Frullania

Frullania; Temporal range: Cenomanian–Present PreꞒ Ꞓ O S D C P T J K Pg N
	Frullania tamarisci
Scientific classification
Kingdom:	Plantae
Division:	Marchantiophyta
Class:	Jungermanniopsida
Order:	Porellales
Suborder:
Family:	Frullaniaceae; Lorch
Genus:	Frullania; Raddi
Species
	See text

Frullania dilatata

Frullania is the only genus of liverworts in family Frullaniaceae. It contains the following species:

A

Taylor, 1846
Caspary, 1887
Gerola, 1947
Hattori, 1986
Stephani, 1916
Hodgson, 1949
Stephani, 1910
Stephani, 1911
Verdoorn, 1930
Frullania alstonii var. pfleidereri Hattori, 1972
Nees In G., L. & N., 1845
Kamimura, 1968
Stephani, 1916
Stephani, 1910
Jack & Stephani In Stephani, 1894
Ångström, 1873
Mitten, 1863
Frullania angulata F. Serratoides Vanden Berghen, 1983
Frullania angulata var. laciniata Demaret & Vanden Berghen, 1950
Stephani, 1908
Hodgson, 1949
Hattori, 1980
Horikawa, 1929
Gerola, 1947
Mitten, 1879
Frullania apicalis var. camerunensis Vanden Berghen, 1976
(R., B. & N.) Nees In G., L. & N., 1845
Frullania apiculata var. goebelii Schiffner, 1893
Frullania apiculata var. guianensis Lindenberg & Gottsche, 1851
Frullania apiculata var. laxa Nees In G., L. & N., 1845
Stephani, 1911
Hattori & Lin, 1985
Schuster, 1983
Hattori, 1972
Frullania appendistipula var. spinifera Hattori, 1974
(Sprengel) Gottsche, 1863
Frullania arecae var. spiniloba (Stephani) Yuzawa, 1991
Herzog & Clark In Clark, 1954
Verdoorn, 1932
Stephani, 1911
Frullania armitiana var. inflexula Hattori, 1988
Frullania armitiana var. longe-Attenuata (Hattori) Hattori, 1980
Stephani, 1924
Frullania asagrayana Montagne, 1842
Stephani, 1910
(Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
Frullania aterrima var. lepida Hodgson, 1949
(Swartz) Montagne, 1839
Frullania atrata var. flaccida (Weber) Nees In G., L. & N., 1845
Frullania atrata var. mexicana Nees In G., L. & N., 1845
Taylor Ex Spruce, 1884
Stephani, 1911
Hattori, 1985
Sim-Sim, Sergio, Mues & Kraut, 1995

B

Stephani, 1910
Stephani, 1916
Gottsche Ex Stephani, 1894
Frullania baladina var. edentata Hattori, 1984
Grolle, 1985
Hattori, 1977
Stephani, 1916
Stephani, 1911
Stephani In Stephani & Watts, 1914
Inoue In Hattori, 1975
Hattori, 1974
Stephani, 1894
(Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
Hattori, 1971
Spruce, 1884
Hattori, 1984
Spruce, 1884
Stephani, 1910
Austin, 1869
Reiner, 1988
Hattori, 1978
Lindenberg In G., L. & N., 1845
Massalongo, 1885
Spruce, 1889
Raddi, 1822
Frullania brasiliensis var. cylindrica (Gottsche In Lehmann) Spruce, 1884
Frullania brasiliensis var. cylindrica F. Lindigii (Gottsche) Spruce, 1884
Gottsche In G., L. & N., 1845
Stephani, 1894
Evans, 1897
Stephani, 1894
(Sprengel) Drège, 1843
Herzog, 1942
Stephani, 1910

C

Hattori, 1980
Beauverd In Stephani, 1924
Stephani, 1894
Frullania caffraria F. Anomala Vanden Berghen, 1976
Ångström, 1873
Ångström, 1876
Gottsche Ex Stephani, 1894
(Austin Ex Underwood) Evans, 1897
Sande Lacoste, 1853
Frullania campanulata var. caduca Verdoorn, 1930
Frullania campanulata var. malesiaca (Verdoorn) Hattori, 1975
Gottsche Ex Stephani, 1910
Gottsche In G., L. & N., 1845
Stephani, 1911
Kamimura & Hattori In Hattori & Kamimura, 1973
Grolle, 1985
Stephani, 1911
(Nees In Martius) Montagne, 1839
Gola, 1907
De Notaris, 1865
Hattori & Gao, 1985
Hattori & Lin, 1985
(Schuster) Schuster, 1992
Stephani, 1910
Stephani, 1887
Stephani, 1894
Gola, 1914
Beauverd In Stephani, 1924
Lindenberg & Gottsche In G., L. & N., 1847
Gottsche In G., L. & N., 1845
(Nees & Montagne) Nees In G., L. & N., 1845
(Hooker & Taylor) Taylor In G., L. & N., 1845
Stephani, 1911
Verdoorn, 1932
Gottsche Ex Stephani, 1894
Gottsche Ex Stephani, 1911
Stephani, 1911
Stephani, 1910
Stephani, 1910
Stephani, 1911
Beauverd In Stephani, 1924
Lindenberg & Hampe In Hampe, 1851
Frullania convoluta var. ampliata Herzog, 1953
Lindenberg In G., L. & N., 1845
(Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
Frullania cordistipula var. (Beta) mutica Nees In G., L. & N., 1845
Frullania cordistipula var. (Gamma) regularis Nees In G., L. & N., 1845
Frullania cordistipula var. dentistipula Hattori, 1986
Stephani, 1911
Stephani, 1910
Stephani, 1894
Jack & Stephani, 1892
Spruce Ex Stephani In Stephani, 1911
Stephani, 1894
Yuzawa & Hattori Ex Konstantinova, Potemkin & Schijakov, 1992
Hattori, 1981
Taylor, 1846
Stephani, 1924
Nees In G., L. & N., 1845
Stephani, 1911
Colenso, 1889
Stephani, 1911
Frullania curvistipula var. falcatidenta Hattori, 1982
Frullania curvistipula var. lamii Verdoorn, 1930
Frullania curvistipula var. latistipula Hattori, 1978
Stephani, 1911
Stephani, 1910
(Schwägrichen) Schwaegrichen In G., L. & N., 1845

D

Gradstein & Uribe In Uribe, 2004
Hampe In G., L. & N., 1845
F. Dorsoblastos (Hattori) Hattori & Lin, 1985
Frullania davurica F. Microphylla (Massalongo) Hattori & Lin, 1985
Frullania davurica var. chichibuensis (Kamimura) Hattori, 1976
Frullania davurica var. concava Chang In Gao & Chang, 1981
Stephani Ex Hattori, 1974
Spruce, 1884
Mitten, 1862
Arnell, 1959
Subsp. Andamana Hattori Ex Singh In Nath & Asthana, 2001
Stephani Ex Evans, 1906
Hattori, 1974
Frullania dentata var. secernens Hattori, 1988
Hattori & Streimann, 1985
Hattori, 1975
Mitten In Hooker, 1855
Gottsche Ex Lehmann, 1844
Mitten, 1863
Frullania dilatata Subsp. (Dilatata 1982) (L.) Dumortier, 1835
Frullania dilatata Subsp. (Dilatata 1982) F. Fuscovirens Jorgensen, 1934
Frullania dilatata Subsp. (Dilatata 1982) var. anomala Corbiere, 1889
Frullania dilatata Subsp. (Dilatata 1982) var. macrotus Nees, 1838
Frullania dilatata Subsp. (Dilatata 1982) var. subtilissima Nees, 1838
Frullania dilatata Subsp. Asiatica Hattori, 1982
(Lehmann & Lindenberg) Drège, 1843
Nees In G., L. & N., 1845
Stephani, 1897
Austin, 1879
Uribe, 2006
Stephani, 1894
Stephani In Dusen, 1905
Stephani, 1910
Frullania duthiana var. laevis Hattori In Ohashi, 1975
Frullania duthiana var. szechuanensis Hattori & Gao, 1985

E

(Subsp. Eboracensis 1992) Gottsche Ex Lehmann, 1844
Frullania eboracensis Subsp. Virginica (Gottsche In Lehmann) Schuster, 1992
Hattori, 1974
Hattori, 1988
(Sprengel) Sprengel In G., L. & N., 1845
Frullania ecklonii F. Robustior Gottsche, 1863
Frullania ecklonii F. Tenerior Sprengel, 1847
Frullania ecklonii var. huitamalcensis Gottsche, 1863
Frullania ecklonii var. rufescens Sprengel, 1847
Stephani, 1911
Lehmann, 1857
Hattori, 1977
Hattori, 1983
Frullania epiphylla Subsp. Fijiensis Hattori, 1985
Stephani, 1911
(Nees In Martius) Montagne, 1839
Frullania ericoides F. Ericoides (Nees) Verdoorn Ex Piippo, 1990
Frullania ericoides var. planescens (Verdoorn) Hattori, 1984
Hattori, 1974
Verdoorn, 1930
Frullania errans var. angulistipula Hattori, 1972
Beauverd Ex Stephani, 1924
Hattori & Thaithong, 1978
Mitten, 1861
Stephani, 1897
Hattori, 1975
Frullania eymae var. crispidentata Hattori & Streimann, 1985

F

Taylor Ex Lehmann, 1844
Gottsche In G., L. & N., 1845
Hattori, 1986
Hattori & Piippo, 1986
Frullania falsisinuata F. Parvistylata (Hattori) Hattori & Piippo, 1986
Frullania falsisinuata var. crispidentata Hattori & Piippo, 1986
Stephani, 1894
Frullania fauriana F. Emarginata Kamimura, 1952
Zhu & So, 1997
Stephani, 1910
Frullania fertilis var. major Massalongo, 1885
Taylor Ex Spruce, 1884
Spruce, 1884
Hattori, 1983
(Taylor) Taylor In G., L. & N., 1845
Howe, 1894
Stephani, 1910
(Hooker & Taylor) Taylor In G., L. & N., 1845
Hattori, 1987
Stephani, 1910
Frullania fusco-Virens var. gemmipara (Schuster & Hattori In Hattori) Hattori & Lin, 1985

G

Vanden Berghen, 1976
Bai & Gao, 1999
(Nees & Montagne) Nees & Montagne In G., L. & N., 1845
Frullania gaudichaudii F. Hasseltii (Sande Lacoste) Verdoorn, 1930
Hattori & Thaithong In Hattori, Thaithong & Kitagawa, 1977
(Nees) Nees In Montagne, 1840
Frullania gibbosa var. major Gottsche, 1863
Stephani, 1911
Massalongo, 1897
Frullania giraldiana var. handelii (Verdoorn) Hattori, 1972
Hattori & Streimann, 1985
(Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg Ex Montagne, 1838
Hattori, 1977
Subsp. (Gracilis 1978) (Reinwardt, Blume & Nees) Gottsche In G., L. & N., 1845
Frullania gracilis Subsp. (Gracilis 1978) var. brevior Nees In G., L. & N., 1845
Frullania gracilis Subsp. (Gracilis 1978) var. vittata Hattori, 1986
Frullania gracilis Subsp. Zennoskei Hattori & Thaithong, 1978
Yuzawa, Mues & Hattori, 1987
Gottsche, 1864
Stephani, 1911
Hattori & Piippo, 1986
Lindenberg In G., L. & N., 1845
Gottsche In G., L. & N., 1846
Hattori, 1972
Stephani, 1910
Stephani, 1911
Gottsche Ex Stephani, 1911
Stephani, 1911
Stephani, 1911

H

Lindenberg In G., L. & N., 1846
Spruce, 1884
Hattori & Lin, 1986
Stephani, 1911
Stephani, 1889
Stephani, 1910
Frullania hamatiloba F. Grosse-Appendiculata Hattori, 1944
Frullania hamatiloba var. latistipula Hattori, 1975
Grolle In Grolle & Meister, 2004
Herzog & Clark In Clark & Schultz, 1953
Verdoorn In Handel-Mazzetti, 1930
Hattori, 1981
Stephani, 1911
Herzog, 1942
Lindenberg In G., L. & N., 1845
Frullania hasskarliana var. gracilis Hattori, 1986
Frullania hasskarliana var. integribractea (Verdoorn) Hattori, 1975
Frullania hasskarliana var. parvidentata Hattori, 1986
Godfrey & Godfrey, 1980
Udar & Nath, 1981
Konrat & Braggins, 2003
Miller, 1953
Stephani, 1911
Hattori & Kamimura, 1973
Stephani, 1910
Stephani, 1911
Hattori, 1955
Schiffner, 1889
Hattori, 1984
Frullania hicksiae F. Litoralis Hattori, 1988
Yuzawa & Hattori In Yuzawa & Koike, 1994
Stephani, 1910
Kamimura, 1982
Hattori, 1980
Spruce, 1884
Hattori & Griffin Iii, 1978
Hattori, 1976
Stephani In Stephani & Watts, 1914
Hattori, 1976
Frullania huerlimannii var. dioica Hattori, 1984
Vanden Berghen, 1976
Spruce, 1890
Nees In G., L. & N., 1843
Hattori, 1984

I

Stephani, 1911
Stephani, 1896
Hattori In Mizutani, 1979
Frullania incisoduthiana var. parva Hattori In Mizutani, 1979
Stephani, 1924
Verdoorn, 1930
Frullania inconstans F. Integrior Verdoorn, 1930
Frullania inconstans var. grossedentata Kamimura & Hattori, 1973
Mitten In Hooker, 1855
Hattori, 1988
Gottsche In G., L. & N., 1845
Frullania inflata var. communis Schuster, 1985
Frullania inflata var. dioica Hattori & Thaithong, 1978
Mitten, 1861
Hattori, 1984
Hattori, 1980
(Nees) Nees In G., L. & N., 1845
Frullania integristipula var. emarginata Verdoorn, 1929
Frullania intermedia Subsp. (Intermedia 1980) F. Billardieriana (Nees & Montagne In Montagne) Verdoorn, 1930
Frullania intermedia Subsp. (Intermedia 1980) var. non-Apiculata Hattori, 1975
Frullania intermedia Subsp. (Intermedia) (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
Frullania intermedia Subsp. Morokensis (Stephani) Hattori, 1980
Colenso, 1889
(Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
Hampe Ex Stephani, 1911
Hattori & Kamimura, 1973
Hattori, 1980
Hattori & Piippo, 1986
Hattori, 1972

J

Subsp. (Jackii 1959) Gottsche In Gottsche & Rabenhorst, 1863
Frullania jackii Subsp. (Jackii 1959) F. Depauperata Grolle, 1970
Hattori, 1986
Gottsche Ex Stephani, 1910
Loitlesberger, 1894
Stephani, 1894
Gottsche In G., L. & N., 1845
Frullania junghuhniana var. bisexualis Hattori, 1976
Frullania junghuhniana var. tenella (Sande Lacoste) Grolle & Hattori In Hattori, 1982
Frullania junghuhniana var. tenella F. Monoica (Hattori) Hattori, 1986

K

Subsp. (Kagoshimensis 1985) Stephani, 1910
Frullania kagoshimensis Subsp. (Kagoshimensis 1985) var. minor Kamimura, 1961
Frullania kagoshimensis Subsp. Hunanensis (Hattori) Hattori, 1985
Hattori, 1986
Piippo & Hattori In Piippo & Tan, 1992
Verdoorn, 1932
Hattori, 1984
Nees Ex Stephani, 1911
Hattori, 1978
(Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
Frullania kunzei var. maritima Schuster, 1991

L

Hampe In G., L. & N., 1845
Bai & Gao, 2000
Stephani, 1910
Singh & Singh, 2005
Spruce, 1890
Spruce, 1884
Herzog, 1948
Spruce, 1884
Frullania laxiflora var. crossii Spruce, 1884
Austin, 1869
Verdoorn, 1930
Hattori & Piippo, 1986
Vanden Berghen, 1976
Stephani, 1910
Nees In G., L. & N., 1845
Gottsche, 1863
Gottsche Ex Lehmann, 1844
Frullania lindenbergii var. (Beta ?) Fusca Gottsche In G., L. & N., 1847
Stephani, 1911
Stephani, 1897
Hattori, 1981
Stephani, 1911
(Hooker) Hooker & Nees In G., L. & N., 1845
Stephani, 1910
Stephani In Renauld & Cardot, 1891
Frullania longistipula var. apiculata Demaret & Vanden Berghen, 1950
Yuzawa & Hattori, 1988
Pearson, 1891
Frullania loricata var. laxa Pearson, 1891
Stephani, 1908
Hattori & Lin, 1985

M

Stephani, 1894
Frullania macgregorii var. rostellula (Hattori) Hattori, 1982
(Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
Hattori, 1980
Taylor, 1846
Gottsche, 1882
Stephani, 1924
Spruce, 1884
(Sprengel) Weber & Nees In G., L. & N., 1845
Frullania magellanica var. diminutiva Herzog, 1938
Grolle, 2003
Hattori, 1977
Miller, 1981
Stephani, 1911
Austin, 1869
Svihla, 1958
Hattori, 1973
(Hodgson) Hattori, 1983
Spruce, 1884
Hattori, 1974
Stephani, 1911
Lindenberg In G., L. & N., 1845
Frullania meyeniana var. dioica Hattori, 1977
Verdoorn, 1929
Frullania microauriculata var. rotundior Verdoorn, 1929
Gola, 1923
Gottsche, 1863
(Gottsche In Gottsche & Rabenhorst) Pearson, 1894
Frullania microphylla F. Corticicola Schiffner, 1936
Frullania microphylla var. deciduifolia Grolle, 1970
Clark & Svihla, 1950
Pearson, 1922
Subsp. (Minor 1975) var. integribracteola Hattori, 1975
Frullania minor Subsp. Recurviloba Hattori, 1975
Jack & Stephani In Stephani, 1892
Lindenberg & Gottsche In G., L. & N., 1847
Kamimura & Hattori, 1973
Subsp. Breviramea (Stephani) Verdoorn, 1930
Frullania moniliata Subsp. Moniliata Crandall-Stotler , Geissler & Stotler, 1987
Frullania moniliata Subsp. Moniliata F. Minshanensis (Hattori) Crandall-Stotler Et Alii, 1987
Frullania moniliata Subsp. Moniliata var. elongatistipula (Verdoorn) Crandall-Stotler Et Alii, 1987
Frullania moniliata Subsp. Moniliata var. vietnamica (Hattori) Crandall-Stotler Et Alii, 1987
(Hooker & Taylor) Taylor In G., L. & N., 1845
Frullania monocera var. depauperata Hattori, 1984
Frullania monocera var. schiffneri (Verdoorn) Hattori, 1979
Stephani, 1900
Gottsche In G., L. & N., 1845
Stephani, 1910
Lindenberg & Gottsche In G., L. & N, 1847
Frullania moritziana var. (Beta) Mexicana Gottsche, 1863
Hattori & Streimann, 1985
Stephani, 1911
(Lehmann & Lindenberg In Lehmann) Lehmann & Linde,nberg
Frullania mucronata var. submutica Lehmann & Lindenberg In G., L. & N. 1845 In G., L. & N., 1845
Subsp. (Multilacera 1987) Stephani, 1911
Frullania multilacera Subsp. (Multilacera 1987) var. lacerissima Hattori, 1975
Frullania multilacera Subsp. Gracilior Hattori, 1987
Hattori, 1987
Stephani, 1894
Stephani, 1911

N

Stephani, 1911
Engel In Engel & Merrill, 1999
Hattori, 1979
(Sprengel) Lehmann & Lindenberg In G., L. & N., 1845
Taylor, 1846
Hodgson, 1949
(R., B. & N.) Nees In G., L. & N., 1845
Frullania nigricaulis var. elongata Verdoorn, 1930
Frullania nisquallensis Sullivant, 1849
Hattori, 1982
Stephani, 1894
Frullania nobilis var. cochleata (Stephani) Hattori In Hattori & Streimann, 1985
(Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
Stephani, 1911
Hattori, 1981
Schiffner, 1890

O

Hampe In G., L. & N., 1843
Frullania oakesiana Subsp. (Oakesiana 1992) Austin, 1870
Frullania oakesiana Subsp. Takayuensis (Stephani) Schuster, 1992
Hattori, 1982
Mitten, 1879
Stephani, 1924
Kamimura & Hattori, 1973
Spruce, 1890
Kamimura, 1982
i Vanden Berghen, 1976
Austin, 1869
Sande Lacoste, 1855
Spruce, 1883
(Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
Frullania ornithocephala F. Magnilobula Hattori, 1976
Frullania ornithocephala F. Retusa Hattori, 1976
Frullania ornithocephala F. Teres (Sande Lacoste) Verdoorn, 1929
Frullania ornithocephala var. maior Nees, 1845
Frullania ornithocephala var. minor Nees, 1845
Frullania ornithocephala var. pilosa Verdoorn, 1929
Frullania ornithocephala var. tuberculosa Hattori, 1974
Notaris, 1855
(Hattori) Hattori In Iwatsuki & Hattori, 1956
Frullania osumiensis var. dentata Kamimura, 1970

P

Hattori, 1978
Stephani, 1911
Hattori, 1988
Gottsche Ex Stephani, 1894
Stephani, 1911
Hattori, 1975
Verdoorn, 1930
Stephani, 1911
Lehmann & Lindenberg In G., L. & N., 1845
Frullania paradoxa F. Cornistipula Herzog, 1952
(Schuster) Schuster, 1992
Hattori & Thaithong, 1978
Frullania pariharii F. Intermedia Hattori & Thaithong, 1978
Stephani, 1910
Stephani, 1910
Stephani, 1910
Mitten In Hooker, 1855
Stephani, 1911
Frullania pauciramea var. pauciramella Hattori & Piippo, 1986
Hattori & Piippo, 1986
Stephani, 1911
Stephani, 1897
Frullania pentapleura Subsp. (Pentapleura 1988) Taylor, 1846
Gottsche In G., L. & N., 1846
Stephani, 1916
Stephani, 1911
Mitten, 1861
Hattori, 1977
Stephani, 1911
Frullania piliflora var. appendiculata Herzog, 1951
Stephani, 1911
Hattori, 1980
Frullania piptophylla var. minor Hattori, 1986
Hattori, 1980
I Stephani, 1892
Sullivant, 1849
Stephani, 1910
Herzog, 1952
Colenso, 1889
Gottsche, 1864
Thaithong & Hattori, 1977
Colenso, 1889
Taylor, 1846
Frullania polyptera var. angustata (Mitten) Hattori, 1975
Frullania polysticta Lindenberg In G., L. & N., 1845
Hattori & Koike In Koike, 1994
Fulford & Sharp, 1990
Taylor, 1846
Hattori & Streimann, 1985
Hattori & Streimann, 1985
Hattori, 1982
Hattori, 1986
Hattori, 1986
Hattori, 1986
Hattori, 1980
Frullania pseudoschensiana var. darjeelingensis Hattori, 1981
Montagne, 1843
Herzog, 1954
Verdoorn, 1930
Stephani, 1911
Reimers, 1931
Stephani, 1911
Mitten, 1871
(Hooker & Taylor) Taylor In G., L. & N., 1845
Grolle In Grolle & Meister, 2004
Stephani, 1910

Q

Stephani, 1910
(Nees & Montagne) Nees & Montagne In Montagne In Alcide D'orbigny, 1839
Frullania quillotensis F. Flaccida Herzog, 1954
Frullania quillotensis F. Leiantha Herzog, 1954

R

Stephani, 1911
(Nees) Montagne, 1842
Hattori, 1975
Sande Lacoste, 1853
Frullania reflexistipula var. novocaledonica Hattori, 1977
Frullania reflexistipula var. squarrosa Hattori & Piippo, 1986
Schiffner, 1890
Stephani, 1910
Verdoorn, 1930
Hattori, 1872
Stephani, 1911
Stephani, 1894
Gottsche, 1864
Subsp. (Repandistipula 1975) Sande Lacoste, 1853
Frullania repandistipula Subsp. Queenslandica Hattori, 1987
Frullania repandistipula Subsp. Spinibractea Hattori, 1975
Frullania reptans var. integristipula Nicholson, 1925
Mitten, 1861
Frullania retusa var. gymnantha Hattori & Thaithong, 1978
Frullania retusa var. hirsuta Hattori & Thaithong, 1978
Herzog Ex Verdoorn In Handel-Mazzetti, 1930
Hattori, 1980
Spruce, 1884
Stephani, 1910
Spruce, 1884
(Raddi) Ångström, 1876
Hampe Ex Lehmann, 1838
Piippo & Hattori, 1992
Mitten In Hooker, 1867
(Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
Gottsche Ex Stephani, 1889
Frullania rubella var. elongata (Stephani) Hattori, 1983
Hattori, 1972
Stephani, 1924

S

Hattori, 1976
Schuster, 1983
Gottsche, 1864
Stephani, 1911
Stephani, 1897
var. minor Kamimura, 1982
Hattori & Koike In Koike, 1994
Hattori, 1976
Hattori, 1977
Montagne, 1843
Yuzawa & Hattori, 1989
Massalongo, 1897
Verdoorn, 1929
Nees In G., L. & N., 1845
Frullania schimperi var. laciniata Vanden Berghen, 1976
(Caspary) Grolle, 1981
Hattori, 1988
Hattori, 1972
Hattori, 1987
Frullania sebastianopolitana var. (Delta) Gottsche, 1864
Pearson, 1890
Gola, 1914
Lindenberg & Gottsche In G., L. & N., 1847
var. (Beta) Glabra Lindenberg & Gottsche In G., L. & N., 1847
Sim-Sim, Fontihna, Mues & Lion, 2000
Gottsche Ex Stephani, 1889
Stephani, 1894
Subsp. (Serrata 1982) Gottsche In G., L. & N., 1845
Frullania serrata Subsp. (Serrata 1982) F. Crispulodentata Verdoorn, 1929
Frullania serrata Subsp. (Serrata 1982) F. Vogelkopensis Hattori, 1982
Frullania serrata Subsp. (Serrata 1982) var. ceramensis Hattori, 1986
Frullania serrata Subsp. (Serrata 1982) var. hamatispina (Hattori) Hattori, 1982
Frullania serrata Subsp. (Serrata 1982) var. oceanica Verdoorn, 1937
Frullania serrata Subsp. (Serrata 1982) var. pertenuis (Nees) Gottsche In G., L. & N., 1845
Frullania serrata Subsp. Grolleana Grolle & Piippo, 1984
Frullania serrata Subsp. Spinistipula (Hattori) Hattori, 1982
Hattori, 1988
Stephani, 1894
Svihla, 1957
Udar & Kumar, 1983
Hattori, 1974
Hattori, 1979
Stephani In Stephani & Watts, 1914
Stephani In Levier, 1906
Hattori & Lin, 1985
Subsp. Sinuata 1975 Sande Lacoste, 1853
Frullania sinuata Subsp. Sinuata F. Parvistylata Hattori In Grolle & Piippo, 1984
Mitten In Balfour, 1888
Colenso, 1889
Stephani, 1910
Herzog, 1949
Reiner, 1988
Hattori, 1980
Spruce, 1884
Lin In Lin & Chen, 1997
(Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
Taylor, 1846
Hattori, 1979
Hattori, 1972
Stephani, 1911
Stephani, 1894
Spruce, 1884
(Hooker & Taylor) Taylor In G., L. & N., 1845
Stephani, 1910
Hattori, 1987
Spruce, 1884
Hattori, 1983
(Schuster) Schuster, 1992
Hattori, 1974
Stephani, 1910
Stephani, 1911
Frullania subdentata F. Longistipula Hattori, 1978
Massalongo In Levier, 1906
Hodgson, 1949
Hattori, 1987
Stephani, 1894
Hattori & Koike In Koike, 1994
Hattori, 1973
Frullania subnigricaulis var. subtruncata Hattori, 1975
Hattori, 1986
Hattori, 1980
Hattori, 1977
Herzog, 1954
Hattori, 1972
Hattori, 1972
Lindenberg In G., L. & N., 1845
Stephani, 1910
Stephani, 1910
Hattori & Thaithong, 1978
(Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
Hattori, 1983

T

(Hattori, Thaithong & Kitawana) Hattori In Hattori & Lin, 1985
Horikawa, 1934
Frullania tamarisci (L.) Dumortier, 1835
Frullania tamarisci var. atlantica Schiffner, 1936
Frullania tamarisci var. atrovirens Carrington, 1863
Frullania tamarisci var. commutata Nees In G., L. & N., 1845
Frullania tamarisci var. cornubica Carrington In Carrington & Pearson, 1878
Frullania tamarisci var. laxa Nees In G., L. & N., 1845
Frullania tamarisci var. mediterranea (De Notaris) Nees In G., L. & N., 1845
Frullania tamarisci var. robusta Lindberg, 1875
Frullania tamarisci var. sardoa (De Notaris) De Notaris, 1865
Frullania tamarisci var. schiffneri Nicholson, 1933
Stephani, 1910
Stephani, 1910
Schuster, 1983
(Weber) Nees, 1838
Frullania teneriffae F. Explanata (Macvicar) Bisang, Schumacker, Sérgio & Grolle, 1988
Frullania teneriffae var. germana (Taylor) Bisang, Schumacker, Sérgio & Grolle, 1988
Stephani, 1911
Gottsche In G., L. & N., 1846
Frullania ternatensis var. non-Appendiculata Hattori, 1974
Nees & Montagne In Montagne, 1838
Hattori, 1988
Hattori, 1976
Hattori & Thaithong, 1977
Hattori, 1975
Colenso, 1889
Gottsche, 1864
Sande Lacoste, 1856
Mitten, 1862
Clark, Jovet-Ast & Frye, 1947
(Lehmann & Lindenberg) Drège, 1843
Lindenberg & Gottsche In G., L. & N., 1847
Herzog, 1942
Caspary, 1887
Hattori & Lin, 1985
Clark & Frye, 1952
Lindenberg & Gottsche In G., L. & N., 1847
Hattori & Thaithong, 1978

U

Nath & Singh, 2006
Mitten Ex Stephani, 1911
Kamimura, 1961
Schiffner Ex Stephani, 1894
Frullania usambarana var. reducta Vanden Berghen, 1976
Stephani, 1897
Stephani, 1894

V

Mitten In Seemann, 1871
(Swartz) Nees In G., L. & N., 1846
Jack & Stephani In Stephani, 1894
Stephani, 1910
Lehmann, 1857
Pocs In Bizot & Pocs, 1979
Kamimura & Hattori, 1973
Stephani, 1910
Caspary, 1887
Stephani, 1894
Hattori, 1974
Hattori, 1973
Stephani, 1910
Yuzawa & Hattori, 1988
Hattori, 1974
F. Denticulata Hattori, 1980
Hattori, 1987

W

Stephani, 1910
Konrat & Braggins, 2005
Hattori & Lin, 1985
Hattori, 1974
Frullania warnckeana var. dentosa Hattori, 1977
Stephani, 1911
Stephani, 1910
Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa, 1991

Y

Stephani, 1911
Stephani, 1894
Frullania yuennanensis var. siamensis (Hatt. et al.) Hattori & Lin, 1985
Kamimura, 1970
Hattori, 1981

Z

Hattori & Lin, 1985
Hattori, 1984

Extinct species[]

†Frullania baerlocheri Heinrichs et al., 2012 Burmese amber, Myanmar, Cenomanian
†Frullania cretacea Hentschel et al., 2009 Burmese amber, Myanmar, Cenomanian
†Frullania partita Li et al, 2020 Burmese amber, Myanmar, Cenomanian
†Frullania pinnata Heinrichs et al., 2017 Burmese amber, Myanmar, Cenomanian

References[]

Media related to Frullania at Wikimedia Commons

Taxon identifiers
Frullania	Wikidata: Q3753784 Wikispecies: Frullania APNI: 120367 BioLib: 91537 EoL: 36257 EPPO: 1FKBG GBIF: 2688736 iNaturalist: 56519 IRMNG: 1037849 ITIS: 15200 NBN: NHMSYS0000309901 NCBI: 95763 NZOR: ddcb9edb-4f6f-48a8-9397-520b172b8eba PLANTS: FRULL Tropicos: 35002031 WFO: wfo-4000015089
Frullaniaceae	Wikidata: Q22809340 Wikispecies: Frullaniaceae APNI: 120771 FloraBase: 22953 GBIF: 2269 IRMNG: 104573 ITIS: 14188 NBN: NHMSYS0000309909 NCBI: 400721 NZOR: 2050ebcf-1055-4a31-a56c-91f3a53d301c Tropicos: 35002634 WFO: wfo-7000000739

Frullania Temporal range: Cenomanian–Present PreꞒ Ꞓ O S D C P T J K Pg N

Frullania tamarisci
Scientific classification
Kingdom:	Plantae
Division:	Marchantiophyta
Class:	Jungermanniopsida
Order:	Porellales
Suborder:
Family:	Frullaniaceae Lorch
Genus:	Frullania Raddi
Species
See text