Nemastomatidae
Nemastomatidae Temporal range:
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Nemastoma bimaculatum from Belgium | |
Scientific classification | |
Kingdom: | Animalia
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Phylum: | Arthropoda
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Class: | Arachnida
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Order: | |
Suborder: | |
Superfamily: | |
Family: | Nemastomatidae Simon, 1872
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Species | |
See text | |
Diversity | |
c. 170 species |
The Nemastomatidae are a family of harvestmen with about 170 described species in 16 recent genera. Several fossil species and genera are known.
Unlike some related currently recognized families, the Nemastomatidae are monophyletic.[1]
Description[]
Members of the Nemastomatidae range in body length from about one to almost six millimeters. Their chelicerae are of normal proportions, but the pedipalps are very elongated and thin in some groups. Leg length is likewise variable.[1]
Distribution[]
The subfamily Ortholasmatinae (ten species in two genera) occur on both sides of the Pacific Ocean: in western North America from Mexico to British Columbia and eastern Asia (Japan and northern Thailand. The other subfamily, Nemastomatinae, occur all over Europe up to Iceland and the Caucasus, in the Atlas Mountains of North Africa, from Anatolia to northern Iran, with a few species found outside this region, in Central Asia and the Himalayas. Most species are restricted to very small mountainous southern regions.[1]
Relationships[]
The Nemastomatidae are probably a sister group to the genera Dicranolasma (Dicranolasmatidae) and (Trogulidae).[1]
Name[]
The genus name Nemastoma is a combination of Ancient Greek nema "thread" and stoma "mouth", referring to the elongated pedipalps.[1]
Species[]
Subfamily Nemastomatinae[]
- Kratochvíl & Miller, 1958
- Silhavý, 1966
- (Ljovuschkin & Starobogatov, 1963)
- Levushkii, 1972
- (Lebedinskiy, 1914) — Crimea
- Giljarovia Kratochvíl & Miller, 1958
- Kratochvíl & Miller, 1958
- (Kratochvíl & Miller, 1958)
- Gruber, 1976
- Nemastoma C. L. Koch, in Hahn & C. L. Koch 1836
- subgenus Nemastoma
- Roewer, 1951
- Roewer, 1951
- Roewer, 1962
- Roewer, 1951
- Roewer, 1927
- Simon, 1879 — Spain
- Nemastoma bacilliferum bacilliferum Simon, 1879 — Pyrenees
- Nemastoma bacilliferum simoni Roewer, 1914
- Nemastoma bacilliferum simplex Simon, 1913
- Nemastoma bacilliferum dipentatum Rambla, 1959
- Roewer, 1914 — central Europe
- Nemastoma bidentatum bidentatum Roewer, 1914
- Nemastoma bidentatum sparsum Gruber & Martens, 1968
- Nemastoma bidentatum relictum Gruber & Martens, 1968
- Roewer, 1951
- Roewer, 1951
- Grese, 1911 — Crimea
- Roewer, 1917 — Bosnia
- Roewer, 1951
- Simon, 1907 — Spain
- Roewer, 1951
- † Menge, 1854 — fossil
- Roewer, 1917 — Korfu
- Roewer, 1951
- † Koch & Berendt, 1854 — Baltic amber fossil: Oligocene
- Canestrini, 1873 — Italy
- Mello-Leitão, 1936 — Spain
- Roewer, 1959
- Roewer, 1951
- Roewer, 1917 — Transcaucasus
- Roewer, 1951
- Kraus, 1959
- Redikorzev, 1936 — Russia
- Roewer, 1923 — Serbia
- , in Lendl 1894 — Hungary
- L. Koch, 1867
- Hadzi, 1973
- Redikorzev, 1936 — Russia
- Kulczynski, 1909 — Portugal
- Kraus, 1959
- Roewer, 1951
- Roewer, 1951
- Roewer, 1917 —
- Roewer, 1951
- Caporiacco, 1940 — Italy
- Wankel, 1851
- (H. Lucas, 1847) — Algeria
- Roewer, 1959
- Roewer, 1951
- Schenkel, 1947 — Albania
- Simon, 1913 — Algeria
- Roewer, 1951
- Roewer, 1923 — Serbia
- Hadzi, 1973
- Simon, 1913 — Spain
- Roewer, 1917 — Serbia and Bosnia
- Kraus, 1961
- Nosek, 1904 — Montenegro
- Roewer, 1951
- Roewer, 1923 — Serbia
- Roewer, 1951
- Roewer, 1951
- Simon, 1879 — Pyrenees
- Roewer, 1951
- Roewer, 1917 — Spain
- Roewer, 1951
- Roewer, 1951
- Simon, 1881 — Alps
- Roewer, 1951
- Simon, 1879 — Pyrenees
- Roewer, 1951
- Gruber & Martens, 1968
- Roewer, 1951
- Simon, 1911 — Spain
- Giltay, 1933 — Balkans
- Kraus, 1961
- L. Koch, 1869 — Greece
- Roewer, 1951
- Simon, 1885 — Greece
- † Roewer, 1939 — fossil: Baltic amber
- Redikorzev, 1936 — Russia
- Hadzi, 1973
- Roewer, 1914 — Bosnia
- Gruber & Martens, 1968
- (C. L. Koch, 1835) — central Europe
- Nemastoma triste triste C. L. Koch in Herrich-Schäffer, 1835
- (Wankel, 1861)
- † Koch & Berendt, 1854 — fossil: Oligocene
- Roewer, 1951
- Roewer, 1927
- subgenus Dromedostoma
- Roewer, 1951
- Nemastoma acrospinosum acrospinosum Roewer, 1951
- Nemastoma acrospinosum pretneri Hadzi, 1973
- Hadzi, 1973
- Hadzi, 1973
- Hadzi, 1973
- Hadzi, 1973
- Hadzi, 1973
- Hadzi, 1973
- Hadzi, 1973
- Roewer, 1926 — cave in Balkans
- Roewer, 1951
- Hadzi, 1973
- subgenus Lugubrostoma
- Hadzi, 1973
- Nemastoma lugubre (Müller, 1776)
- Nemastoma lugubre lugubre (Müller, 1776)
- Nemastoma lugubre bimaculatum (Fabricius, 1775) — Europe
- Nemastoma lugubre unicolor Roewer, 1914 — southern Europe
- Hadzi, 1973
- Redikorzew, 1936
- (C. L. Koch, 1835)
- Avram, 1973
- (Mcheidze, 1952)
- (Roewer, 1951)
- (Nowicki, 1870)
- (Perty, 1833) — Europe
- Paranemastoma quadripunctatum quadripunctatum (Perty, 1833)
- Paranemastoma quadripunctatum aurosum L. Koch, 1869
- Paranemastoma quadripunctatum armatum Kulczynski, 1909 — southeastern Europe
- Paranemastoma quadripunctatum thessalum Simon, 1885 — Greece
- Paranemastoma quadripunctatum werneri Kulczynski, 1903 — Asia Minor
- Paranemastoma quadripunctatum humerale C. L. Koch, 1839 — southeastern Europe
- Starega, 1978
- (Hermann, 1871) — central Europe
- (Roewer, 1951)
- (Roewer, 1951)
- (Roewer, 1951)
- (Roewer, 1951)
- Paranemastoma aurigerum aurigerum Roewer, 1951
- Paranemastoma aurigerum joannae Starega, 1976
- Kratochvíl & Miller, 1958
- Kratochvíl & Miller, 1958
- (Ausserer, 1867) — central Europe
- (Canestrini, 1872) — , Italy
- (Redikorzev, 1936) — Russia
- Carinostoma Kratochvíl & Miller, 1958
- (Roewer, 1914) — southeastern Europe
- Carinostoma elegans (Sørensen, 1894) — Hungary
- Carinostoma elegans elegans (Sørensen, 1894)
- Carinostoma elegans batorligetiense (Szalay, 1951)
- (Hadzi, 1940)
- Kratochvíl & Miller, 1958
- subgenus Centetostoma
- (Simon, 1881) — Alps
- subgenus Cretostoma
- (Roewer, 1927)
- Roewer, 1951
- Roewer, 1962
- Tedeschi & Sciaky, 1997
- Tedeschi & Sciaky, 1997
- Tedeschi & Sciaky, 1997
- Avram, 1969
- Avram, 1969
- Roewer, 1958
- Roewer, 1951
- Roewer, 1951
- Mitostoma chrysomelas (Hermann, 1804) — Europe
- Mitostoma chrysomelas chrysomelas (Hermann, 1804)
- Mitostoma chrysomelas michielii Hadzi, 1973
- Mitostoma chrysomelas multidenticulatum Hadzi, 1973
- Mitostoma chrysomelas alpinum Hadzi, 1931 — Yugoslavia
- (Fage, 1946) — Italy
- Hadzi, 1973
- Avram, 1970
- Hadzi, 1973
- Avram, 1970
- Avram, 1970
- Hadzi, 1973
- Silhavý, 1946 — Dalmatia, Montenegro
- Mitostoma olgae olgae Silhavý, 1946
- Mitostoma olgae decorum Silhavý, 1946
- Mitostoma olgae kratochvili Silhavý, 1946
- Mitostoma olgae zorae Hadzi, 1973
- Marcellino, 1977
Subfamily Ortholasmatinae[]
- Cladolasma Suzuki, 1963
- Cladolasma ailaoshan Zhang, Zhao & Zhang, 2018 — China
- Cladolasma angka (Schwendinger & Gruber, 1992) — Thailand
- Cladolasma damingshan Zhang & Zhang, 2013 — China
- Cladolasma parvulum Suzuki, 1963 — Japan
- Dendrolasma Banks, 1894
- Dendrolasma mirabile Banks, 1894 — Washington
- P. J. Schwendinger & J. Gruber, 1992 — Thailand
- W. A. Shear & J. Gruber, 1983 — California
- (Suzuki, 1963)
- † Giribet & Dunlop, 2005
- † Giribet & Dunlop, 2005 — fossil: Cretaceous (Cenomanian) Burmese amber,[2]
- Ortholasma Banks, 1894
- (Goodnight & Goodnight, 1942) — Mexico
- Ortholasma coronadense Cockerell, 1916 — Coronados Islands
- W. A. Shear & J. Gruber, 1983 — California
- Ortholasma pictipes Banks, 1911 — California
- Ortholasma rugosum Banks, 1894 — California
- Silhavý, 1973
incertae sedis[]
- Mello-Leitão, 1936
- Mello-Leitão, 1936 — Spain
- Hadzinia Silhavý, 1966
- (Hadzi, 1940)
- Acromitostoma Roewer, 1951
- (Roewer, 1917) — Spain: Granada
- Roewer, 1951
- Kratochvíl & Miller, 1958
- (Roewer, 1917)
- (Roewer, 1951)
- W. Starega, 1986 — Tajikistan
- Gruber, 1976
- Starega, 1976
- (Roewer, 1916)
- † Haupt, 1956 — fossil
- † Haupt, 1956 — Permian
Footnotes[]
References[]
- Joel Hallan's Biology Catalog: Nemastomatidae
- Giribet, G.; Dunlop, J. (2005). "First identifiable Mesozoic harvestman (Opiliones: Dyspnoi) from Cretaceous Burmese amber" (PDF). Proceedings of the Royal Society B. 272 (1567): 1007–1013. doi:10.1098/rspb.2005.3063. PMC 1039256. PMID 1599874.
- Pinto-da-Rocha, R., Machado, G. & Giribet, G. (eds.) (2007): Harvestmen - The Biology of Opiliones. Harvard University Press ISBN 0-674-02343-9
Wikimedia Commons has media related to Nemastomatidae. |
- Harvestman families