Pellaea glabella

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Smooth cliffbrake
Pellaea glabella fronds.jpg

Secure (NatureServe)
Scientific classification edit
Kingdom: Plantae
Clade: Tracheophytes
Division: Polypodiophyta
Class: Polypodiopsida
Order: Polypodiales
Family: Pteridaceae
Genus: Pellaea
Species:
P. glabella
Binomial name
Pellaea glabella
Mett. ex Kuhn
Synonyms
  • Pellaea atropurpurea (L.) Link var. glabella (Mett. ex Kuhn) F.C.Gates
  • Pellaea atropurpurea (L.) Link var. bushii Mack.

Pellaea glabella is the smooth cliffbrake. For much of pteridological history, it was regarded as a reduced form or variety of Pellaea atropurpurea. P. glabella is known to exist in two cryptic species, one diploid and one tetraploid. The diploid reproduces sexually, while the tetraploid is normally apogamous. It is now known that the tetraploid form of the species is one of the parents of the original hybrid P. × atropurpurea that became the apogamous species.

P. glabella is epipetric, normally growing on well-weathered limestone. It favors more exposed sites than P. atropurpurea.

This species is distinguished by its sessile or nearly sessile pinnae and smooth, not hairy, stipes.

P. glabella has been assigned a total of four subspecies:

  • Pellaea glabella subsp. glabella Mettenius ex Kuhn
  • Pellaea glabella subsp. missouriensis (G. J. Gastony) Windham
  • Pellaea glabella subsp. occidentalis (E. E. Nelson) Windham
  • Pellaea glabella subsp. simplex (Butters) A. Löve & D. Löve

The subspecies glabella and simplex are the tetraploids, while missouriensis and occidentalis are the diploids. Glabella and missouriensis have hairlike scales near the midrib, while simplex and occidentalis are completely glabrous.

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