Permocupedidae

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Permocupedidae
Temporal range: Cisuralian–Anisian
Permocupedidae.jpg
Fossil and life restoration of Permocupes sojanensis
Scientific classification e
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Suborder: †Protocoleoptera
Superfamily: †Permocupedoidea
Family: †Permocupedidae
Martynov, 1933
Genera
  • See text
Synonyms
  • Taldycupedidae

Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the Middle Permian. They became rare in the Late Permian, with only one species known from the Triassic, from the Middle Triassic (Anisian) Röt Formation of Germany.[1] They are thought to have been xylophagous (wood eating), which is presumed to be the ancestral ecology of beetles.[2] One monotypic with the species , was described in 2014 from late Permian fossils in found in China.[3]

Taxonomy[]

Kirejtshuk (2020) included the following genera in an expanded (sensu lato) family, with the traditional (sensu stricto) Permocupedidae and related Taldycupedidae as subfamilies Permocupedinae and Taldycupedinae respectively.[4]

Subfamily Permocupedinae Martynov, 1932

  • Geertsema & van den Heever, 1996 South Africa: Western Cape (Kenmoore Farm), Whitehill Formation, Permian, Cisuralian, Sakmarian/Artinskian, −290.1–279.5 Ma.
    • Geertsema & van den Heever, 1996 (Type species)
  • Rohdendorf, 1961 (syn Palaeocupes Rohdendorf, 1961) Russia, Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma
    • Rohdendorf, 1961 (Type species)
  • Ponomarenko, 1969 Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Ponomarenko, 1969 (Type species)
  • Rohdendorf, 1961Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Rohdendorf, 1961 (Type species)
  • Rohdendorf, 1961Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma).
    • Ponomarenko, 1969
    • Rohdendorf, 1961 (Type species)
  • Ponomarenko & Bashkuev, 2018 Germany: Gambach am Main (Lower Franconia), Röt Formation Middle Triassic, Olenekian/Anisian, −247.2–242.0 Ma.
    • Ponomarenko & Bashkuev, 2018 (Type species)
  • Rohdendorf, 1961 Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Kemerovo Region (Tykhta River, Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma.
    • Ponomarenko, 1969
    • (Rohdendorf, 1961)
    • Rohdendorf, 1961 (Type species)
  • Rohdendorf, 1961. Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Rohdendorf, 1961 (Type species)
  • Rohdendorf, 1961Russia: Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Orenburg Region (Kargala), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Sweverodvinian), −265.0–259.0 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. One undescribed species of this genus was recorded from Indian of the Lower Permian.
    • Ponomarenko, 1963
    • Ponomarenko, 1963
    • Ponomarenko, 1963
    • Rohdendorf, 1961 (Type species)
    • Rohdendorf, 1961
  • Ponomarenko, Yan & Huang, 2014 China: Anhui (Houdong, Yinping Mountain, SW Chaohu City), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian/Vyatkian), −265.0–259.0 Ma.
    • Ponomarenko, Yan & Huang, 2014 (Type species)
  • Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 Russia, Primorsky Krai (Cape, Novosilsky, Russky Island), Lower Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
    • Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 (Type species)
  • Martynov, 1932 (syn Permocupoides Martynov, 1932) Russia: Arkhangel’sk Region (Letopala River and Soyana), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma: Tatarstan (Mendeleevsk District, Tikhie Gory), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • (Ponomarenko, 1963
    • Ponomarenko, 2013
    • Martynov, 1932
    • (Ponomarenko, 1963
  • Kirejtshuk, 2020. Brazil: Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
    • (Pinto, 1987) (Type species)
  • Rohdendorf, 1961 (syn Tomiocupes Rohdendorf, 1961, Tricupes Rohdendorf, 1961) Russia: Vologda Region (Isady locality, Mutovino), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinskian/Vyatkian), −259.0–254.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma; Kemerovo Region (Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • (Rohdendorf, 1961)
    • (Rohdendorf, 1961)
    • Ponomarenko, 2013
    • Ponomarenko, 2013
    • Rohdendorf, 1961
    • Rohdendorf, 1961
    • Pinto, 1987 Brazil (Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Kungurian, −279.5–272.5 Ma) considered indeterminate to genus by Kirejtshuk (2020)
  • Ponomarenko, 2004 Russia: Orenburg Region (Novo-Aleksandrovka), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian), −265.0–259.0 Ma.
    • Ponomarenko, 2004 (Type species)
  • Ponomarenko, 1969 Russia: Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
    • Ponomarenko, 1963 (Type species)

Subfamily Taldycupedinae Rohdendorf, 1961

  • Kirejtshuk, 2020 Beaufort Group, South Africa, Late Permian (Changhsingian)
    • (Ponomarenko, 2005) (Type species)
    • (Ponomarenko, 2005)
  • Tillyard, 1916 Blackstone Formation, Australia Late Triassic (Norian)
    • Tillyard, 1916 (Type species)
    • Dunstan, 1923
    • Dunstan, 1923
  • Ponomarenko, 2015 (syn Lobanovia Kirejtshuk, 2020)
    • Ponomarenko, 2015 , Vologda, Russia, Late Permian (Changhsingian)
    • Ponomarenko, 2015 Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian)
    • (Ponomarenko, 2015) (Type species) , Vologda, Russia, Late Permian (Wuchiapingian)
  • Dunstan, 1923 Blackstone Formation, Australia Late Triassic (Norian)
    • Dunstan, 1923
    • Dunstan, 1923 (Type species)
  • Rohdendorf, 1961 (syn Cryptocupes Rohdendorf, 1961, Taldycupidium Rohdendorf, 1961)
    • (Rohdendorf, 1961) , Kemerovo Oblast, Russia, Late Permian
    • Ponomarenko, 2013 Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian)
    • Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
    • Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
    • Ponomarenko et al. 2014 , Anhui, China, Middle Permian (Capitanian)
    • Ponomarenko, 1969 , Kazakhstan, Late Permian (Changhsingian)
    • Ponomarenko, 2006 , Krasnoyarsk, Russia, Late Permian (Changhsingian)
    • Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
  • Rohdendorf, 1961 (syn Stegocupes Rohdendorf, 1961
    • Rohdendorf, 1961 (Type species) , Kemerovo, Russia, Middle Permian (Wordian) , Kemerovo, Russia, Middle Permian (Roadian)
    • Rohdendorf, 1961 Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian)
    • Ponomarenko, 2013 . Orenburg, Middle Permian (Capitanian)
  • Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
    • Rohdendorf, 1961
    • Rohdendorf, 1961
    • Rohdendorf 1961
    • "" Rohdendorf 1961 (nomen dubium)
  • Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
    • Rohdendorf, 1961
  • Rohdendorf, 1961 (nomen dubium) Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian

References[]

  1. ^ Ponomarenko, Alexandr G.; Bashkuev, Alexei S. (December 2018). "First Triassic record of the beetle family Permocupedidae (Insecta: Coleoptera): a peculiar example of a Lazarus taxon". PalZ. 92 (4): 587–591. doi:10.1007/s12542-018-0426-4. ISSN 0031-0220.
  2. ^ Zhao, Xianye; Yu, Yilun; Clapham, Matthew E; Yan, Evgeny; Chen, Jun; Jarzembowski, Edmund A; Zhao, Xiangdong; Wang, Bo (2021-11-08). Perry, George H; Fikacek, Martin (eds.). "Early evolution of beetles regulated by the end-Permian deforestation". eLife. 10: e72692. doi:10.7554/eLife.72692. ISSN 2050-084X.
  3. ^ A. G. Ponomarenko, E. V. Yan and Di-ying Huang (2014). "New beetles (Coleoptera) from the terminal Middle Permian of China". Paleontological Journal 48 (2): 191–200. doi:10.1134/S0031030114010109.
  4. ^ Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.
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