of a molecular phylogenetic study. Most species are native to Africa, with two found in the Indian subcontinent.[2]

Description[]

Species of Equilabium are herbaceous or soft-wooded shrubs, rarely woody shrubs. The herbaceous species may be annual or perennial. The leaves are opposite. The inflorescences are "thryses" – compound structures in which the flowers are arranged on secondary branches. Individual flowers have stalks (pedicels). The sepals form a two-lipped funnel shape, the upper lip having four lobes, the lower lip one lobe. The petals form a two-lipped tube, with an S-shaped basal portion. There are four stamens, whose filaments are not fused together. The style is divided into two parts (bifid). The nutlets are ovoid.[2]

Equilabium and Plectranthus species are distinguished from Coleus by having the stem (pedicel) of the calyx attached symmetrically to the base of the calyx tube, rather than opposite the upper lip, and having the corolla lobes more or less equal in length. Equilabium species can be distinguished from Plectranthus by the truncated shape of the throat of the calyx and by the usually S-shaped tube of the corolla, which is parallel-sided at the base.[2]

Taxonomy[]

The genus Equilabium was first described in 2018 as the result of a molecular phylogenetic study. Equilabium was split off from the genus Plectranthus, which was discovered not to be monophyletic. Only the names of the genus and the type species, , were formally published at the time.[3] The names of 41 more species were published in 2019.[2]

Phylogeny[]

In 2019, Paton et al. published a summary cladogram for the subtribe Plectranthinae,[2] based on an earlier 2018 study.[3] The new genus Equilabium was established and Coleus revived.[2] In the version below, the three genera accepted in these studies that were formerly included in a broad circumscription of Plectranthus are highlighted.

Tribe Ocimeae
 

Subtribe Ociminae

 
 
Subtribe Plectranthinae
 
 

Alvesia (3 spp.)

 
 
 

Aeollanthus (42 spp.)

 
 
 
 

Tetradenia + Thorncroftia (29 spp.)

 
 

Plectranthus (72 spp.)

 
 
 

Capitanopsis (6 spp.)

 
 

Equilabium (42 spp.)

 
 
 
 
 

Coleus (294 spp.)

 
 
 
 

Species[]

Paton et al. (2019) list 42 species:[2][4]

  • (Brummitt & Seyani) Mwany., Culham & A.J.Paton (syn. Plectranthus acaulis) – Malawi to Zambia (Nyika Plateau)
  • (Lukhoba & A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus agnewii) – E. Tropical Africa
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus annuus) – S. Tanzania to Mozambique
  • (Lukhoba & A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus caespitosus) – Kenya to N. Tanzania
  • (Launert) Mwany. & A.J.Paton (syn. Plectranthus candelabriformis) – Tanzania to N. Namibia
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus cinereus) – Kenya to Tanzania
  • (Brenan) Mwany. & A.J.Paton (syn. Plectranthus dissectus) – S. Malawi
  • (Codd) Mwany. & A.J.Paton (syn. Plectranthus dolomiticus) – Zimbabwe to Limpopo
  • (E.A.Bruce) Mwany. & A.J.Paton (syn. Coleus equisetiformis) – Tanzania to N. Zambia
  • (Vatke) Mwany. & A.J.Paton (syns Coleus flaccidus, Plectranthus flaccidus) – S. Somalia to Mozambique, Comoros
  • (Hook. f.) Mwany. & A.J.Paton (syn. Plectranthus glandulosus) – Widespread in Tropical Africa
  • (Gürke) Mwany. & A.J.Paton (syn. Plectranthus goetzei) – SW. Tanzania to Zambia
  • (Suess.) Mwany. & A.J.Paton (syn. Plectranthus gracilis) – S. Tanzania to S. Tropical Africa
  • (Briq.) Mwany. & A.J.Paton (syn. Plectranthus intrusus) – Democratic Republic of the Congo
  • (Lebrun & L. Touss) Mwany. & A.J.Paton (syn. Plectranthus janthinothryx) – Democratic Republic of the Congo and Uganda
  • (Wickens & B.Mathew) A.J.Paton (syn. Plectranthus jebel-marrae) – Sudan
  • (Gürke) Mwany. & A.J.Paton (syn. Plectranthus kamerunensis) – Nigeria to Cameroon, E. Tropical Africa
  • (Benth.) Mwany. & A.J.Paton (syn. Plectranthus laxiflorus) – Ethiopia to South Africa
  • (Baker) Mwany. & A.J.Paton (syn. Plectranthus longipes) – Eritrea to Rwanda and Tanzania
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus mafiensis) – Tanzania (Mafia I.)
  • (Baker) Mwany. & A.J.Paton (syn. Plectranthus masukensis) – Kenya to N. Zambia
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus megafolius) – Tanzania (type)
  • (Aiton) Mwany. & A.J.Paton (syn. Ocimum molle, Plectranthus mollis) – Indian Subcontinent to N. Myanmar
  • (Gürke) Mwany. & A.J.Paton (syn. Plectranthus orbicularis) – E. Tanzania (incl. Zanzibar, Pemba)
  • (Oliv.) Mwany. & A.J.Paton (syn. Plectranthus parvus) – Uganda to N. Zambia
  • (Baker) Mwany. & A.J.Paton (syn. Plectranthus pauciflorus) – Democratic Republic of the Congo, East Africa, Zambia
  • (Benth.) Mwany. & A.J.Paton (syn. Plectranthus petiolaris) – S. Mozambique to South Africa
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus pinetorum) – Malawi and E. Zimbabwe
  • (Baker) Mwany. & A.J.Paton (syn. Plectranthus pubescens) – SW. Tanzania to Mozambique
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus pulcherissimus) – Democratic Republic of the Congo and Zambia
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus radiatus) – S. Tanzania
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus rungwensis) – Tanzania (Mt. Rungwe)
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus scopulicola) – Tanzania (W. Usambara Mts.)
  • (N.E.Br.) Mwany. & A.J.Paton (syn. Plectranthus selukwensis) – Zambia to Zimbabwe
  • (A.J.Paton, Friis & Sebsebe) A.J.Paton (syn. Plectranthus spananthus) – Ethiopia
  • (Baker) Mwany. & A.J.Paton (syn. Plectranthus stenophyllus) – S. Tanzania to S. Tropical Africa
  • (Baker) Mwany. & A.J.Paton (syn. Plectranthus stenosiphon) – S. Malawi to C. Mozambique and Zimbabwe
  • (Gilli) Mwany. & A.J.Paton (syn. Plectranthus stolzii) – SW. Tanzania to N. Malawi
  • (Benth.) Mwany., Smitha & A.J.Paton (syn. Plectranthus subincisus) – S. India, Sri Lanka?
  • (A.J.Paton) Mwany. & A.J.Paton (syn. Plectranthus vesicularis) – Tanzania to N. Mozambique
  • (Brummitt & Seyani) Mwany., Culham & A.J.Paton (syn. Plectranthus viphyensis) – Tanzania to S. Tropical Africa
  • (S.Moore) Mwany. & A.J. Paton (syn. Plectranthus wollastonii) – E. Central Tropical Africa to S. Kenya.

References[]

  1. ^ "Equilabium Mwany., A.J.Paton & Culham", The International Plant Names Index, retrieved 2020-07-29
  2. ^ a b c d e f g h Paton, Alan J.; Mwanyambo, Montfort; Govaerts, Rafaël H.A.; Smitha, Kokkaraniyil; Suddee, Somran; Phillipson, Peter B.; Wilson, Trevor C.; Forster, Paul I. & Culham, Alastair (2019), "Nomenclatural changes in Coleus and Plectranthus (Lamiaceae): a tale of more than two genera", PhytoKeys (129): 1–158, doi:10.3897/phytokeys.129.34988, PMC 6717120, PMID 31523157
  3. ^ a b Paton, A.; Mwanyambo, M. & Culham, A. (2018). "Phylogenetic study of Plectranthus, Coleus and allies (Lamiaceae): Taxonomy, distribution and medicinal use" (PDF). Botanical Journal of the Linnean Society. 188 (4): 355–376. doi:10.1093/botlinnean/boy064.
  4. ^ "Equilabium Mwany., A.J. Paton & Culham". Tropicos. Missouri Botanical Garden. Retrieved 2020-07-29.
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