List of ant genera

Ants (family Formicidae in the order Hymenoptera) are the most species-rich of all social insects, with more than 12,000 described species and many others awaiting description.^[1] Formicidae is divided into 21 subfamilies, of which 17 are extant and four subfamilies are extinct, described from fossils. In total more than 300 genera have been described.^[2] Ants have come to occupy virtually all major terrestrial habitats, with the exception of tundra and cold ever-wet forests. They display a wide range of social behaviors, foraging habits and associations with other organisms, which has generated scientific and public interest.^[1]

The following is a list of worldwide ant genera organised by subfamily.

Key[]

Genus name	The scientific name of the genus.
Binomial authority	The binomial authority—the person who first described the species using an available scientific name.
Classified	The year the genus was formally described and classified.
No. of species	The number of species, both extant and extinct, that are members of the genus.
Type species	The type species that was designated by the original author, unless noted otherwise. Extinct taxa are indicated by a †.
Example image	An image (if available) of a species who is a member of the genus.
References	Most citations provided are the first publications that describe the genus; additional citations are added if the original source does not contain all provided information.

Subfamilies[]

Agroecomyrmecinae[]

The subfamily Agroecomyrmecinae represents two extant genera and two fossil genera, once widespread in both hemispheres during the early Tertiary.^[3][4] The subfamily was originally classified as Agroecomyrmecini, a Myrmicinae tribe until English myrmecologist Barry Bolton raised the tribe to subfamily status in 2003.^[5][6] Most specimens collected are from Central America and Mexico, although one type specimen of an Agroecomyrmecinae species was collected from Ghana.^[7]

Agroecomyrmecinae Carpenter, 1930
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Agroecomyrmex	Wheeler	1910	1	†Myrmica duisburgi	†Agroecomyrmex duisburgi	[8][9]
Ankylomyrma	Bolton	1973	1	Ankylomyrma coronacantha	Ankylomyrma coronacantha	[10]
†Eulithomyrmex	Carpenter	1935	2	†Lithomyrmex rugosus	†Eulithomyrmex rugosus	[11]
Tatuidris	Brown & Kempf	1968	1	Tatuidris tatusia	Tatuidris tatusia	[4]

Amblyoponinae[]

The subfamily Amblyoponinae represents nine extant genera and one fossil genus. Established by Swiss myrmecologist Auguste Forel in 1893, these ants are specialist predators, distributed worldwide in the tropics.^[12][13][14]

Subfamily Amblyoponinae Forel, 1893 – 10 genera, 146 species[15]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Adetomyrma	Ward	1994	9	Adetomyrma venatrix	Adetomyrma venatrix	[16]
Amblyopone	Erichson	1842	10	Amblyopone australis	Amblyopone australis	[17]
†Casaleia	Pagliano & Scaramozzino	1990	4	†Protamblyopone inversa	†Casaleia eocenica	[18]
Fulakora	Mann	1919	25		Fulakora celata	[19][20]
Myopopone	Roger	1861	2	Myopopone maculata	Myopopone castanea	[21][22]
Mystrium	Roger	1862	14		Mystrium mysticum	[23]
Onychomyrmex	Emery	1895	4	Onychomyrmex hedleyi	Onychomyrmex hedleyi	[24]
Prionopelta	Mayr	1866	22		Prionopelta punctulata	[25]
Stigmatomma	Roger	1859	54		Stigmatomma denticulatum	[26]
Xymmer	Santschi	1914	1	Stigmatomma muticum	Xymmer muticus	[27]

Aneuretinae[]

The subfamily Aneuretinae represents one extant genus and eight fossil genera established by Italian entomologist Carlo Emery in 1913. Only a single species of this subfamily is extant, the Sri Lankan relict ant (Aneuretus simoni), endemic to Sri Lanka.^[28]

Subfamily Aneuretinae Emery, 1913 – nine genera, 12 species[29]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Aneuretellus	Dlussky	1988	1	†Aneuretellus deformis	N/A	[30]
Aneuretus	Emery	1893	1	Aneuretus simoni	Aneuretus simoni	[31]
†	Dlussky & Perfilieva	2014	1	†	N/A	[32]
†Burmomyrma	Dlussky	1996	1	†Burmomyrma rossi	†Burmomyrma rossi	[33]
†Cananeuretus	Engel & Grimaldi	2005	1	†Cananeuretus occidentalis	N/A	[34]
†	Carpenter	1930	2	†	†Mianeuretus mirabilis	[5]
†Paraneuretus	Wheeler	1915	3	†Paraneuretus tornquisti	†Paraneuretus tornquisti	[9]
†	Wheeler	1915	1	†	†Pityomyrmex tornquisti	[9]
†	Wheeler	1915	1	†	†Protaneuretus succineus	[9]

Apomyrminae[]

The subfamily Apomyrminae contains the single genus Apomyrma which only has one species, the rare subterranean ant Apomyrma stygia from West Africa. Several undescribed species are known to exist, all from tropical Africa.^[35]

Subfamily Apomyrminae Dlussky & Fedoseeva, 1988 – one genus, one species[36]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Apomyrma	Brown, Gotwald & Levieux	1970	1	Apomyrma stygia	Apomyrma stygia	[37]

Brownimeciinae[]

The subfamily Brownimeciinae contains the single genus Brownimecia which only has one species, Brownimecia clavata. It was described in 1997 after a fossilised specimen was collected from Cretaceous amber from New Jersey, and was initially placed in the subfamily Ponerinae.^[38] The species was later classified into its own subfamily in 2003 by Barry Bolton.^[6]

Subfamily †Brownimeciinae Bolton, 2003 – one genus, one species[39]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Brownimecia	Grimaldi, Agosti & Carpenter	1997	1	†Brownimecia clavata	†Brownimecia clavata	[38]

Dolichoderinae[]

The subfamily Dolichoderinae was established by Forel in 1878, which represents 28 extant genera and 20 fossil genera. The subfamily presents a great diversity of species throughout the world, mainly in the tropics. Most species are generalised scavengers, but some are predacious.^[40][41]

Subfamily Dolichoderinae Forel, 1878 – 48 genera, 843 species[42]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Alloiomma	Zhang	1989	2	†Alloiomma changweiensis	N/A	[43]
Anillidris	Santschi	1936	1	Anillidris bruchi	Anillidris bruchi	[44]
Anonychomyrma	Donisthorpe	1947	26	Anonychomyrma myrmex	Anonychomyrma gilberti	[45]
Aptinoma	Fisher	2009	2	Aptinoma mangabe	Aptinoma mangabe	[46]
Arnoldius	Dubovikoff	2005	3	Bothriomyrmex flavus	Arnoldius flavus	[47]
†Asymphylomyrmex	Wheeler	1915	1	†Asymphylomyrmex balticus	†Asymphylomyrmex balticus	[9]
Axinidris	Weber	1941	21	Axinidris acholli	Axinidris acholli	[48]
Azteca	Forel	1878	86		Azteca xanthochroum	[49]
Bothriomyrmex	Emery	1869	24	Bothriomyrmex costae	Bothriomyrmex costae	[50]
†Chronomyrmex	McKellar, Glasier & Engel	2013	1	†Chronomyrmex medicinehatensis	†Chronomyrmex medicinehatensis	[51]
Chronoxenus	Santschi	1919	6	Bothriomyrmex myops	Chronoxenus myops	[52]
†Ctenobethylus	Brues	1939	1	†Ctenobethylus succinalis	†Ctenobethylus goepperti	[53]
Doleromyrma	Forel	1907	2	Tapinoma darwinianum	Doleromyrma darwiniana	[54]
Dolichoderus	Lund	1831	181	Formica attelaboides	Dolichoderus attelaboides	[55]
Dorymyrmex	Mayr	1866	60	Dorymyrmex flavescens	Dorymyrmex flavescens	[25]
Ecphorella	Forel	1909	1	Tapinoma wellmani	Ecphorella wellmani	[56]
†Elaeomyrmex	Carpenter	1930	2	†Elaeomyrmex gracilis	†Elaeomyrmex gracilis	[5]
†Elaphrodites	Zhang	1989	2	†Elaphrodites scutulatus	N/A	[43]
†Eldermyrmex	Heterick & Shattuck	2011	1	†Iridomyrmex oblongiceps	†Eldermyrmex oblongiceps	[57]
†Emplastus	Donisthorpe	1920	12	†Emplastus emeryi	†Emplastus britannicus	[58]
†Eotapinoma	Dlussky	1988	3	†Eotapinoma gracilis	N/A	[30]
†Eurymyrmex	Zhang, Sun & Zhang	1994	1	†Eurymyrmex geologicus	N/A	[59]
Forelius	Emery	1888	18	Iridomyrmex mccooki	Forelius mccooki	[60]
Froggattella	Forel	1902	2	Acantholepis kirbii	Froggattella kirbii	[61]
Gracilidris	Wild & Cuezzo	2006	2	Gracilidris pombero	Gracilidris pombero	[62]
Iridomyrmex	Mayr	1862	84	Formica detecta	Iridomyrmex purpureus	[22][63]
†Kotshkorkia	Dlussky	1981	1	†Kotshkorkia laticeps	N/A	[64]
†	Lapolla & Greenwalt	2015	1	†	†Ktunaxia jucunda	[65]
Leptomyrmex	Mayr	1862	29	Formica erythrocephala	Leptomyrmex erythrocephalus	[63]
†Leptomyrmula	Emery	1913	1	†Leptomyrmula maravignae	†Leptomyrmula maravignae	[66]
Linepithema	Mayr	1866	20	Linepithema fuscum	Linepithema fuscum	[25]
Liometopum	Mayr	1861	28	Liometopum microcephalum	Liometopum microcephalum	[67]
Loweriella	Shattuck	1992	1	Loweriella boltoni	Loweriella boltoni	[68]
†Miomyrmex	Carpenter	1930	2	†Miomyrmex impactus	†Miomyrmex impactus	[5]
Nebothriomyrmex	Dubovikov	2004	1	Nebothriomyrmex majeri	Nebothriomyrmex majeri	[69]
Ochetellus	Shattuck	1992	7	Ochetellus glaber	Ochetellus glaber	[70]
Papyrius	Shattuck	1992	2	Papyrius nitidus	Papyrius nitidus	[70]
†Petraeomyrmex	Carpenter	1930	1	†Petraeomyrmex minimus	†Petraeomyrmex minimus	[5]
Philidris	Shattuck	1992	9	Philidris cordata	Philidris cordata	[70]
†Proiridomyrmex	Dlussky & Rasnitsyn	2003	2	†Proiridomyrmex vetulus	†Proiridomyrmex rotundatus	[65]
†Protazteca	Carpenter	1930	5	†Protazteca elongata	†Protazteca elongata	[5]
Ravavy	Fisher	2009	1	Ravavy miafina	Ravavy miafina	[46]
Tapinoma	Förster	1850	74	Tapinoma erraticum	Tapinoma erraticum	[71]
Technomyrmex	Mayr	1872	97		Technomyrmex strenuus	[72]
Turneria	Forel	1895	8	Turneria bidentata	Turneria bidentata	[73]
†Usomyrma	Dlussky et al.	2014	1	†Usomyrma mirabilis	†Usomyrma mirabilis	[74]
†Yantaromyrmex	Dlussky & Dubovikoff	2013	5	†Yantaromyrmex geinitzi	†Yantaromyrmex geinitzi	[75]
†Zherichinius	Dlussky	1988	2	†Zherichinius horribilis	†Zherichinius horribilis	[30]

Dorylinae[]

The subfamily Dorylinae was established by Leach in 1815, which represents 27 extant genera and one fossil genus. Many species of ant in this subfamily are known as army ants that are distributed in the Old World and New World.^[76][77]

Subfamily Dorylinae Leach, 1815 – 28 genera, 693 species[77][78]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acanthostichus	Mayr	1887	24		Acanthostichus serratulus	[79]
Aenictogiton	Emery	1901	7		Aenictogiton fossiceps	[80]
Aenictus	Shuckard	1840	184		Aenictus ambiguus	[81]
Cerapachys	Smith	1857	5		Cerapachys antennatus	[22][82]
Cheliomyrmex	Mayr	1870	4		Cheliomyrmex morosus	[83]
Chrysapace	Crawley	1924	3[a]		Chrysapace jacobsonii	[84]
Cylindromyrmex	Mayr	1870	13		Cylindromyrmex striatus	[83]
Dorylus	Fabricius	1793	60		Dorylus helvolus	[85]
Eburopone	Borowiec	2016	1		Eburopone wroughtoni	[77]
Eciton	Latreille	1804	12	Eciton hamatum	Eciton hamatum	[86][87]
Eusphinctus	Emery	1893	2		Eusphinctus furcatus	[31]
Labidus	Jurine	1807	7	Labidus coecus	Labidus coecus	[88]
Leptanilloides	Mann	1923	19		Leptanilloides biconstricta	[89]
Lioponera	Mayr	1879	73		Lioponera longitarsus	[90]
Lividopone	Bolton & Fisher	2016	1		Lividopone livida	[91]
Neivamyrmex	Borgmeier	1940	128	Neivamyrmex nigrescens	Neivamyrmex nigrescens	[92]
Neocerapachys	Borowiec	2016	2		Neocerapachys neotropicus	[77]
Nomamyrmex	Borgmeier	1936	2		Nomamyrmex esenbeckii	[93]
Ooceraea	Roger	1862	11	Ooceraea fragosa	Ooceraea fragosa	[23]
Parasyscia	Emery, in André	1882	50		Parasyscia piochardi	[94]
†Procerapachys	Wheeler	1915	3	†	†Procerapachys annosus	[9]
Simopone	Forel	1891	39		Simopone grandidieri	[95]
Sphinctomyrmex	Mayr	1866	3	Sphinctomyrmex stali	Sphinctomyrmex stali	[96]
Syscia	Roger	1861	5	Syscia typhla	Syscia typhla	[21]
Tanipone	Bolton & Fisher	2012	10		Tanipone hirsuta	[97]
Vicinopone	Bolton & Fisher	2012	1	Vicinopone conciliatrix	Vicinopone conciliatrix	[97]
Yunodorylus	Xu	2000	4		Yunodorylus sexspinus	[98]
Zasphinctus	Wheeler	1918	20		Zasphinctus turneri	[99]

Ectatomminae[]

The subfamily Ectatomminae represents four extant genera and three fossil genera, established in 1895 by Carlo Emery. They are distributed in tropical and warm climates in the New World and Old World, as well as the Indo-Australian regions.^[41]

Subfamily Ectatomminae Emery, 1895 – seven genera, 278 species[100]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†	Dlussky	1999	1	†	N/A	[101]
Ectatomma	Smith	1858	16	Ectatomma tuberculatum	Ectatomma tuberculatum	[22][102]
†	Wheeler	1915	1	†	†Electroponera dubia	[9]
Gnamptogenys	Roger	1863	143		Gnamptogenys sulcata	[103][104]
†Pseudectatomma	Dlussky & Wedman	2012	2	†Pseudectatomma eocenica	†Pseudectatomma eocenica	[13]
Rhytidoponera	Mayr	1862	108		Rhytidoponera araneoides	[63][104]
Typhlomyrmex	Mayr	1862	7		Scyphodon anomalum	[63]

Formiciinae[]

The subfamily Formiciinae represents one extinct genus of ants dating back to the Eocene.^[b] Ants of the genus Titanomyrma are the largest ants ever known, with queen specimens the size of small hummingbirds. Fossils have been collected from the state of Wyoming and in Germany.^[106][107]

Subfamily †Formiciinae Lutz, 1986 – one, six species[108]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Formicium[b]	Westwood	1854	3	None[c]	N/A	[109]
†Titanomyrma	Archibald, Johnson, Mathewes & Greenwood	2011	3	†Titanomyrma gigantea	†Titanomyrma gigantea	[106]

Formicinae[]

The subfamily Formicinae represents 51 extant genera and 30 fossil genera that are globally distributed. Established by French zoologist Pierre André Latreille in 1809, the subfamily has more than 3,000 described species, placing it as the second largest ant subfamily. Despite this, the hyperdiverse genus Camponotus is the most diverse group of ants in the world, with more than 1,100 species described.^[41][110]

Subfamily Formicinae Latreille, 1809 – 82 genera, 3,243 species[111]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acropyga	Roger	1862	41	Acropyga acutiventris	Acropyga acutiventris	[23]
Agraulomyrmex	Prins	1983	2		Agraulomyrmex meridionalis	[112]
Alloformica	Dlussky	1969	4		Alloformica aberrans	[113]
Anoplolepis	Santschi	1914	9	Anoplolepis gracilipes	Anoplolepis gracilipes	[114]
Aphomomyrmex	Emery	1899	1	Aphomomyrmex afer	Aphomomyrmex afer	[115]
†Attopsis	Heer	1850	1	†Attopsis longipennis	Attopsis longipennis	[116]
Bajcaridris	Agosti	1994	3		Bajcaridris theryi	[117]
Brachymyrmex	Mayr	1868	44	Brachymyrmex patagonicus	Brachymyrmex patagonicus	[118]
Bregmatomyrma	Wheeler	1929	1	Bregmatomyrma carnosa	Bregmatomyrma carnosa	[119]
Calomyrmex	Emery	1895	9		Calomyrmex laevissimus	[120]
†	Steinbach	1967	5	†	†Camponotites silvestris	[121]
Camponotus	Mayr	1861	1131	Camponotus ligniperda	Camponotus ligniperda	[22][67]
Cataglyphis	Förster	1850	91	Cataglyphis bicolor	Cataglyphis bicolor	[122]
†	Dlussky	2008	2	†	†Cataglyphoides constrictus	[123]
†Chimaeromyrma	Dlussky	1988	1	†	N/A	[30]
Cladomyrma	Wheeler	1920	13		Cladomyrma hewitti	[124]
†	Dlussky	2008	1	†	N/A	[123]
†	Hong	2002	1	†	N/A	[125]
Dinomyrmex	Ashmead	1905	1	Formica gigas	N/A	[126]
†	Wheeler	1915	2	†	†Drymomyrmex fuscipennis	[9]
Echinopla	Smith	1857	33		Echinopla melanarctos	[82][127]
†	Hong	2002	2	†	N/A	[125]
Euprenolepis	Emery	1906	8	Euprenolepis procera	Euprenolepis procera	[128]
†	Hong	2002	1	†	N/A	[125]
Formica	Linnaeus	1758	234	Formica rufa	Formica rufa	[129]
†	Hong	2002	1	†	N/A	[125]
Gesomyrmex	Mayr	1868	17	†	†Gesomyrmex hoernesi	[130]
Gigantiops	Roger	1863	1	Gigantiops destructor	Gigantiops destructor	[131]
†	Wheeler	1915	1	†	†Glaphyromyrmex oligocenicus	[9]
†	Dlussky & Putyatina	2014	1	†	N/A	[132]
†	Hong	2002	1	†	N/A	[125]
Iberoformica	Tinaut	1990	2	Iberoformica subrufa	Iberoformica subrufa	[133]
†	Heer	1850	2	†	N/A	[116]
†	Grimaldi & Agosti	2000	1	†	†Kyromyrma neffi	[134]
Lasiophanes	Emery	1895	6		Lasiophanes atriventris	[135]
Lasius	Fabricius	1804	134	†	†Formica nigra	[136]
Lepisiota	Santschi	1926	81		Lepisiota rothneyi	[137]
†	Hong	2002	1	†	N/A	[125]
†	Donisthorpe	1920	2	†	†Leucotaphus gurnetensiss	[58]
†	Hong	2002	1	†	N/A	[125]
†	Hong	2002	2	†	N/A	[125]
†	Hong	2002	1	†	N/A	[125]
Melophorus	Lubbock	1883	23	Melophorus bagoti	Melophorus bagoti	[138]
Myrmecocystus	Wesmael	1838	29	Myrmecocystus mexicanus	Myrmecocystus mexicanus	[139]
Myrmecorhynchus	André	1896	3		Myrmecorhynchus emeryi	[140]
Myrmelachista	Roger	1863	56		Myrmelachista kraatzii	[103]
Myrmoteras	Forel	1893	41	Myrmoteras binghamii	Myrmoteras binghamii	[141]
Notoncus	Emery	1895	6		Notoncus ectatommoide	[24]
Notostigma	Emery	1920	2	Notostigma carazzii	Notostigma carazzii	[142]
Nylanderia	Emery	1906	110		Nylanderia vividula	[128]
Oecophylla	Smith	1860	15	Oecophylla smaragdina	Oecophylla smaragdina	[143]
Opisthopsis	Dalla Torre	1893	13		Opisthopsis respiciens
†	Hong	2002	1	†	N/A	[125]
†	Hong	2002	1	†	N/A	[125]
†	Hong	2002	1	†	N/A	[125]
Overbeckia	Viehmeyer	1916	1	Overbeckia subclavata	Overbeckia subclavata	[144]
Paraparatrechina	Donisthorpe	1947	36		Paraparatrechina pallida	[145]
Paratrechina	Motschoulsky	1863	5	Paratrechina currens	Paratrechina longicornis	[127][146]
Petalomyrmex	Snelling	1979	1	Petalomyrmex phylax	Petalomyrmex phylax	[147]
Plagiolepis	Mayr	1861	72	Formica pygmaea	Plagiolepis pygmaea	[67]
Polyergus	Latreille	1804	14	Formica rufescens	Polyergus rufescens	[86]
Polyrhachis	Smith	1857	698		Polyrhachis bihamata	[82]
Prenolepis	Mayr	1861	17		Prenolepis nitens	[22][67]
†	Wheeler	1915	1	†	†Prodimorphomyrmex primigenius	[9]
Proformica	Ruzsky	1902	25		Proformica nasuta	[148]
Prolasius	Forel	1892	19		Prolasius advena	[149]
†	Dlussky	1967	1	†	N/A	[150]
†Protrechina	Wilson	1985	1	†	†Protrechina carpenteri	[151]
†Pseudocamponotus	Carpenter	1930	1	†	†Pseudocamponotus elkoanus	[5]
Pseudolasius	Emery	1887	50		Pseudolasius familiaris	[22][152]
Pseudonotoncus	Clark	1934	2		Pseudonotoncus hirsutus	[153]
Rossomyrmex	Arnol'di	1928	4	Rossomyrmex proformicarum	Rossomyrmex proformicarum	[154]
Santschiella	Forel	1916	1		Santschiella kohli	[155]
†	Wheeler	1915	1	†	†Sicilomyrmex corniger	[9]
†	Hong	2002	1	†	N/A	[125]
†	Hong	2002	1	†	N/A	[125]
Stigmacros	Forel	1905	49		Stigmacros froggatti	[156]
Tapinolepis	Emery	1925	14		Tapinolepis tumidula	[157]
Teratomyrmex	McAreavey	1957	3		Teratomyrmex greavesi	[158]
†	Hong	2002	2	†	N/A	[125]
Zatania	LaPolla, Kallal & Brady	2012	6		Zatania albimaculata	[159]

Haidomyrmecinae[]

The subfamily Haidomyrmecinae contains 9 fossil genera of specialized ants described from Cretaceous ambers. The Subfamily was previously treated as the tribe Haidomyrmecini and placed within Sphecomyrminae. The tribe was elevated to a subfamily in 2020.^[160]

Haidomyrmecinae Bolton, 2003
Genus name	Genus authority	Described	No. of species	Type species	Image	Ref(s)
†	Perrichot et al.	2020	1	†		[160]
†Ceratomyrmex	Perrichot, Wang & Engel	2016	1	†Ceratomyrmex ellenbergeri	†Ceratomyrmex ellenbergeri	[161]
†	Perrichot et al.	2020	1	†		[160]
†	Perrichot et al.	2020	1	†		[160]
†Haidomyrmex	Dlussky	1996	3	†Haidomyrmex cerberus	†Haidomyrmex cerberus	[33]
†Haidomyrmodes	Perrichot et al.	2008	1	†Haidomyrmodes mammuthus	†Haidomyrmodes mammuthus	[162]
†Haidoterminus	McKellar, Glasier & Engel	2013	1	†Haidoterminus cippus	†Haidoterminus cippus	[163]
†Linguamyrmex	Barden & Grimaldi	2017	3	†Linguamyrmex vladi	†Linguamyrmex vladi	[160]
†	Perrichot et al.	2020	1	†		[160]

Heteroponerinae[]

The subfamily Heteroponerinae represents three extant genera of ants, established in 2003 when Barry Bolton divided the subfamily Ponerinae into six subfamilies.^[41] These ants are known from the Neotropics of Central America and South America while Aulacopone relicta is from Azerbaijan.^[164]

Subfamily Heteroponerinae Bolton, 2003 – three genera, 33 species[165]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acanthoponera	Mayr	1862	4	Acanthoponera mucronata	Acanthoponera mucronata	[63]
Aulacopone	Arnol'di	1930	1	Aulacopone relicta	Aulacopone relicta	[166]
Heteroponera	Mayr	1887	28	Heteroponera carinifrons	Heteroponera carinifrons	[79]

Leptanillinae[]

The subfamily Leptanillinae represents nine extant genera of ants, established in 1910 by Carlo Emery. They are subterranean ants from Africa, Europe and a single species known from Australia. Studies about their biology is minimal.^[167][168]

Subfamily Leptanillinae Emery, 1910 – nine genera, 64 species[169]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Anomalomyrma	Bolton	1990	3		Anomalomyrma taylori	[170]
Furcotanilla	Xu	2012	1	Furcotanilla furcomandibula	N/A	[171]
Leptanilla	Emery	1870	45		Leptanilla revelierii	[172]
Noonilla	Petersen	1968	1	Noonilla copiosa	Noonilla copiosa	[173]
Opamyrma	Yamane, Bui & Eguchi	2008	1	Opamyrma hungvuong	Opamyrma hungvuong	[174]
Phaulomyrma	Wheeler & Wheeler	1930	1	Phaulomyrma javana	Phaulomyrma javana	[175]
Protanilla	Taylor	1990	9		Protanilla rafflesi	[170]
Scyphodon	Brues	1925	1		Scyphodon anomalum	[176]
Yavnella	Kugler	1987	2	Yavnella argamani	Yavnella argamani	[177]

Martialinae[]

The subfamily Martialinae contains the single genus Martialis which only has one species, Martialis heureka. The ant was discovered in 2000 Amazon rainforest near Manaus, Brazil. Described in 2008, the ant belongs to the oldest known distinct lineage to have diverged from the ancestors of all other ants.^[178]

Subfamily Martialinae Rabeling & Verhaagh, 2008 – one genus, one species[179]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Martialis	Rabeling & Verhaagh	2008	1	Martialis heureka	Martialis heureka	[178]

Myrmeciinae[]

The subfamily Myrmeciinae represents two extant genera and five fossil genera that were once found worldwide.^[d] Established by Carlo Emery in 1877, the extant genera are restricted to Australia, New Caledonia and New Zealand.^[181] The notorious ant genus Myrmecia is known for their venomous stings and aggression, which has caused several human deaths in sensitive people.

Subfamily Myrmeciinae Emery, 1877 – seven genera, 111 species[182]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Archimyrmex	Cockerell	1923	4	†Archimyrmex rostratus	†Archimyrmex rostratus	[183]
†Avitomyrmex	Archibald, Cover & Moreau	2006	3	†Avitomyrmex mastax	N/A	[184]
†Macabeemyrma	Archibald, Cover & Moreau	2006	1	†Macabeemyrma ovata	†Macabeemyrma ovata	[184]
Myrmecia	Fabricius	1804	95	Myrmecia gulosa	Myrmecia gulosa	[87][136]
†Myrmeciites[d]	Archibald, Cover & Moreau	2006	0	None[c]	†Myrmeciites "incertae sedis"	[184]
Nothomyrmecia	Clark	1934	1	Nothomyrmecia macrops	Nothomyrmecia macrops	[185]
†Prionomyrmex	Mayr	1868	3	†Prionomyrmex longiceps	†Prionomyrmex longiceps	[130]
†Ypresiomyrma	Archibald, Cover & Moreau	2006	4	†Ypresiomyrma orbiculata	†Ypresiomyrma orbiculata	[184]

Myrmicinae[]

The subfamily Myrmicinae was established by Lepeletier de Saint-Fargeau in 1835. It represents 142 extant genera and 36 fossil genera that are distributed globally. It is the largest subfamily of the Formicidae, with more than 6,758 species described. The seed-harvesting ants and fungus-growing ants are well known among the Myrmicines.^[41][186]

Subfamily Myrmicinae Lepeletier de Saint-Fargeau, 1835 – 178 genera, 6,758 species[186]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Acanthognathus	Mayr	1887	8	Acanthognathus ocellatus	Acanthognathus ocellatus	[79]
Acanthomyrmex	Emery	1893	17	Acanthomyrmex luciolae	Acanthomyrmex concavus	[31]
Acromyrmex	Mayr	1865	32	Formica hystrix	Acromyrmex balzani	[187]
Adelomyrmex	Emery	1897	30		Adelomyrmex biroi	[188]
Adlerzia	Forel	1902	1	Monomorium froggatti	Adlerzia froggatti	[61]
†Afromyrma	Dlussky, Brothers & Rasnitsyn	2004	1	†Afromyrma petrosa	N/A	[189]
†Agastomyrma	Dlussky, Rasnitsyn & Perfilieva	2015	1	†	N/A	[190]
Allomerus	Mayr	1878	8	Allomerus decemarticulatus	Allomerus decemarticulatus	[191]
Ancyridris	Wheeler	1935	2		Ancyridris polyrhachioides	[192]
Anillomyrma	Emery	1913	2	Monomorium decamerum	Anillomyrma decamera	[193]
Aphaenogaster	Mayr	1853	204			[194]
Apterostigma	Mayr	1865	47		Apterostigma pilosum	[187]
Aretidris	General	2015	2		Aretidris buenaventei	[195]
Atopomyrmex	André	1889	3		Atopomyrmex mocquerysi	[196]
Atta	Fabricius	1804	17	Formica cephalotes	Atta cephalotes	[136]
†Attaichnus	Laza	1982	1	None	N/A	[197]
Austromorium	Shattuck	2009	2	Xiphomyrmex flavigaster	Austromorium flavigaster	[198]
Baracidris	Bolton	1981	3			[199]
Bariamyrma	Lattke	1990	1	Bariamyrma hispidula	Bariamyrma hispidula	[200]
Basiceros	Schulz	1906	8			[201]
†	Dlussky, Rasnitsyn & Perfilieva	2015	3	†	N/A	[190]
†	Radchenko & Dlussky	2013	2	†	N/A	[202]
Blepharidatta	Wheeler	1915	4		Blepharidatta brasiliensis	[203]
†	Radchenko & Dlussky	2012	1	†	N/A	[204]
Bondroitia	Forel	1911	2		Bondroitia lujae	[205]
†	Hong	2002	1	†	N/A	[125]
Calyptomyrmex	Emery	1887	37		Calyptomyrmex beccarii	[152]
Cardiocondyla	Emery	1869	70	Cardiocondyla elegans		[206]
Carebara	Westwood	1840	212			[207]
Cataulacus	Smith	1853	68	Cataulacus taprobanae		[208]
†	Carpenter	1930	1	†	†Cephalomyrmex rotundatus	[5]
Cephalotes	Latreille	1802	135	Formica atrata	Cephalotes atratus	[209]
Chimaeridris	Wilson	1989	2		Chimaeridris boltoni	[210]
†	Hong	2002	1	†	N/A	[125]
Colobostruma	Wheeler	1927	16			[211]
Crematogaster	Lund	1831	496	Formica scutellaris	Crematogaster scutellaris	[55]
Cryptomyrmex	Fernández	2004	2		Cryptomyrmex longinodus	[212]
Cyatta	Sosa-Calvo et al.	2013	1	Cyatta abscondita	Cyatta abscondita	[213]
Cyphoidris	Weber	1952	4		Cyphoidris spinosa	[214]
Cyphomyrmex	Mayr	1862	41		Cyphomyrmex minutus	[63]
Dacatria	Rigato	1994	1	Dacatria templaris	Dacatria templaris	[215]
Dacetinops	Brown & Wilson	1957	7		Dacetinops cibdelus	[216]
Daceton	Perty	1833	2	Formica armigera	Daceton armigerum	[217]
Diaphoromyrma	Fernández, Delabie & Nascimento	2009	1	Diaphoromyrma sofiae	Diaphoromyrma sofiae	[218]
Dicroaspis	Emery	1908	2		Dicroaspis cryptocera	[219]
Dilobocondyla	Santschi	1910	19			[220]
Diplomorium	Mayr	1901	1	Diplomorium longipenne	Diplomorium longipenne	[221]
Dolopomyrmex	Cover & Deyrup	2007	1	Dolopomyrmex pilatus	Dolopomyrmex pilatus	[222]
†Electromyrmex	Wheeler	1910	1	†Electromyrmex klebsi	†Electromyrmex klebsi	[223]
†	Mayr	1868	1	†	†Enneamerus reticulatus	[130]
†	Wilson	1985	1	†	†Eocenidris crassa	[151]
†	Dlussky & Radchenko	2006	4	†	†Eocenomyrma orthospina	[224]
†	Hong	1974	1	†	N/A	[225]
Epelysidris	Bolton	1987	1	Epelysidris brocha	Epelysidris brocha	[226]
Epopostruma	Forel	1895	19			[73]
Eurhopalothrix	Brown & Kempf	1961	53		Eurhopalothrix bolaui	[227]
Eutetramorium	Emery	1899	3		Eutetramorium mocquerysi	[228]
†	Dlussky & Radchenko	2006	1	†	N/A	[229]
Formicoxenus	Mayr	1855	7	Myrmica nitidula	Formicoxenus nitidulus	[230]
Formosimyrma	Terayama	2009	1	Formosimyrma lanuyensis	Formosimyrma lanuyensis	[231]
†	Hong	2002	1	†	N/A	[125]
Gaoligongidris	Xu	2012	1	Gaoligongidris planodorsa	N/A	[232]
Gauromyrmex	Menozzi	1933	2		Gauromyrmex bengakalisi	[233]
Goniomma	Emery	1895	8		Goniomma collingwoodi	[234]
Harpagoxenus	Forel	1893	3	Myrmica sublaevis	Harpagoxenus sublaevis	[235]
Huberia	Forel	1890	2	Tetramorium striatum	Huberia striata	[236]
Hylomyrma	Forel	1912	13		Hylomyrma balzani	[237]
†	Emery	1891	1	†	†Hypopomyrmex bombiccii	[238]
†	Wilson	1985	1	†	†Ilemomyrmex caecus	[239]
Indomyrma	Brown	1986	2	Indomyrma dasypyx	Indomyrma dasypyx	[240]
Ishakidris	Bolton	1984	1	Ishakidris ascitaspis	Ishakidris ascitaspis	[241]
Kalathomyrmex	Klingenberg & Brandão	2009	1	Myrmicocrypta emeryi	Kalathomyrmex emeryi	[242]
Kartidris	Bolton	1991	6		Kartidris nyos	[243]
Kempfidris	Fernández, Feitosa & Lattke	2014	1	Monomorium inusuale	Kempfidris inusualis	[244]
Lachnomyrmex	Wheeler	1910	16		Lachnomyrmex scrobiculatus	[8]
Lasiomyrma	Terayama & Yamane	2000	4		Lasiomyrma gedensis	[245]
Lenomyrmex	Fernández & Palacio	1999	6			[246]
Leptothorax	Mayr	1855	19	Formica acervorum	Leptothorax acervorum	[230]
Liomyrmex	Mayr	1865	1		Liomyrmex gestroi	[187]
†	Mayr	1867	2	†	N/A	[247]
Lophomyrmex	Emery	1892	13	Oecodoma quadrispinosa		[248]
Lordomyrma	Emery	1897	34		Lordomyrma furcifera	[188]
Malagidris	Bolton & Fisher	2014	6		Malagidris belti	[249]
Manica	Jurine	1807	6	Formica rubida	Manica rubida	[88]
Mayriella	Forel	1902	9		Mayriella abstinens	[61]
Megalomyrmex	Forel	1885	44		Megalomyrmex leoninus	[250]
Melissotarsus	Emery	1877	4		Melissotarsus beccarii	[251]
Meranoplus	Smith	1853	90	Cryptocerus bicolor	Meranoplus bicolor	[208]
Mesostruma	Brown	1948	9		Mesostruma turneri	[252]
Messor	Forel	1890	110	Formica barbara	Messor barbarus	[253]
Metapone	Forel	1911	18	Metapone greeni		[254]
Microdaceton	Santschi	1913	4		Microdaceton exornatum	[255]
†	Zhang	1989	1	†	N/A	[43]
Monomorium	Mayr	1855	372		Monomorium pharaonis	[230]
Mycetagroicus	Brandão & Mayhé-Nunes	2001	4		Mycetagroicus cerradensis	[256]
Mycetarotes	Emery	1913	4		Mycetarotes parallelus	[193]
Mycetophylax	Emery	1913	3		Mycetophylax conformis	[193]
Mycetosoritis	Wheeler	1907	5		Mycetosoritis hartmanni	[257]
Mycocepurus	Forel	1893	6	Atta smithii	Mycocepurus smithii	[258]
Myrmecina	Curtis	1829	51	Myrmecina latreillii	Myrmecina graminicola	[259]
†	Dlussky & Rasnitsyn	2003	6	†	N/A	[260]
Myrmica	Latreille	1804	211	Formica rubra	Myrmica rubra	[86]
Myrmicaria	Saunders	1842	32	Myrmicaria brunnea	Myrmicaria brunnea	[261]
Myrmicocrypta	Smith	1860	27		Myrmicocrypta squamosa	[262]
Myrmisaraka	Bolton & Fisher	2014	2		Myrmisaraka producta	[249]
Nesomyrmex	Wheeler	1910	57			[8]
Novomessor	Emery	1915	3	Aphaenogaster cockerelli	Novomessor cockerelli	[263]
Ochetomyrmex	Mayr	1878	2		Ochetomyrmex semipolitus	[191]
Octostruma	Forel	1912	34		Octostruma iheringi	[264]
Ocymyrmex	Emery	1886	37			[265]
†	Hong	2002	1	†	N/A	[125]
Orectognathus	Smith	1853	29		Orectognathus antennatus	[208]
Oxyepoecus	Santschi	1926	21	Oxyepoecus bruchi	Oxyepoecus bruchi	[266]
†	Wilson	1985	1	†	†Oxyidris antillana	[239]
Oxyopomyrmex	André	1881	12		Oxyopomyrmex oculatus	[267]
†	Wheeler	1915	1	†	†Parameranoplus primaevus	[9]
Paramycetophylax	Kusnezov	1956	1	Sericomyrmex bruchi	Paramycetophylax bruchi	[268]
†	Dlussky	1981	4	†	†Paraphaenogaster hooleyana	[64]
Paratopula	Wheeler	1919	11	Atopomyrmex ceylonicus		[269]
Patagonomyrmex	Johnson & Moreau	2016	3			[270]
Perissomyrmex	Smith	1947	6		Perissomyrmex snyderi	[271]
Peronomyrmex	Viehmeyer	1922	3			[272]
Phalacromyrmex	Kempf	1960	1	Phalacromyrmex fugax	Phalacromyrmex fugax	[273]
Pheidole	Westwood	1839	1007		Pheidole rhea	[274]
Pilotrochus	Brown	1978	1	Pilotrochus besmerus	Pilotrochus besmerus	[275]
†	Dlussky & Radchenko	2009	1	†	N/A	[276]
Podomyrma	Smith	1859	52			[277]
Poecilomyrma	Mann	1921	2		Poecilomyrma senirewae	[278]
Pogonomyrmex	Mayr	1868	69	Formica badia	Camponotus badius	[118]
Pristomyrmex	Mayr	1866	60		Pristomyrmex punctatus	[96]
Proatta	Forel	1912	1	Proatta butteli	Proatta butteli	[279]
Procryptocerus	Emery	1887	45			[152]
Propodilobus	Branstetter	2009	1		Propodilobus orientale	[280]
Protalaridris	Brown	1980	1	Protalaridris armata	Protalaridris armata	[281]
†	Dlussky & Radchenko	2009	1	†	†Protomyrmica atavia	[276]
Pseudoatta	Gallardo	1916	1	Pseudoatta argentina	Pseudoatta argentina	[282]
†	Hong	2002	1	†	N/A	[125]
†	Hong	2002	1	†	N/A	[125]
Recurvidris	Bolton	1992	11	None		[283]
Rhopalomastix	Forel	1900	6		Rhopalomastix rothneyi	[284]
Rhopalothrix	Mayr	1870	16		Rhopalothrix ciliata	[285]
Rogeria	Emery	1894	40		Rogeria curvipubens	[286]
Romblonella	Wheeler	1935	9		Romblonella opaca	[192]
Rostromyrmex	Rosciszewski	1994	1	Rostromyrmex pasohensis	Rostromyrmex pasohensis	[287]
Rotastruma	Bolton	1991	2		Rotastruma recava	[243]
Royidris	Bolton & Fisher	2014	15		Royidris robertsoni	[249]
Secostruma	Bolton	1988	1	Secostruma lethifera	Secostruma lethifera	[288]
Sericomyrmex	Mayr	1865	19	Sericomyrmex opacus	Sericomyrmex amabilis	[187]
†	Hong	2002	2	†	N/A	[125]
Solenopsis	Westwood	1840	205	Solenopsis mandibularis	Solenopsis geminata	[207]
†	Dlussky & Rasnitsyn	2003	3	†	†Solenopsites rossi	[260]
†	Hong	2002	1	†	N/A	[125]
Stegomyrmex	Emery	1912	5			[289]
Stenamma	Westwood	1839	85		Stenamma westwoodii	[274]
Stereomyrmex	Emery	1901	3	Stereomyrmex horni	Stereomyrmex horni	[290]
†	Mayr	1868	1	†	†Stigmomyrmex venustus	[130]
†	Wheeler	1915	1	†	†Stiphromyrmex robustus	[9]
Strongylognathus	Mayr	1853	24	Eciton testaceum	Strongylognathus testaceus	[291][292]
Strumigenys	Smith	1860	842			[262]
Syllophopsis	Santschi	1915	20		Syllophopsis modesta	[293]
Talaridris	Weber	1941	1	Talaridris mandibularis	Talaridris mandibularis	[48]
Temnothorax	Mayr	1861	395		Temnothorax unifasciatus	[67]
Terataner	Emery	1912	12		Terataner foreli	[289]
Tetheamyrma	Bolton	1991	1	Tetheamyrma subspongia	Tetheamyrma subspongia	[243]
Tetramorium	Mayr	1855	569	Formica caespitum	Tetramorium caespitum	[230]
Trachymyrmex	Forel	1893	48	Atta septentrionalis	Trachymyrmex septentrionalis	[258]
Tranopelta	Mayr	1866	2		Tranopelta gilva	[25]
Trichomyrmex	Mayr	1865	18	Trichomyrmex rogeri	Trichomyrmex destructor	[187]
Tropidomyrmex	Silva, Feitosa, Brandão & Diniz	2009	1	Tropidomyrmex elianae	Tropidomyrmex elianae	[294]
Tyrannomyrmex	Fernández	2003	3	Tyrannomyrmex rex		[295]
Veromessor	Forel	1917	9		Veromessor andrei	[296]
Vitsika	Bolton & Fisher	2014	14		Vitsika crebra	[249]
Vollenhovia	Mayr	1865	62		Vollenhovia emeryi	[187]
Vombisidris	Bolton	1991	17		Vombisidris philax	[243]
Wasmannia	Forel	1893	11	Tetramorium auropunctatum	Wasmannia auropunctata	[297]
†	Hong	2002	1	†	N/A	[125]
Xenomyrmex	Forel	1885	4		Xenomyrmex stollii	[250]
†	Bolton	2003	1	†	N/A	[6]

Paraponerinae[]

The subfamily Paraponerinae contains a single genus Paraponera. This genus has two species, one of which was found in Dominican amber from the Miocene.^[298] The extant species, Paraponera clavata, is found in Central America and South America, and the pain from its sting is said to be greater than any other insect sting on earth.^[299]

Subfamily Paraponerinae Emery, 1901 – one genus, two species[300]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Paraponera	Smith	1858	2	Paraponera clavata	Paraponera clavata	[102]

Ponerinae[]

The subfamily Ponerinae was established by Lepeletier de Saint-Fargeau in 1835, which represents 47 extant genera and 12 fossil genera. The subfamily is among the most diverse in the family Formicidae, with more than 1,000 species described. They are mostly distributed in the tropics and subtropics.^[6][41]

Subfamily Ponerinae Lepeletier de Saint-Fargeau, 1835 – 59 genera, 1,287 species[301]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Afropone	Dlussky et al.	2004	2	Afropone oculata	N/A	[189]
Anochetus	Mayr	1861	122	Anochetus ghilianii	Anochetus ghilianii	[67]
†Archiponera	Carpenter	1930	1	†Archiponera wheeleri	†Archiponera wheeleri	[5]
Asphinctopone	Santschi	1914	3		Asphinctopone silvestrii	[27]
Austroponera	Schmidt & Shattuck	2014	3		Austroponera rufonigra	[302]
Belonopelta	Mayr	1870	2		Belonopelta attenuata	[285]
Boloponera	Fisher	2006	1	Boloponera vicans	Boloponera vicans	[303]
Bothroponera	Mayr	1862	37		Bothroponera pumicosa	[63]
Brachyponera	Emery	1900	19		Brachyponera croceicornis	[304]
Buniapone	Schmidt & Shattuck, 2014	2014	1	Buniapone amblyops	Buniapone amblyops	[302]
Centromyrmex	Mayr	1866	15		Centromyrmex bohemanni	[96]
†Cephalopone	Dlussky & Wedmann	2012	2	†Cephalopone potens	†Cephalopone potens	[13]
Cryptopone	Emery	1893	24	Cryptopone testacea	Cryptopone testacea	[31]
†Cyrtopone	Dlussky & Wedmann	2012	4	†Cyrtopone microcephala	†Cyrtopone microcephala	[13]
Diacamma	Mayr	1862	24	Diacamma rugosum	Diacamma rugosum	[63]
Dinoponera	Roger	1861	8	Dinoponera gigantea	Dinoponera gigantea	[21]
Dolioponera	Brown	1974	1	Dolioponera fustigera	Dolioponera fustigera	[305]
Ectomomyrmex	Mayr	1867	28		Ectomomyrmex javanus	[22][306]
Emeryopone	Forel	1912	5		Emeryopone buttelreepeni	[279]
†	Hong	2002	1	†	N/A	[125]
†	Hong	2002	1	†	N/A	[125]
Euponera	Forel	1891	27		Euponera sikorae	[95]
Feroponera	Bolton & Fisher	2008	1	Feroponera ferox	Feroponera ferox	[307]
Fisheropone	Schmidt & Shattuck	2014	1	Fisheropone ambigua	Fisheropone ambigua	[302]
†	Hong	2002	1	†	N/A	[125]
Hagensia	Forel	1901	2		Hagensia havilandi	[308]
Harpegnathos	Jerdon	1851	7	Harpegnathos saltator	Harpegnathos saltator	[309]
Hypoponera	Santschi	1938	154		Hypoponera abeillei	[310]
Iroponera	Schmidt & Shattuck	2014	1	Iroponera odax	Iroponera odax	[302]
Leptogenys	Roger	1861	308	Leptogenys falcigera	Leptogenys falcigera	[21]
Loboponera	Bolton & Brown	2002	9		Loboponera vigilans	[311]
†	Hong	2002	1	†	N/A	[125]
Mayaponera	Schmidt & Shattuck	2014	1	Mayaponera constricta	Mayaponera constricta	[302]
Megaponera	Mayr	1862	1	Megaponera analis	Megaponera analis	[63]
Mesoponera	Emery	1900	20	Mesoponera melanaria	Mesoponera melanaria	[312]
†Messelepone	Dlussky & Wedmann	2012	1	†Messelepone leptogenoides	†Messelepone leptogenoides	[13]
Myopias	Roger	1861	40	Myopias amblyops	Myopias amblyops	[21]
Neoponera	Emery	1901	57		Neoponera villosa	[313]
Odontomachus	Latreille	1804	70		Odontomachus haematodus	[86]
Odontoponera	Mayr	1862	2		Odontoponera denticulata	[63]
Ophthalmopone	Forel	1890	5		Ophthalmopone berthoudi	[236]
Pachycondyla	Smith	1858	36		Pachycondyla crassinoda	[102]
Paltothyreus	Mayr	1862	1		Paltothyreus tarsatus	[313]
Parvaponera	Schmidt & Shattuck	2014	4		Parvaponera darwinii	[302]
Phrynoponera	Wheeler	1920	5	Phrynoponera gabonensis	Phrynoponera gabonensis	[124]
Platythyrea	Roger	1863	44	Platythyrea punctata	Platythyrea punctata	[22][103]
Plectroctena	Smith	1858	17	Plectroctena mandibularis	Plectroctena mandibularis	[102]
Ponera	Latreille	1804	62		Ponera coarctata	[86][274]
†	Dlussky & Rasnitsyn	2003	14	†	†Ponerites eocenicus	[260]
Promyopias	Santschi	1914	1		Promyopias silvestrii	[27]
†Protopone	Dlussky	1988	7	†Protopone primigena	†Protopone primigena	[30]
Psalidomyrmex	André	1890	6		Psalidomyrmex foveolatus	[314]
Pseudoneoponera	Donisthorpe	1943	18		Pseudoneoponera verecundae	[315]
Pseudoponera	Emery	1900	6		Pseudoponera stigma	[304]
Rasopone	Schmidt & Shattuck	2014	11		Rasopone ferruginea	[302]
Simopelta	Mann	1922	22		Simopelta jeckylli	[316]
Streblognathus	Mayr	1862	2		Streblognathus aethiopicus	[63]
†	Dlussky & Perfilieva	2014	6	†	†Taphopone macroptera	[32]
Thaumatomyrmex	Mayr	1887	12		Thaumatomyrmex mutilatus	[79]

Proceratiinae[]

The subfamily Proceratiinae was established by Italian entomologist Carlo Emery in 1895, which represents three extant genera and one extinct genus. Found worldwide, these ants are mainly encountered in tropical and subtropical areas. Little is known about their biology.^[41][317]

Subfamily Proceratiinae Emery, 1895 – four genera, 155 species[318]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†Bradoponera	Mayr	1868	4	†	†Bradoponera meieri	[130]
Discothyrea	Roger	1863	37	Discothyrea testacea	Discothyrea testacea	[103]
Probolomyrmex	Mayr	1901	26		Probolomyrmex filiformis	[221]
Proceratium	Roger	1863	88	Proceratium silaceum	Proceratium silaceum	[103]

Pseudomyrmecinae[]

The subfamily Pseudomyrmecinae was established by M.R. Smith in 1952,^[319] which represents three genera of ants that are primarily arboreal nesting ants in the tropical and subtropical regions.^[320] They are found in Africa, Asia, Australia, North America and South America.^[321]

Subfamily Pseudomyrmecinae Smith, 1952 – three genera, 251 species[322]
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
Myrcidris	Ward	1990	1	Myrcidris epicharis	Myrcidris epicharis	[320]
Pseudomyrmex	Lund	1831	147	Pseudomyrmex gracilis	Pseudomyrmex gracilis	[319][323]
Tetraponera	Smith	1852	103	Tetraponera nigra	Tetraponera nigra	[324]

Sphecomyrminae[]

The subfamily Sphecomyrminae contains 9 fossil genera of stem ants. Most fossilized ants from Cretaceous amber were placed in this subfamily, however revisions in 2017 and 2020 removed several genera and added former members of the subfamily Armaniinae.^{[1][325][160]}

Subfamily Sphecomyrminae Wilson & Brown, 1967
Genus name	Binomial authority	Classified	No. of species	Type species	Example image	Ref(s)
†	Dlussky	1983	4	†	N/A	[326]
†Boltonimecia	Borysenko	2017	1	†	†Boltonimecia canadensis	[327]
†	Dlussky	1975	2	†	N/A	[328]
†Gerontoformica	Nel & Perrault	2004	13	†Gerontoformica cretacica	†Gerontoformica cretacica	[329]
†	Dlussky, Brothers & Rasnitsyn	2004	2	†	†Orapia rayneri	[189]
†	Dlussky	1983	2	†	N/A	[326]
†Sphecomyrma	Wilson & Brown	1967	2	†Sphecomyrma freyi	†Sphecomyrma freyi	[330]
†Zigrasimecia	Barden & Grimaldi	2013	2	†Zigrasimecia tonsora	†Zigrasimecia tonsora	[325]

Incertae sedis[]

There are several ant genera where their taxonomic placement is uncertain (incertae sedis). These genera have not yet been assigned to any subfamily within Formicidae; 16 genera are currently listed as incertae sedis.^[2]

Incertae sedis genera
Genus name	Genus authority	Described	No. of species	Type species	Image	Ref(s)
†	Dlussky	1975	2	†	N/A	[328]
†Baikuris	Dlussky	1987	4	†Baikuris mandibularis	†Baikuris mandibularis	[331]
†	Scudder	1877	1	†	N/A	[5][332]
†Camelomecia	Barden & Grimaldi	2016	1	†Camelomecia janovitzi	†Camelomecia janovitzi	[333]
†	Dlussky	1975	1	†	†Cretopone magna	[334]
†[e]	Hong	2002	1	†	N/A	[6][125]
†Dlusskyidris	Bolton	1994	1	†Dlusskyidris zherichini	†Dlusskyidris zherichini	[328][336]
†[f]	Hong	2002	1	†	N/A	[125]
†	Cockerell	1921	6	†	†Eoformica pinguis	[260][338]
†	Martins-Neto & Mendes	2002	1	†	N/A	[339]
†Klondikia	Dlussky & Rasnitsyn	2003	1	†Klondikia whiteae	N/A	[260]
†	Dlussky & Rasnitsyn	2003	3	†	N/A	[260]
†Myanmyrma	Engel & Grimaldi	2005	1	†Myanmyrma gracilis	†Myanmyrma gracilis	[34]
†	Dlussky	1975	1	†	†Petropone petiolata	[334]
†	Dlussky	1983	1	†	N/A	[326]

Formerly included in Formicidae[]

There are several genera which were formerly placed in Formicidae, but have subsequently been removed. These genera are now placed in other families, are considered incertae sedis within Hymenoptera taxonomy, or are considered invalid.^[340]

Excluded genera
Genus name	Genus authority	Described	No. of species	Type species	Current placement	Image	Ref(s)
Condylodon	Lund	1831	1	Condylodon audouini	Unidentifiable, considered invalid by Bolton, 2020	N/A	[6][55]
†	Dlussky	1975	2	†	Incertae sedis in Aculeata	N/A	[328][327]
	Buckley	1866	1		Unidentifiable, considered invalid by Bolton, 2020	N/A	[341][342]
†	Dlussky	1999	1	†	Incertae sedis in Aculeata	N/A	[343]

See also[]

• List of ant subfamilies

Notes[]

References[]