2011 in paleontology

From Wikipedia, the free encyclopedia
List of years in paleontology (table)
In paleobotany
2012
2013
2014
In arthropod paleontology
2009
2010
2011
2012
2013
2014
In paleoentomology
In paleomalacology
2013
2014
In reptile paleontology
In archosaur paleontology
2009
2010
2011
2012
2013
In mammal paleontology
2009
2010
2011
2012
2013
2014
In paleoichthyology
2010
2011

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2011.

Protozoa[]

New taxa[]

Name Novelty Status Authors Age Unit Location Notes Images

Vetufebrus[2]

Gen et sp nov

Valid

Poinar

Burdigalian

Dominican amber

 Dominican Republic

A Plasmodiidae vectored by the bat fly Enischnomyia stegosoma

Vetufebrus ovatus

Plants[]

Angiosperms[]

Name Novelty Status Authors Age Unit Location Notes Images

Palmoxylon lametaei[3]

Species

Valid

Dutta, Ambwani, & Estrada-Ruiz

Maastrichtian

Lameta Formation

 India

A petrified palm wood

Leefructus[4]

Gen. et sp.

Valid

Sun, Dilcher, Wang, & Chen

Early Cretaceous

Yixian Formation

 China

A basal eudicot

Gymnosperms[]

Name Novelty Status Authors Age Unit Location Notes Images

Qataniaria[5]

Gen. et sp.

Valid

Krassilov & Schrank

Albian

 Israel

[6]

Gen. et sp.

Valid

Kunzmann, Mohr, Wilde & Bernardes-de-Oliveira

Aptian

Crato Formation

 Brazil

Pteridophytes[]

Name Novelty Status Authors Age Unit Location Notes Images

Equisetum thermale[7]

sp. nov

Valid

Channing, Zamuner, Edwards, & Guido

Callovian - Tithonian

La Matilde Formation

 Argentina

oldest crown group Equisetum species

Nematoda[]

Name Novelty Status Authors Age Unit Location Notes Images

Formicodiplogaster[8]

Gen et sp nov

valid

Poinar

Burdigalian

Dominican amber

 Dominican Republic

Diplogasteridae nematode, parasite of ant genus Azteca;
Type species Formicodiplogaster myrmenema

Formicodiplogaster myrmenema

Lobopods[]

Name Novelty Status Authors Age Unit Location Notes Images

Diania cactiformis[9]

Valid

Liu et al.

Lower Cambrian

Maotianshan shale

 China

A lobopodian.

Mureropodia apae

Valid

Gamez Vintaned, Linan, & Zhuravlev

Early Cambrian

Murero

 Spain

A lobopodian.

Siberion lenaicus

Valid

Dzik

Early Cambrian

 Russia

A lobopodian.

Vetulicolians[]

Name Novelty Status Authors Age Unit Location Notes Images

Ooedigera peeli[10]

gen. nov

valid

Zinther, et al.

Early Cambrian

Sirius Passet

 Greenland

earliest known vetulicolian described

Artist's reconstruction

Molluscs[]

Arthropods[]

Fishes[]

Amphibians[]

Newly named lepospondyls[]

Name Novelty Status Authors Age Unit Location Notes Images

Kirktonecta[11]

Gen. et sp.

Valid

  • Clack

Viséan

East Kirkton Quarry

  •  Scotland

A microsaur.

Tambaroter[12]

Gen. et sp.

Valid

Early Permian

Tambach Formation

  •  Germany

An ostodolepid.

Newly named temnospondyls[]

Name Novelty Status Authors Age Unit Location Notes Images

Acheloma dunni[13]

Species

Valid

  • Polley
  • Reisz

Early Permian

  •  USA

A trematopid.

Calmasuchus[14]

Gen. et sp.

Valid

  • Fortuny
  • Galobart
  • De Santisteban

early-middle Anisian

  •  Spain

A capitosaurian.

Rotaryus[15]

Gen. et sp.

Valid

Early Permian

Tambach Formation

  •  Germany

A trematopid.

Rubeostratilia[16]

Gen. et sp.

Valid

  • Bourget
  • Anderson

Early Permian

  •  USA

An amphibamid.

Rubeostratilia texensis.

Newly named lissamphibians[]

Name Novelty Status Authors Age Unit Location Notes Images

Urupia[17]

Gen. et sp.

Valid

  • Skutschas
  • Krasnolutskii

Bathonian

Itat Formation

  •  Russia

A basal stem salamander.

Basal reptiles[]

Newly named captorhinids[]

Name Novelty Status Authors Age Unit Location Notes Images

Gansurhinus[18]

Gen. et sp.

Valid

Middle Permian

  •  China

A moradisaurine .

Newly named basal diapsids[]

Name Novelty Status Authors Age Unit Location Notes Images

Orovenator[19]

Gen. et sp.

Valid

Sakmarian

  •  USA

The oldest and most basal neodiapsid.

Newly named ichthyosaurs[]

Name Novelty Status Authors Age Unit Location Notes Images

Sveltonectes[20]

Gen. et sp.

Valid

Barremian

  •  Russia

An ophthalmosaurid.

Lepidosauromorphs[]

Newly named saurosphargids[]

Name Novelty Status Authors Age Unit Location Notes Images

Sinosaurosphargis[21]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

  •  China

An incertae sedis.

Newly named sauropterygians[]

Name Novelty Status Authors Age Unit Location Notes Images

Abyssosaurus[22]

Gen. et sp.

Valid

  • Berezin

Late Hauterivian

  •  Russia

An aristonectid.

Diandongosaurus[23]

Gen. et sp.

Valid

  • Shang
  • Wu
  • Li

Middle Anisian

Guanling Formation

  •  China

A relative of nothosauroids.[24]

Dianopachysaurus[25]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

  •  China

A keichousaurid pachypleurosaur.

Hauffiosaurus tomistomimus[26]

Species

Valid

Early Toarcian

  •  England

A second species of Hauffiosaurus.

Marmornectes[27]

Gen. et sp.

Valid

  • Ketchum
  • Benson

Callovian

Oxford Clay Formation

  •  United Kingdom

A pliosaurid.

Westphaliasaurus[28]

Gen. et sp.

Valid

  • Schwermann
  • Sander

Pliensbachian

Höxter district

  •  Germany

A plesiosaurid plesiosaur.

Zarafasaura[29]

Gen. et sp.

Valid

Maastrichtian

  •  Morocco

An elasmosaurid plesiosaur.

Newly named lizards[]

Name Novelty Status Authors Age Unit Location Notes Images

Brasiliguana[30]

Gen. et sp.

Valid

  • Nava
  • Martinelli

Turonian/Santonian

Adamantina Formation

  •  Brazil

An iguanian lizard.

Cryptolacerta[31]

Gen. et sp.

Valid

Lutetian

Messel pit

  •  Germany

An amphisbaenian lizard.

Latoplatecarpus[32]

Gen. et sp.

Valid

  • Konishi
  • Caldwell

Campanian

Pierre Shale Formation
Demopolis Chalk Formation

  •  Canada
  •  USA

A plioplatecarpine mosasaur. Its type species is Latoplatecarpus willistoni; "Plioplatecarpus" nichollsae (Cuthbertson et al.., 2007) is a second species of Latoplatecarpus.

[33][34]

Gen. et sp. nov

Valid

  • Smith

Eocene (Chadronian)

  •  United States

A member of Corytophanidae. The type species is Oreithyia oaklandi.

Plesioplatecarpus[32]

Gen. et comb.

Valid

  • Konishi
  • Caldwell

Middle Coniacian to middle Santonian.

Niobrara Chalk
Eutaw Formation

  •  USA

A plioplatecarpine mosasaur, a new genus for "Clidastes" planifrons (Cope, 1874).

[33]

Gen. et sp. nov

Valid

  • Smith

Eocene (Chadronian)

  •  United States

A relative of the desert iguana. The type species is Queironius praelapsus.

[33]

Gen. et comb. nov

Valid

  • Smith

Eocene (Chadronian)

  •  United States

A member of Polychrotidae; a new genus for "Polychrus" charisticus Smith (2006).

Newly named snakes[]

Name Novelty Status Authors Age Unit Location Notes Images

Madtsoia pisdurensis[35]

Species

Valid

  • Mohabey
  • Head
  • Wilson

Late Cretaceous

  •  India

A madtsoiid snake.

Turtles[]

Newly named turtles[]

Name Novelty Status Authors Age Unit Location Notes Images

Albertwoodemys[36]

Gen. et sp.

Valid

Early Oligocene

Jebel Qatrani Formation

  •  Egypt

A podocnemidid.

Brontochelys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

  •  Pakistan

A podocnemidid, a new genus for "" gaffneyi.

Cordichelys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Eocene

Qasr el-Sagha beds

  •  Egypt

A podocnemidid, a new genus for "Podocnemis" antiqua.

Gilmoremys[37]

Gen. et comb.

Valid

  • Joyce
  • Lyson

Maastrichtian

Lance Formation
Hell Creek Formation

  •  USA

A softshell turtle, new genus for "Aspideretes" lancensis (Gilmore, 1928).

[38]

Sp. nov

Valid

  • Ehret & Bourque

Late Pleistocene

  •  United States

An emydid, a species of Graptemys.

Hispaniachelys[39]

Gen. et sp.

Valid

Oxfordian

  •  Spain

A paracryptodire or a basal testudine.

Hoplochelys clark[40]

Species

Valid

Maastrichtian

Hell Creek Formation

  •  USA

A kinosternoid, a species of Hoplochelys.

Lapparentemys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Paleocene

Santa Lucía Formation

  •  Bolivia

A podocnemidid, a new genus for "" vilavilensis.

Latentemys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Uncertain, possibly Miocene.

Uncertain, possibly .

  •  Egypt?

A podocnemidid.

Lemurchelys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

  •  Egypt

A podocnemidid.

Mogharemys[36]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Miocene

  •  Egypt

A podocnemidid, a new genus for "Sternothaerus" blanckenhorni.

[41]

Gen. et comb. nov.

Valid

  • Vitek

Eocene

Washakie Formation

  •  USA

A trionychid, a new genus for "Trionyx" uintaensis.

Patagoniaemys[42]

Gen. et sp.

Valid

  • Sterli
  • De la Fuente

Campanian - Maastrichtian

La Colonia Formation

A stem turtle.

Peiropemys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

  •  Brazil

A podocnemidid.

Pricemys[36]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

  •  Brazil

A podocnemidid.

Prochelidella cerrobarcinae[43]

Species

Valid

Aptian/Albian

Cerro Barcino Formation

The oldest known chelid, a species of Prochelidella.

Selenemys[44]

Gen. et sp.

Valid

  • Pérez-García
  • Ortega

Upper Kimmeridgian

Lusitanian Basin

  •  Portugal

A pleurosternid.

Sokatra[45]

Gen. et sp.

Valid

  • Gaffney
  • Krause

Maastrichtian

Maevarano Formation

A pleurodiran.

Trionyx ikoviensis[46]

Species

Valid

Middle Eocene

  •  Ukraine

A species of Trionyx sensu lato.

[47]

Gen. et sp. nov

Valid

Late Miocene-Pliocene (Hemphillian-Blancan)

Ellensburg Formation

  •  USA

A member of Emydidae belonging to the subfamily Emydinae. The type species is Wilburemys yakimensis.

Archosauromorphs[]

Newly named basal archosauromorphs[]

Name Novelty Status Authors Age Unit Location Notes Images

Archeopelta[48]

Gen. et sp.

Valid

  • Desojo
  • Ezcurra
  • Schultz

Late Ladinian/Early Carnian

Santa Maria Formation

  •  Brazil

A member of Archosauriformes of uncertain phylogenetic placement. Originally classified as a doswelliid, but subsequently argued to be an erpetosuchid archosaur.[49]

Archeopelta arborensis

Augustaburiania[50]

Gen. et sp.

Valid

  • Sennikov

Early Triassic

Don River Basin

  •  Russia

The earliest tanystropheid.

Kyrgyzsaurus[51]

Gen. et sp.

Valid

  • Alifanov
  • Kurochkin

Late Ladinian/Early Carnian

Madygen Formation

The earliest drepanosaurid.

Protanystropheus[50]

Gen. et comb.

Valid

  • Sennikov

Middle - Late Triassic

  •  Austria
  •  Germany
  •  Netherlands
  •  Poland

A tanystropheid, a new genus for "Tanystropheus" antiquus (von Huene, 1905).

Archosaurs[]

Synapsids[]

Newly named non-mammalian synapsids[]

Name Novelty Status Authors Age Unit Location Notes Images

Annatherapsidus postum[52]

Sp.

Valid

  • Ivakhnenko

Late Permian

  •  Russia

A therocephalian.

Basilodon[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" woodwardi.

Bridetherium[54]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

  •  United Kingdom

A morganucodontan.

Candelariodon[55]

Gen. et sp.

Valid

Middle Triassic

Santa Maria Formation

  •  Brazil

A carnivorous cynodont.

Cryptovenator[56]

Gen. et sp.

Valid

Late Gzhelian

  •  Germany

A sphenacodontid pelycosaur. The type species is Cryptovenator hirschbergeri.

Euptychognathus[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" bathyrhynchus.

Euromycter[57]

Gen. et comb.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

  •  France

A large caseid. A new genus for "Casea" rutena.

Gorochovetzia[52]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

  •  Russia

A therocephalian.

Keyseria[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" benjamini.

Muchia[52]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

  •  Russia

A therocephalian.

Paceyodon[54]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

  •  United Kingdom

A morganucodontan.

Peramodon[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

  •  Russia

A dicynodont. A new genus for "Dicynodon" amalitzkii

Perplexisaurus lepusculus[52]

Species

Valid

  • Ivakhnenko

Middle Permian

  •  Russia

A therocephalian.

Purlovia[52]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

  •  Russia

A therocephalian with a short skull.

Ruthenosaurus[57]

Gen. et sp.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

  •  France

A large caseid. The type species is Ruthenosaurus russellorum

Syops[53]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" vanhoepeni.

Tiarajudens[58]

Gen. et sp.

Valid

Capitanian

Rio do Rasto Formation

  •  Brazil

A basal, saber-toothed herbivorous anomodont. The type species is Tiarajudens eccentricus.

Mammals[]

Footnotes[]

Complete author list[]

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.

  1. ^ Henrici, Martens, Berman, Sumida.
  2. ^ Berman, Henrici, Martens, Sumida, Anderson
  3. ^ Reisz, Liu, Li, Müller.
  4. ^ Fischer, Masure, Arkhangelsky, Godefroit.
  5. ^ Li, Rieppel, Wu, Zhao, Wang.
  6. ^ Liu, Rieppel, Jiang, Aitchison, Motani, Zhang, Zhou, Sun.
  7. ^ Benson, Ketchum, Noè, Gómez-Pérez.
  8. ^ Vincent, Bardet, Suberbiola, Bouya, Amaghzaz, Meslouh.
  9. ^ Müller, Hipsley, Head, Kardjilov, Hilger, Wuttke, Reisz.
  10. ^ Gaffney, Meylan, Wood, Simons and Campos
  11. ^ Slater, Reolid, Wood, Schouten and Benton
  12. ^ Knauss, Joyce, Lyson and Pearson
  13. ^ de la Fuente, Umazano, Sterli and Carballido
  14. ^ Danilov, Zvonok, Syromyatnikova and Udovichenko
  15. ^ De Oliveira, Schultz, Soares, Rodrigues.
  16. ^ Fröbisch, Schoch, Müller, Schindler, Schweiss.
  17. ^ a b Reisz, Maddin, Fröbisch, Falconnet.
  18. ^ Cisneros, Abdala, Rubidge, Dentzien-Dias, Bueno.

References[]

  1. ^ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
  2. ^ Poinar, G. (2011). "Vetufebrus ovatus n. gen., n. sp. (Haemospororida: Plasmodiidae) vectored by a streblid bat fly (Diptera: Streblidae) in Dominican amber". Parasites & Vectors. 4 (229): 1–5. doi:10.1186/1756-3305-4-229. PMC 3253689. PMID 22152687.
  3. ^ Dutta, D.; Ambwani, K.; Estrada-Ruiz, E. (2011). "Late Cretaceous palm stem Palmoxylon lametaei sp. nov. from Bhisi Village, Maharashtra, India" (PDF). Revista Mexicana de Ciencias Geológicas. 28 (1): 1–9.[permanent dead link]
  4. ^ Ge Sun; David L. Dilcher; Hongshan Wang; Zhiduan Chen (2011). "A eudicot from the Early Cretaceous of China". Nature. 471 (7340): 625–628. Bibcode:2011Natur.471..625S. doi:10.1038/nature09811. PMID 21455178. S2CID 205224036.
  5. ^ Valentin Krassilov; Eckart Schrank (2011). "New Albian macro- and palynoflora from the Negev (Israel) with description of a new gymnosperm morphotaxon" (PDF). Cretaceous Research. 32 (1): 13–29. doi:10.1016/j.cretres.2010.10.001. Archived from the original (PDF) on 2012-03-30. Retrieved 2011-08-20.
  6. ^ Kunzmann, L.; Mohr, B.A.R.; Wilde, V.; Bernardes-de-Oliveira, M.E.C. (2011). "A putative gnetalean gymnosperm Cariria orbiculiconiformis gen. nov. et spec. nov. from the Early Cretaceous of northern Gondwana". Review of Palaeobotany and Palynology. 165 (1–2): 75–95. doi:10.1016/j.revpalbo.2011.02.005.
  7. ^ Channing, A.; Zamuner, A.; Edwards, D.; Guido, D. (2011). "Equisetum thermale sp. nov. (Equisetales) from the Jurassic San Agustin hot spring deposit, Patagonia: Anatomy, paleoecology, and inferred paleoecophysiology". American Journal of Botany. 98 (4): 680–697. doi:10.3732/ajb.1000211. PMID 21613167.
  8. ^ Poinar, G.; Heiss, E. (2011). "New Termitaphididae and Aradidae (Hemiptera) in Mexican and Dominican amber" (PDF). Palaeodiversity. 4: 51–62.
  9. ^ Liu, J.; Steiner, M.; Dunlop, J. A.; Keupp, H.; Shu, D.; Ou, Q.; Han, J.; Zhang, Z.; Zhang, X. (2011). "An armoured Cambrian lobopodian from China with arthropod-like appendages". Nature. 470 (7335): 526–530. Bibcode:2011Natur.470..526L. doi:10.1038/nature09704. PMID 21350485. S2CID 4324509.
  10. ^ Vinther, J.; Smith, M. P.; Harper, D. A. T. (2011). "Vetulicolians from the Lower Cambrian Sirius Passet Lagerstätte, North Greenland, and the polarity of morphological characters in basal deuterostomes". Palaeontology. 54 (3): 711–719. doi:10.1111/j.1475-4983.2011.01034.x.
  11. ^ Clack, Jennifer A. (2011). "A new microsaur from the early Carboniferous (Viséan) of East Kirkton, Scotland, showing soft tissue evidence". Special Papers in Palaeontology. 86: 45–55.
  12. ^ Henrici, A.C.; Martens, T.; Berman, D.S.; Sumida, S.S. (2011). "An ostodolepid 'microsaur' (Lepospondyli) from the Lower Permian Tambach Formation of central Germany". Journal of Vertebrate Paleontology. 31 (5): 997–1004. doi:10.1080/02724634.2011.596601. S2CID 129710688.
  13. ^ Polley, Brendan P.; Reisz, Robert R. (2011). "A new Lower Permian trematopid (Temnospondyli: Dissorophoidea) from Richards Spur, Oklahoma". Zoological Journal of the Linnean Society. 161 (4): 789–815. doi:10.1111/j.1096-3642.2010.00668.x.
  14. ^ Fortuny, Josep; Galobart, Àngel; De Santisteban, Carles (2011). "A new capitosaur from the Middle Triassic of Spain and the relationships within the Capitosauria" (PDF). Acta Palaeontologica Polonica. 56 (3): 553–66. doi:10.4202/app.2010.0025. S2CID 55068128. Archived from the original (PDF) on 2011-05-22.
  15. ^ David S. Berman; Amy C. Henrici; Thomas Martens; Stuart S. Sumida; Jason S. Anderson (2011). "Rotaryus gothae, a New Trematopid (Temnospondyli: Dissorophoidea) from the Lower Permian of Central Germany". Annals of Carnegie Museum. 80 (1): 49–65. doi:10.2992/007.080.0106. S2CID 84780478.
  16. ^ Bourget, Hélène; Anderson, Jason S. (2011). "A new amphibamid (Temnospondyli: Dissorophoidea) from the Early Permian of Texas". Journal of Vertebrate Paleontology. 31 (1): 32–49. doi:10.1080/02724634.2011.539652. S2CID 85856972.
  17. ^ P.P. Skutschas; S.A. Krasnolutskii (2011). "A new genus and species of basal salamanders from the Middle Jurassic of western Siberia, Russia" (PDF). Proceedings of the Zoological Institute of the Russian Academy of Sciences. 315 (2): 167–175.
  18. ^ Robert R. Reisz; Jun Liu; Jin-Ling Li; Johannes Müller (2011). "A new captorhinid reptile, Gansurhinus qingtoushanensis, gen. et sp. nov., from the Permian of China". Naturwissenschaften. 98 (5): 435–441. Bibcode:2011NW.....98..435R. doi:10.1007/s00114-011-0793-0. PMID 21484260. S2CID 20274349.
  19. ^ Robert R. Reisz; Sean P. Modesto; Diane M. Scott (2011). "A new Early Permian reptile and its significance in early diapsid evolution". Proceedings of the Royal Society B. 278 (1725): 3731–3737. doi:10.1098/rspb.2011.0439. PMC 3203498. PMID 21525061.
  20. ^ Fischer, V.; Masure, E.; Arkhangelsky, M.S.; Godefroit, P. (2011). "A new Barremian (Early Cretaceous) ichthyosaur from western Russia". Journal of Vertebrate Paleontology. 31 (5): 1010–1025. doi:10.1080/02724634.2011.595464. hdl:2268/92828. S2CID 86036325.
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