Coptoclavidae

Coptoclavidae Temporal range: Carnian–Aptian PreꞒ Ꞓ O S D C P T J K Pg N
Holcoptera giebeli
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Adephaga
Family:	†Coptoclavidae , 1961
Subfamilies
Coptoclavinae Hispanoclavininae Coptoclavidae (incertae sedis)

Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga.^[1][2] The Coptoclavidae lived from the Triassic to the Early Cretaceous.^[3] Coptoclavidae is a clade of .^[4][5][6] The increase in the teleost fish population and the competition of Gyrinidae and Dytiscidae is believed to have caused their extinction.^[7]

Taxonomy[]

• † Lin 1992 , China, Carnian
• † Bode 1953 Posidonia Shale, Germany, Toarcian
• †subfamily Charonoscaphinae Ponomarenko 1977
  • † Ponomarenko 1977 Karabastau Formation, Kazakhstan, Callovian
  • † Ponomarenko 1977 Karabastau Formation, Kazakhstan, Callovian
• † Ping 1928 , Upper Triassic
• †subfamily Coptoclavinae Ponomarenko 1961
  • † Ponomarenko 1987
    • †Bolbonectes intermedius Ponomarenko 1987 , Russia, Tithonian , Russia, Barremian
    • †Bolbonectes occidentalis Ponomarenko 1993 , Russia, Oxfordian
    • †Bolbonectus lithographicus Ponomarenko and Martínez-Delclòs 2000 La Pedrera de Rúbies Formation, Spain, Barremian
  • † Ping 1928 , Russia, Hauterivian Dabeigou Formation, China, Hauterivian, , Russia, Barremian , , , , Jiufotang Formation, , Aptian Zaza Formation, , , , Russia, Aptian
  • † Soriano et al. 2007 Las Hoyas, Spain, Barremian
  • † Ponomarenko and Martínez-Delclòs 2000 Las Hoyas, Spain, Barremian
• †subfamily Coptoclaviscinae Soriano et al. 2007
  • †Coptoclavella Ponomarenko 1980
    • †Coptoclavella elegans Ponomarenko 1980 , Mongolia, Aptian
    • †Coptoclavella inexpecta Soriano et al. 2007 La Pedrera de Rúbies Formation, Spain, Barremian
    • †Coptoclavella jurassica Ponomarenko 2014 Sharteg, Mongolia, Tithonian
    • †Coptoclavella minor Ponomarenko 1980 , Russia, Callovian, , Russia, Hauterivian Mogotuin Formation, Mongolia, Aptian Turga Formation, Russia, Aptian
    • †Coptoclavella purbeckensis Ponomarenko et al. 2005, Durlston Formation, United Kingdom, Berriasian
    • †Coptoclavella vittata Ponomarenko 1986 , Mongolia, Aptian
    • †Coptoclavella striata Ponomarenko 1986 Gurvan-Eren Formation, Mongolia, Aptian
  • † Ponomarenko 1987 Laiyang Formation, China, Aptian Tsagaantsav Formation, Mongolia, Early Cretaceous
  • † Ponomarenko 2015 Röt Formation, Germany, Anisian, Hassberge Formation, Germany, Carnian
• † Lin 1992 Huangshanjie Formation, China, Carnian
• † Handlirsch 1906
  • †Holcoptera alisonae Kelly et al. 2017 Charmouth Mudstone Formation, United Kingdom, Sinemurian
  • †Holcoptera giebeli Handlirsch 1906 Lilstock Formation, United Kingdom, Rhaetian, Charmouth Mudstone Formation, United Kingdom, Sinemurian,
  • †Holcoptera pigmentatus Kelly et al. 2017 Lilstock Formation, United Kingdom, Rhaetian Portland Formation, Connecticut, Hettangian Mount Toby Formation, Massachusetts, Hettangian
  • †Holcoptera schlotheimi Giebel 1856 (=Holcoptera confluens Cockerell 1915) , Italy, Norian, Lilstock Formation, United Kingdom, Rhaetian, Blue Lias, United Kingdom, Hettangian, Portland Formation, Connecticut, Hettangian
  • †Holcoptera solitensis Kelly et al. 2017 Cow Branch Formation, North Carolina, Norian
• †subfamily Hispanoclavinae Soriano et al. 2007
  • † Soriano et al. 2007 Las Hoyas, Spain, Barremian
• †subfamily Timarchopsinae Wang et al. 2010
  • † Germar 1842 Solnhofen, Germany, Tithonian
  • † Wang et al. 2009 Daohugou, China, Callovian
  • † Germar 1839 Solnhofen, Germany, Tithonian
  • † Ponomarenko 1977 Karabastau Formation, Kazakhstan, Callovian
  • † Soriano et al. 2007 Las Hoyas, Spain, Barremian
  • † Prokin and Ponomarenko 2013 Hassberge Formation, Germany, Carnian
  • † Deichmüller 1886 Solnhofen, Germany, Tithonian
  • † Ponomarenko 1977 , Russia, Toarcian, , Russia, Callovian, , Ichetuy Formation, Russia, Oxfordian, , Russia, Tithonian , Bukachacha Formation, Russia, Barremian, , Turga Formation, Russia, Aptian
  • † Brauer et al. 1889
    • †Timarchopsis aquaticus Ponomarenko 1977 Cheremkhovskaya Formation, Russia, Toarcian, Ichetuy Formation, Russia, Oxfordian
    • †Timarchopsis cyrenaicus Ponomarenko 1977 Algeria, Hauterivian
    • †Timarchopsis czekanowskii Brauer et al. 1889 Cheremkhovskaya Formation, Russia, Toarcian
    • †Timarchopsis gigas Ponomarenko 1977 Karabastau Formation, Kazakhstan, Callovian
    • †Timarchopsis gobiensis Ponomarenko 1987 Formation, Mongolia, Tithonian
    • †Timarchopsis latus Ponomarenko 1977 , Kyrgyzstan, Toarcian
    • †Timarchopsis longus Ponomarenko 2014 Sharteg, Mongolia, Tithonian
    • †Timarchopsis mongolicus Ponomarenko 1985 , Mongolia, Bathonian
    • †Timarchopsis sainshandensis Ponomarenko 1987 , Mongolia, Aalenian
  • † Hong 1995 , China, Bathonian

References[]

This Adephaga-related article is a stub. You can help Wikipedia by .

• v
• t

This Jurassic insect-related article is a stub. You can help Wikipedia by .

• v
• t

This prehistoric beetle-related article is a stub. You can help Wikipedia by .

• v
• t