Orthophlebiidae

Orthophlebiidae Temporal range: Middle Triassic–Early Cretaceous PreꞒ Ꞓ O S D C P T J K Pg N
Species of Orthophlebia and Mesopanorpa from the Middle Jurassic of China
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Mecoptera
Superfamily:
Family:	†Orthophlebiidae Handlirsch, 1906
Genera
See text

Orthophlebiidae is an extinct family of scorpionflies known from the Triassic to Cretaceous, belonging to the superfamily . The family is poorly defined and is probably paraphyletic, representing many primitive members of Panorpoidea with most species only known from isolated wings, and has such been considered a wastebasket taxon.^[1][2]

Systematics[]

Based on^[3]

The genus Protorthophlebia has been moved to the separate family Protorthophlebiidae.^[4]

• † Riek 1950 Hawkesbury Sandstone, Australia, Anisian, Magyden Formation, Kyrgyzstan, Carnian, Koonwarra Fossil Bed, Australia, Aptian
• † Jell and Duncan 1986 Koonwarra Fossil Bed, Australia, Aptian
• † Handlirsch 1906
  • †Mesopanorpa angarensis Martynov 1927 , Russia, Toarcian
  • †Mesopanorpa brodiei Tillyard 1933 Whitby Mudstone, United Kingdom, Toarcian, , China, Sinemurian
  • †Mesopanorpa brooksorum Jarzembowski and Soszyńska-Maj, 2018 Weald Clay, United Kingdom, Barremian
  • †Mesopanorpa densa Zhang 1996 Badaowan Formation, China, Sinemurian
  • †Mesopanorpa enormis Lin 1986 , China, Bajocian
  • †Mesopanorpa fanshanensis Ren 1995 Lushangfen Formation, China, Aptian
  • †Mesopanorpa felix Martynov 1927 Karabastau Formation, Kazakhstan, Oxfordian
  • †Mesopanorpa formosa Bode 1953 Posidonia Shale, Germany, Toarcian
  • †Mesopanorpa gambra Lin 1980 , China, Barremian
  • †Mesopanorpa hartungi Brauer et al. 1889 Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian, Cheremkhovskaya Formation, Russia, Toarcian
  • †Mesopanorpa incerta Martynov 1927 Cheremkhovskaya Formation, Russia, Toarcian
  • †Mesopanorpa kuliki Martynova 1948 Badaowan Formation, China, Sinemurian, Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
  • †Mesopanorpa kuschmurunensis Martynova 1956, Kushmurun Formation, Russia, Early Jurassic
  • †Mesopanorpa maculata Handlirsch 1939 "Green Series", Germany, Toarcian
  • †Mesopanorpa martynovae Sukacheva 1990 , Russia, Tithonian
  • †Mesopanorpa monstrosa Zhang 1996 Badaowan Formation, China, Sinemurian
  • †Mesopanorpa obtusa Bode 1953 Posidonia Shale, Germany, Toarcian
  • †Mesopanorpa unica Sukatsheva 1985 Ichetuy Formation, Russia, Oxfordian
  • †Mesopanorpa unicolor Martynov 1937 , Kyrgyzstan, Toarcian
  • †Mesopanorpa yaojiashanensis Lin 1980 Laocun Formation, China, Barremian
• † Willmann 1989 Dzhil Formation, Kyrgyzstan, Hettangian-Sinemurian
• † Lin 1976 , China, Barremian
• † Bode 1953 Posidonia Shale, Germany, Toarcian
• † Handlirsch 1906 Lulworth Formation, United Kingdom, Berriasian
• Subfamily Gigaphlebiinae Soszyńska-Maj and Krzemiński, 2018
  • † Soszyńska-Maj and Krzemiński, 2018 , Kyrgyzstan, Hettangian-Sinemurian, Daohugou, China, Callovian, Karabastau Formation, Kazakhstan, Oxfordian
  • Soszyńska-Maj and Krzemiński, 2018 Daohugou, Tiaojishan Formation, China, Callovian
• †, Soszyńska-Maj & Krzemiński 2019^[4] Daohugou, China, Callovian
• † Westwood 1845 Middle Triassic-Early Cretaceous, Eurasia
  • †Orthophlebia (Dolichophlebia) Hong and Zhang 2004
    • †Orthophlebia (Dolichophlebia) ladinica Hong 2009 , China, Ladinian
    • †Orthophlebia (Dolichophlebia) xiaofangzhangziensis Hong 1983 Jiulongshan Formation, China, Callovian
  • †Orthophlebia aequalis Martynov 1937 Kyzyl-Kiya, Kyrgyzstan, Pliensbachian
  • †Orthophlebia anglica Handlirsch 1939 United Kingdom, Rhaetian
  • †Orthophlebia angustata Martynov 1937 Sulyukta Formation, Kyrgyzstan, Toarcian
  • †Orthophlebia bella Handlirsch 1939 United Kingdom, Rhaetian/Hettangian
  • †Orthophlebia bifurcata Giebel 1856 Lulworth Formation, United Kingdom, Berriasian
  • †Orthophlebia bolboica Sukacheva 1990 , Russia, Tithonian
  • †Orthophlebia brunsvicensis Bode 1905 Posidonia Shale, "Green Series" Germany, Toarcian
  • †Orthophlebia capillata Whalley 1985 Charmouth Mudstone Formation, England, Sinemurian
  • †Orthophlebia colorata Zhang 1996 , China, Sinemurian
  • †Orthophlebia compacta Bode 1953 Posidonia Shale, Germany, Toarcian
  • †Orthophlebia confusa Willmann 1977 Lilstock Formation, United Kingdom, Rhaetian
  • †Orthophlebia curta Martynova 1948 , Kyrgyzstan, Hettangian-Sinemurian
  • †Orthophlebia deformis Lin 1986 , China, Bajocian
  • †Orthophlebia effusa Martynova 1948 , Kyrgyzstan, Hettangian-Sinemurian
  • †Orthophlebia elenae Willmann and Novokshonov 1998 Karabastau Formation, Kazakhstan, Oxfordian
  • †Orthophlebia elongata Handlirsch 1939 "Green Series" Germany, Toarcian
  • †Orthophlebia exculpta Zhang 1996 , China, Aalenian/Bajocian
  • †Orthophlebia extensa Martynov 1937 , Kyrgyzstan, Toarcian
  • †Orthophlebia fallerslebensis Bode 1953 Posidonia Shale, Germany, Toarcian
  • †Orthophlebia fracta Sukacheva 1990 , Russia, Aptian
  • †Orthophlebia furcata Handlirsch 1939 England, Rhaetian/Hettangian
  • †Orthophlebia fuscipennis Handlirsch 1906 "Green Series" Germany, Toarcian
  • †Orthophlebia germanica Handlirsch 1906 "Green Series" Germany, Toarcian
  • †Orthophlebia gracilis Handlirsch 1939 Lilstock Formation, England, Rhaetian
  • †Orthophlebia gubini Novokshonov and Sukatsheva 2003 Ulan Malgait Formation, Mongolia, Tithonian
  • †Orthophlebia heidemariae Willmann and Novokshonov 1998 Karabastau Formation, Kazakhstan, Oxfordian
  • †Orthophlebia intermedia Giebel 1856 Lilstock Formation, England, Rhaetian
  • †Orthophlebia jejuna Sukatsheva 1985 , Russia, Toarcian
  • †Orthophlebia karabonica Sukacheva 1990 , Russia, Aptian
  • †Orthophlebia laesa Handlirsch 1939 England, Rhaetian
  • †Orthophlebia latebrosa Sukatsheva 1985 Badaowan Formation, China, Sinemurian , Russia, Toarcian
  • †Orthophlebia latipennisimilis Bode 1953 Posidonia Shale, Germany, Toarcian
  • †Orthophlebia lauta Sukatsheva 1985 , Russia, Toarcian
  • †Orthophlebia liadis Handlirsch 1939 "Green Series" Germany, Toarcian
  • †Orthophlebia liaoningensis Ren 1997 Yixian Formation, China, Aptian
  • †Orthophlebia liassica Mantell 1844 Lilstock Formation, United Kingdom, Rhaetian/Hettangian
  • †Orthophlebia limnophila Handlirsch 1906 "Green Series" Germany, Toarcian
  • †Orthophlebia lithographica Willmann and Novokshonov 1998 Solnhofen Limestone, Germany, Tithonian
  • †Orthophlebia longicauda Willmann and Novokshonov 1998 Karabastau Formation, Kazakhstan, Oxfordian
  • †Orthophlebia luanpingensis Hong 1983 Jiulongshan Formation, China, Callovian
  • †Orthophlebia maculata Martynov 1927 , Kyrgyzstan, Hettangian-Sinemurian, Karabastau Formation, Kazakhstan, Oxfordian
  • †Orthophlebia mongolica Sukatsheva 1985 , Mongolia, Bathonian
  • †Orthophlebia nana Handlirsch 1939 United Kingdom, Rhaetian/Hettangian
  • †Orthophlebia nervulosa Qiao et al. 2012 Jiulongshan Formation, China, Callovian
  • †Orthophlebia obunca Sukatsheva 1985 Itat Formation, Russia, Bathonian
  • †Orthophlebia parvula Handlirsch 1939 United Kingdom, Rhaetian/Hettangian
  • †Orthophlebia phryganoides Martynov 1925 Karabastau Formation, Kazakhstan, Oxfordian
  • †Orthophlebia pictipennis Tillyard 1933 Lilstock Formation, United Kingdom, Rhaetian/Hettangian
  • †Orthophlebia pulchra Martynova 1956 , Kazakhstan, Early Jurassic
  • †Orthophlebia pygmaea Handlirsch 1939 United Kingdom, Rhaetian
  • †Orthophlebia quadrimacula Lin 1982 Zhiluo Formation, China, Middle Jurassic
  • †Orthophlebia radialis Handlirsch 1939 "Green Series" Germany, Toarcian
  • †Orthophlebia retorrida Sukatsheva 1985 , Russia, Toarcian
  • †Orthophlebia rossica Martynova 1948 , Kyrgyzstan, Hettangian-Sinemurian
  • †Orthophlebia rotundipennis Martynov 1937 , Kyrgyzstan, Toarcian , China, Bajocian
  • †Orthophlebia shartegica Novokshonov and Sukatsheva 2003 Ulan Malgait Formation, Mongolia, Tithonian
  • †Orthophlebia shurabica Martynov 1937 , Kyrgyzstan, Toarcian
  • †Orthophlebia speciosa Bode 1953 Posidonia Shale, Germany, Toarcian
  • †Orthophlebia stigmatica Handlirsch 1939 Lilstock Formation, United Kingdom, Rhaetian/Hettangian
  • †Orthophlebia varia Martynova 1948 , Kyrgyzstan, Hettangian-Sinemurian
  • †Orthophlebia venosa Martynov 1937 , Kyrgyzstan, Toarcian
  • †Orthophlebia vernacula Martynova 1948 , Kyrgyzstan, Hettangian-Sinemurian
  • †Orthophlebia vicina Handlirsch 1939 "Green Series" Germany, Toarcian
  • †Orthophlebia yangjuanxiangensis Hong 1985 , China, Toarcian
  • †Orthophlebia yaogouensis Hong 1983 Jiulongshan Formation, China, Callovian

References[]