Berothidae

Berothidae Temporal range: Late Jurassic–Recent PreꞒ Ꞓ O S D C P T J K Pg N
sp.
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Neuroptera
Suborder:	Hemerobiiformia
Superfamily:	Mantispoidea
Family:	Berothidae Handlirsch, 1906
Subfamilies
and see text

The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings.^[1] The family was first named by Anton Handlirsch in 1906.^[2]

Berothidae is very closely related to the thorny lacewings which are often included in the Berothidae as subfamily "Rhachiberothinae". They are also occasionally considered to be the distinct family Rhachiberothidae or included as a subfamily in the mantidflies.^[2] The family consists of 22 genera and 100 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions.^[2] Eleven extinct genera with a total of 13 species have been described from the fossil record.^[2]

The larvae are associated with termites.^[1]

Systematics[]

The peculiar genus Lomamyia cannot be robustly assigned to any one subfamily. In addition, a considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or incertae sedis:

• Ren & Guo, 1996 (Late Jurassic; China)
• Microberotha

Extinct genera[]

• † Yang et al. 2019 Burmese amber, Myanmar, Cenomanian
• † Martins-Neto and Vulcano 1990 Crato Formation, Brazil, Aptian
• † Whalley 1980 Lebanese amber, Barremian
• † Martins-Neto and Vulcano 1990 Crato Formation, Brazil, Aptian
• † Pérez-de la Fuente, Peñalver & Engel, 2020, , Albian
• † Yang et al. 2019 Burmese amber, Myanmar, Cenomanian
• † Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
• † Makarkin and Ohl 2015 Baltic amber, Eocene
• † Jepson et al. 2012 Lulworth Formation, United Kingdom, Berriasian
• † Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
• † Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
• † Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
• † Grimaldi 2000 Burmese amber, Myanmar, Cenomanian New Jersey amber, Turonian
• † Khramov 2015 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
• † Yuan et al. 2016 Burmese amber, Myanmar, Cenomanian
• † Yuan et al. 2016 Burmese amber, Myanmar, Cenomanian
• †subfamily Mesithoninae Panfilov 1980
  • † Khramov 2015 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
  • † Panfilov 1980 Karabastau Formation, Kazakhstan, Callovian/Oxfordian Zaza Formation, , Russia, Aptian
  • † Makarkin 1999 Zaza Formation, Russia, Aptian
• †Microberotha Archibald and Makarkin 2004 , Canada, Ypresian
• † Grimaldi 2000 New Jersey amber, Turonian
• † Ren and Guo 1996 Yixian Formation, China, Aptian
• † Huang et al. 2019 Burmese amber, Myanmar, Cenomanian
• † Azar and Nel 2013 Lebanese amber, Barremian
• † Hong 1983 Daohugou, Haifanggou Formation, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian, , Mongolia, Tithonian
• † Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
• † Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
• † Shi et al. 2019 Burmese amber, Myanmar, Cenomanian
• † Makarkin 2017 Green River Formation, United States, Eocene

References[]

• Bugguide.net. Family Berothidae - Beaded Lacewings

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