List of the Paleozoic life of Pennsylvania

This list of the Paleozoic life of Pennsylvania contains the various prehistoric life-forms whose fossilized remains have been reported from within the US state of Pennsylvania and are between 541 and 252.17 million years of age.

A[]

†
- †
†Acidaspis
- †
†
- †
- †
†
†
- † – type locality for species
†
- †
- †
- †
†
- †
†
- †
- † – type locality for species
- † – type locality for species
†
- † – type locality for species
- † – type locality for species
†
†
†Ageleodus
- †
†
- †
- †
†Agnostus
†
- † – type locality for species
- † – type locality for species
Fossilized fronds of the Carboniferous-Early Cretaceous seed fern Alethopteris
†Alethopteris
- †
- †
- †
- †
- †
- †
†
- †
†
- † – type locality for species
- † – type locality for species
†
- †
- †
† – type locality for genus
- † – type locality for species
†
†Amphiscapha
- †
†Amphissites
†
†Ancyrospora
†
- †
†
†
- †
- † – or unidentified comparable form
- †
Restorative model of the Cambrian arthropod Anomalocaris
†Anomalocaris
- †
† – type locality for genus
- † – type locality for species
†
- †
†
†
†
- †
†Aphthoroblattina
- † – type locality for species
†
- †
†
- † – type locality for species
†Archaeopteris
- †Archaeopteris halliana
- †
- Life restoration of the Late Devonian-Carboniferous tree Archaeopteris macilenta with insets detailing its anatomy
  †Archaeopteris macilenta
- †
- † – or unidentified comparable form
- †
†
- †
†
- †
†
- †
†Archimylacris
- †
†
†
- †
†
†
Fossil of the Cambrian trilobite Asaphiscus
†Asaphiscus
- †
† – type locality for genus
- † – type locality for species
†
- †
† – type locality for genus
- † – type locality for species
†
- †
†Athyris
- †
- †
- †Athyris spiriferoides
† – type locality for genus
- † – type locality for species
- † – type locality for species
- † – type locality for species
†
- †
†Atrypa
- †
- Fossilized shell of the Silurian-Late Devonian brachiopod Atrypa reticularis
  †Atrypa reticularis
†
†
- †
- †
- †
- †
†Aviculopecten
- †
- †
- †Aviculopecten occidentalis

B[]

†
†Barbclabornia
- †
†Barinophyton
- †
- †
†Bathyuriscus
†
Fossilized shell of the Silurian-Early Triassic mollusc Bellerophon
†Bellerophon
- †
- †
†Bembexia
- †
†Beyrichoceratoides
- †
†
- †
†
- † – type locality for species
†
- † – type locality for species
†Blattoidea
- † – type locality for species
†
- † – type locality for species
- †
- † – type locality for species
†
- †
- †
- †
- †
- †
†
- †
†Bonneterrina
- † – type locality for species
†
- †
- †
- † – tentative report
†
- †
Fossilized bony armor of the Late Devonian placoderm fish Bothriolepis

Life restoration of the Late Devonian placoderm fish Bothriolepis
†Bothriolepis
- †
- †
†
- †
†
- † – type locality for species
- † – type locality for species
- † – type locality for species
†
†
- †
†
- †
- †
†
- †
†
- †
- †
†
- †
†
- †
- † – informal
†
- †
†Byronia
† – tentative report
- † – or unidentified comparable form

C[]

†
- † – type locality for species
†
- †
- †
- †
†Callixylon
† – type locality for genus
- † – type locality for species
†
- †
†
- †
†Camarotoechia
- †
- †
- †
†
Life restoration of the Cambrian echinoderm Camptostroma
†Camptostroma
- †
†
- † – type locality for species
†
- †
- †
†
- †
†
- †
†Cavusgnathus
†
†Ceramopora
- †
†Ceratopsis
- †
†
† – type locality for genus
- † – type locality for species
†Chancelloria
- †
†Chancia
†
- † – type locality for species
†Chonetes
- †
- †
†
- †
†
Assemblage of fossilized shells of the Ordovician brachiopod Cincinnetina
†Cincinnetina
- †Cincinnetina multisecta
†
- †
†Cleiothyridina
†
- †
- †
†Climacograptus
- †
† – type locality for genus
- † – type locality for species
Coccosteus cuspidatus restoration, shown attacking multiple Palaeospondylus'
†Coccosteus
- † – type locality for species
†Coelacanthus
†
†
- †
†
- †
†
- †
†Composita
- †Composita subtilita
†
†Conotheca
- † – or unidentified comparable form
†
- † – tentative report
†
- † – or unidentified comparable form
†Coosella
- † – type locality for species
- † – type locality for species
†Coosia
- † – type locality for species
†
- †
Fossilized foliage of the Carboniferous-Permian conifer relative Cordaites
†Cordaites
†
- †
†Cornulites
- †Cornulites flexuosus
- † – or unidentified comparable form
† – type locality for genus
- † – type locality for species
†
†Craniops
- †
- †
†
- †
†Crepicephalus
†Crurithyris
- †
†Cryptolithus
- †
- †
†
- †
†Ctenerpeton – type locality for genus
- † – type locality for species
†
- †
Fossils and life restoration of the Silurian eurypterid ("sea scorpion") Ctenopterus
†Ctenopterus
- †
†
†
- †
- †
- †
- †
†Cyclonema
†Cyclopteris
†
- †
†
- †
- †
- †
†
- †
†Cypricardinia
- †Cypricardinia indenta
†
- †
Fossilized shells of the Middle-Late Devonian brachiopod Cyrtospirifer
†Cyrtospirifer
- †
†
- † – or unidentified comparable form
†

D[]

†
- †
†
- †
†
†Deckera
- †
†
†
- †
- † – type locality for species
- †
†
†
- †
†Densignathus – type locality for genus
- † – type locality for species
†
- †
- †
†
- †
- †
- †
†
†
†Diaphragmus
†
†
- †
†
- †
† – type locality for genus
- † – type locality for species
†Diplocaulus – or unidentified comparable form
Life restoration of the Permian amphibian Diploceraspis
†Diploceraspis
- †Diploceraspis burkei
†Diplograptus
- †
†Ditomopyge
- †
†
†
- †
† – tentative report
- †
†
- †
- †
†
- †
- †
- †
- †
- †
†
- †

E[]

†
- †
†
- †
Life restoration of the Permian synapsid (mammal precursor) Edaphosaurus
†Edaphosaurus
- † – type locality for species
†
†Eldredgeops
- †Eldredgeops rana
†
- †
†Elrathia
- † – type locality for species
†Elrathina
†
- †
†
- †
- †
- †
†Emmanuella
- †
†
- † – or unidentified comparable form
- † – or unidentified comparable form
- †
†Endolobus
†Endothyra
†
- † – or unidentified comparable form
- † – or unidentified comparable form
†
- †
†Eoagnostus
- †
†
- †
†
- †
†
- † – type locality for species
†
- †
Life restoration of the Silurian-Early Devonian eurypterid ("sea scorpion") Erettopterus
†Erettopterus
- †
†Eryops
- †
†
†Esmeraldina
- †
†
- †
†
†
†Euomphalus
†
†Euphemites
- †

F[]

† – type locality for genus
- † – type locality for species
Fossil of the Late Ordovician-Permian tabulate coral Favosites
†Favosites
- †Favosites niagarensis
†Fedexia
†Fenestella
- †
- †
†
†
- † – or unidentified comparable form
†
- †
†
- † – or unidentified comparable form
†
- †

G[]

†Ganorhynchus
- † – type locality for species
†
Fossil of the Cambrian trilobite Genevievella
†Genevievella
- † – type locality for species
- † – type locality for species
- † – type locality for species
† – type locality for genus
- † – type locality for species
†
- †
- † – type locality for species
†
- †
†
- †
†Glaukerpeton – type locality for genus
- † – type locality for species
†Glyptocrinus
†
- † – tentative report
†Gnathodus
† – type locality for genus
- † – type locality for species
†Goniophora
- †
†
- †
- †
Fossils of the Middle Ordovician-Early Devonian trilobite Gravicalymene
†Gravicalymene
†Greenops
- †

H[]

†
Life restoration of the Late Devonian eurypterid ("sea scorpion") Hallipterus
†Hallipterus
- †
† – type locality for genus
- † – type locality for species
†Hardieopterus
- †
†Hazelia
- †
†
†Helcionella
†
- † – type locality for species
†
- † – type locality for species
†
†
- †
†Hexactinellid
- †
†
†
- †
†Hindeodus
†
- †
†
- † – or unidentified comparable form
- †
Life restoration of the Middle-Late Devonian placoderm fish Holonema
†Holonema
- † – type locality for species
- †
†Holopea
†Holoptychius
- †
- † – type locality for species
- †
†Holoptychus
- † – type locality for species
- † – type locality for species
- † – type locality for species
†
- † – type locality for species
†
- †
†
- †
†
- †
†
- † – or unidentified comparable form
†Hughmilleria
- †Hughmilleria shawangunk
†Hymenocaris
†
- †
- †
Life restoration of the Late Devonian four-limbed animal precursor Hynerpeton
†Hynerpeton – type locality for genus
- †Hynerpeton bassetti – type locality for species
†Hyolithellus
- †
†Hyolithes
- †
†Hyolithid
†
- †
†
- †
- ��
†
- †

I[]

†
†
- †
† – type locality for genus
- † – type locality for species
†
- † – type locality for species
†Indiana – tentative report
†
†
†Iocrinus
- †
†Irvingella
- †
Life restoration of the Permian amphibian Isodectes
†Isodectes
- † – type locality for species
†Isotelus
- †Isotelus gigas
†

K[]

†
- †
Fossilized heads of the Silurian eurypterid ("sea scorpion") Kiaeropterus (specimens 1-8)
†Kiaeropterus
- †
†
- †
†Kingstonia
- †
- †
- † – type locality for species
- † – type locality for species
- † – type locality for species
- †
†
- † – type locality for species
† – tentative report
†
- †
† – type locality for genus
- † – type locality for species
†Kladognathus
†
†
- †
†
Fossil of the Cambrian trilobite Kootenia
†Kootenia
- †
†
†Kutorgina
- †
- †
†

L[]

Life restoration of the Carboniferous tree-sized club moss relative Lepidodendron

†
- † – type locality for species
†
- †
†
†Lancastria
†
- †
- † – or unidentified comparable form
†Leiorhynchus
†
†
†
†
- †
†
- †
- †
- †
- †
†Lepidostrobus
- †
†Leptaena
- †
- † – or unidentified comparable form
†
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
- †
†Leptomitus
†
Lingula
- †
- †
- †
Illustration of a fossilized shell of the Cambrian-Late Ordovician brachiopod Lingulella
†Lingulella
- † – or unidentified comparable form
†Lingulepis
- †
†Linoproductus
- †
†
- †
†Llanoaspidella – type locality for genus
- † – type locality for species
- † – type locality for species
†Llanoaspis – tentative report
†Lonchocephalus
- † – type locality for species
- † – type locality for species
- † – type locality for species
†
- †
†
- †
†
†
- †
- †
†
Life restoration of the Permian snake-like amphibian Lysorophus showing speculative egg-coiling behavior
†Lysorophus
- †

M[]

† – tentative report
- †
†
- †
†Margaretia
†
†Martinia
†
- †
- †
- †
- †
†
- †
Life restoration of the Late Ordovician eurypterid ("sea scorpion") Megalograptus
†Megalograptus – tentative report
†Megamolgophis
- †
†
†
- †
†
- † – type locality for species
†
†Metacoceras
- †
† – type locality for genus
- † – type locality for species
† – type locality for genus
- † – type locality for species
†Michelinoceras
†Micromitra
- †
†Mimagoniatites – tentative report
†
- †
†
†
- †
†
- † – type locality for species
†
- †
Fossilized shell of the Devonian brachiopod Mucrospirifer
†Mucrospirifer
- †Mucrospirifer mucronatus
†
- †
†
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species

N[]

†
- †
†Nanahughmilleria
- †
†
Fossilized shell of the Early Devonian – Triassic sea snail Naticopsis
†Naticopsis
†
- † – type locality for species
- † – type locality for species
† – type locality for genus
- † – type locality for species
†
- †
†
- † – type locality for species
- † – type locality for species
†
†
† – type locality for genus
- † – type locality for species
†Neospirifer
- †Neospirifer cameratus
†
- †
†
- †
Fronds of the Carboniferous seed fern Neuropteris
†Neuropteris
†Nisusia
- †
Nucula
- †
- †
Nuculana
†
- †
- †
- †
†
- †

O[]

†
- †
- †
- †
†Ogygopsis
- †
†Olenellus
- †
- †
- †
Fossil of the Cambrian trilobite Olenoides
†Olenoides
- † – type locality for species
- † – type locality for species
- †Olenoides serratus
†
- † – type locality for species
†
- †
† – type locality for genus
- † – type locality for species
Life restoration of the Carboniferous-Permian freshwater shark Orthacanthus
†Orthacanthus
- †
- †
- †
†Orthoceras
†
- †
†
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
- † – type locality for species
†
- † – tentative report
- †
†
- †
†
†Orthotheca
- † – type locality for species
†Oryctocephalus
Restoration
†Osteolepis
†Oulodus
†
†
- † – type locality for species
†Ozarkodina

P[]

†Paedeumias
- †
- †
- †
Fossil of the Cambrian trilobite Pagetia
†Pagetia
†Paladin
†
- †
- †
- †
- †
- †
- †
†
- † – type locality for species
†
- †
†Panenka
†
- †
- †
†
- †
†
†
- †
†
- † – type locality for species
†Parahughmilleria
- †
†
- †
†
- †
†
Assemblage of the Early-Middle Devonian brachiopod Paraspirifer
†Paraspirifer
- †
†
- † – or unidentified comparable form
†Paterina
- †
†Pelagiella
†Pemphigaspis
- †
- † – type locality for species
- †
†
Fossilized theca of the Carboniferous blastoid echinoderm ("sea bud") Pentremites
†Pentremites
†
- †
- †
†Peronopsis
†Petalodus
- †
†
Fossil of the Devonian trilobite Phacops rana
†Phacops
†
- †
†Phestia
†
- † – or unidentified comparable form
† – type locality for genus
- † – type locality for species
- † – type locality for species
†
†
- †
†
†
†
- † – type locality for species
- † – type locality for species
†
- †
†
- † – type locality for species
- † – type locality for species
†
- †
Pinna
†
- †
†
- † – type locality for species
- † – type locality for species
†
- †
† – tentative report
- †
†Planutenia
- †
† – type locality for genus
- † – type locality for species
Fossilized shell of the Silurian-Early Triassic sea snail Platyceras
†Platyceras
- †
- †
†
†
- †
†Pleurodictyum
- †
†
- †
†
- †
†
†
- †
- †
†
- † – type locality for species
†
- †
†
- †
†
†
- †
†Productus
Restoration of the Silurian trilobite Proetus
†Proetus
- † – tentative report
- †
†
- †
†
- †
- †
†
†Protobarinophyton
- †
Illustration of a fossil of the Cambrian crustacean Protocaris
†Protocaris
†
- †
†
- †
† – or unidentified comparable form
†Protospongia – tentative report
†
†
- †
†
- †
†
- † – type locality for species
†
- † – type locality for species
†
- †
- † – tentative report
- †
†
- †
† – type locality for genus
- † – type locality for species
†
†
- †
- †
†
†
- †
†
† – type locality for genus
- † – type locality for species
†
†
†
- † – or unidentified comparable form
†
- †
†
- †
- †
- †
†
†Pugnax
- †
†Pustula
- †
†
- †

Q[]

†Quasillites

R[]

†Rafinesquina
- †
†
- †
- †
†
- †
†
- †
†
†
†
Fossil of the Silurian eurypterid ("sea scorpion") Rhinocarcinosoma
†Rhinocarcinosoma
- †
†Rhipidomella
- †
- †
- †
†Rhodea
- † – type locality for species
- † – type locality for species
- † – type locality for species
†Rhombopora
†
- †
†Ribeiria
- † – or unidentified comparable form
†
†
- † – type locality for species
†Ruedemannipterus
- †
†
- †

S[]

Illustration of fossilized shells of the mysterious Cambrian organism Salterella
†Salterella
- †
- †Salterella conulata
- †
- †
†
†
- †
†
- † – informal
Fossilized fin of the Devonian lobe-finned fish Sauripterus
†Sauripterus
- †Sauripterus taylori
† – or unidentified comparable form
†
- † – type locality for species
†Schizodiscus
- †
†Schizodus
- †
†Schizophoria
- †
- †
- †
†
- †
- †
†
- †
†Sciadophyton
†
†Selkirkia
†Septopora
†
†
- †
Fossils of the Cambrian arthropod Sidneyia
†Sidneyia
†Sigillaria
†
- †
†
- †
†
- †
- †
†Solenochilus
- †
†
- † – type locality for species
- † – type locality for species
†
- † – type locality for species
†Sowerbyella
- †
- †
†Spathognathodus
†
- †
- †
†
- † – type locality for species
†
- †
- †
†
- †
- †
- †
†
- †
Fossilized shell of the Late Ordovician-Late Triassic brachiopod Spirifer
†Spirifer
- †
- †
†
†Spyroceras
- †
†
- †
†
- †
† – type locality for genus
- † – type locality for species
†
†
- † – type locality for species
†
†
- † – tentative report
†
- †
†
- †
†
- †
Fossilized shell of the Ordovician-Silurian brachiopod Strophomena
†Strophomena
†
- †
- †
†
- †
†
- †
- †
†
- †
- † – or unidentified comparable form
† – type locality for genus
- † – type locality for species
†
†
- † – type locality for species
- † – type locality for species

T[]

†Tabulipora
†
- †
†
- †
Fossils of the Devonian plant stem morphogenus Taeniocrada
†Taeniocrada
†
- †
†
†
†
- †
†Tentaculites
- †
†
- †
†Tetraxylopteris
†
- †
†
- †
†
- † – type locality for species
†Titusvillia – type locality for genus
- † – type locality for species
†
- †
Restoration showing the top (left) and underside of the Late Ordovician trilobite Triarthrus
†Triarthrus
- †
†Tricrepicephalus
- † – type locality for species
†Trimerus
- †
†
- † – type locality for species
- † – type locality for species
- †
- †
†
- †
†
- †
†
- †
†Truncatiramus – now regarded as a jr. synonym of Erettopterus^[1]^[2]
†
†Tubulella
†Tuzoia
†
- †
- †

U[]

†
- †
- †
Fossil of the Ordovician-Devonian sea star Urasterella
†Urasterella
- †

V[]

†
- †
†
†
- †
- †
†Vinella

W[]

Fossil of the Cambrian trilobite Wanneria
†Wanneria
- †
†
- †
†
- † – or unidentified comparable form
†
- †
†Wilkingia
- †
†Worthenia
- †

X[]

†
- † – type locality for species
†
- †

Y[]

Life restoration of the probable Cambrian mollusc Yochelcionella
†Yochelcionella
- † – type locality for species
- †
- †
†
- †

Z[]

†
†
†
†

References[]

^ Ciurca, Samuel J.; Tetlie, O. Erik (2007). "Pterygotids (Chelicerata; Eurypterida) from the Silurian Vernon Formation of New York". Journal of Paleontology. 81 (4): 725–736. doi:10.1666/pleo0022-3360(2007)081[0725:PEFTSV]2.0.CO;2. ISSN 0022-3360.
^ Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 18.5 http://www.wsc.nmbe.ch/resources/fossils/Fossils18.5.pdf (PDF).

Various Contributors to the Paleobiology Database. "Fossilworks: Gateway to the Paleobiology Database". Retrieved 17 December 2021. {{cite web}}: |author= has generic name (help)

[:5-1] Ciurca, Samuel J.; Tetlie, O. Erik (2007). "Pterygotids (Chelicerata; Eurypterida) from the Silurian Vernon Formation of New York". Journal of Paleontology. 81 (4): 725–736. doi:10.1666/pleo0022-3360(2007)081[0725:PEFTSV]2.0.CO;2. ISSN 0022-3360.

[:0-2] Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 18.5 http://www.wsc.nmbe.ch/resources/fossils/Fossils18.5.pdf (PDF).

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