Urocyclidae

Urocyclidae
	A live individual of
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Heterobranchia
Superorder:	Eupulmonata
Order:	Stylommatophora
Suborder:	Helicina
Infraorder:	Limacoidei
Superfamily:	Helicarionoidea
Family:	Urocyclidae; Simroth, 1889

Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).^[2]

Some species within this family make and use love darts before mating.^{[citation needed]}

Distribution[]

The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.^[3]^[4]

Taxonomy[]

The family Urocyclidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005),^[2] that follows classification of Urocyclinae by Van Goethem (1977):^[5]

subfamily Rhysotininae Schileyko, 2002
subfamily Connolly, 1925 (1912)^[6] - synonyms: Peltatinae Godwin-Austen, 1912;^[7] Trochonanininae Connolly, 1912;^[8] Trochozonitinae Iredale, 1914;^[9] Ledoulxiinae Pilsbry, 1919;^[10] Gymnarionidae Von Mol, 1970;^[11] Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
subfamily Urocyclinae Simroth, 1889
- tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
- tribe Van Goethem, 1977^[5]
- tribe Van Goethem, 1977^[5]
- tribe Van Goethem, 1977^[5]

Genera[]

Genera within the family Urocyclidae include:

unsorted

† Pickford, 2009

Subfamily Rhysotininae Schileyko, 2002

Ancey, 1887^[2]
Thomeonanina Germain, 1909 : synonym of Rhysotina Ancey, 1887 (objective synonym)

Subfamily Sheldoniinae

Binder & Tillier, 1985^[2]
Godwin-Austen, 1883
Van Mol, 1970
Pilsbry, 1919
Morelet, 1884
Degner, 1932
Pilsbry, 1919
Bourguignat, 1890
Van Mol, 1970
van Bruggen & de Winter, 1990
Van Mol, 1970 ^[11]
Van Mol, 1968
Colparion Laidlaw, 1938
Verdcourt, 1956
Pilsbry, 1919
Poirier, 1887
Preston, 1914
Verdcourt, 1998
Germain, 1912
Gymnarion Pilsbry, 1919^[2]
Bourguignat, 1885 †
Godwin-Austen, 1912
Connolly, 1939
Malagarion Tillier, 1979
Pilsbry, 1919
Godwin-Austen, 1912
Verdcourt, 1961
Verdcourt, 1998
Van Mol, 1970
Preston, 1914
Verdcourt, 1961
D. Holyoak & G. Holyoak, 2020
D. Holyoak & G. Holyoak, 2020
Herbert, 2016
Degner, 1932
Herbert, 2017
Van Mol, 1970
Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae^[2]^[12]
Verdcourt, 1961
Van Mol, 1970
Thapsia Albers, 1860
Van Mol, 1970
D. Holyoak & G. Holyoak, 2020
D. Holyoak & G. Holyoak, 2020
Van Goethem, 1975
Mousson, 1869^[2]
Trochozonites Pfeffer, 1883^[2]
Verdcourt, 1998
Verdcourtia Van Mol, 1970
Van Mol, 1970 ^[13]
Zingis E. von Martens, 1878
Pfeffer, 1883^[2]
Bourguignat, 1885:^[2] synonym of Mousson, 1869
Godwin-Austen, 1908:^[2] synonym of Sheldonia Ancey, 1887

Subfamily Urocyclinae

Tribe Dendrolimacini

Heynemann, 1868 - the type genus of the tribe Dendrolimacini^[2]

Tribe Leptichnini

Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini^[2]

Tribe Upembellini

Van Goethem, 1975
Van Goethem, 1969 - the type genus of the tribe Upembellini^[2]

tribe Urocyclini

Van Goethem, 1975
Simroth, 1888^[2]
Van Goethem, 1973
Atrichotoxon Simroth, 1910
Buettneria Simroth, 1910^[14]
Simroth, 1896
Elisolimax Cockerell, 1893
Verdcourt, 2003
Microcyclus Simroth, 1896
Van Goethem, 1975
Van Goethem, 1975
Simroth, 1888
Simroth, 1897
Van Goethem, 1975
Rowson, Paustian & Van Goethem, 2017
Trichotoxon Simroth, 1888
Rowson, Paustian & Van Goethem, 2017
Urocyclus Gray, 1864 - the type genus of the family Urocyclidae^[2]

Cladogram[]

The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade:^[3]

limacoid clade

Staffordioidea

Dyakioidea

Gastrodontoidea

	Euconulidae

	Trochomorphidae

	Gastrodontidae

	Oxychilidae

Parmacelloidea

	Trigonochlamydidae

	Parmacellidae

	Milacidae

Zonitoidea

Zonitidae

Helicarionoidea

Helicarionidae

	Ariophantidae

	Urocyclidae

Limacoidea

Vitrinidae

Boettgerillidae

	Limacidae

	Agriolimacidae

References[]

^ Simroth H. (1899). "Beiträge zur Kenntniss der Nacktschnecken". Nova Acta, Kaiserlich Leopoldinish-Carolinische Deutsche Akademie der Naturforscher (= Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum) 54(1): 1-91, plates 1-4. p. 62.
^ Jump up to: ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
^ Jump up to: ^a ^b Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
^ Herbert, Dai; Kilburn, Dick (2004). Field guide to the land snails and slugs of eastern south Africa (1st ed.). Pietermaritzburg :Natal Museum: Natal Museum. ISBN 0-620-32415-5.
^ Jump up to: ^a ^b ^c ^d Van Goethem J. (1977). "Révision systématique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)". Musée Royal de l'Afrique Centrale , Annales, Sciences zoologiques 218: 355 pp., 4 plates.
^ Connolly (1925). Annals and Magazine of Natural History (9)15: 467.
^ Godwin-Austen H. H. (1912). "A review of South-African Land-Mollusca belonging to the Family Zonitidae". Annals and Magazine of Natural History (8)9: 122-139. Plate 1-7. p. 124.
^ Connolly (1912). Annals of the South African Museum 11(3): 101.
^ Iredale (1914). Proceedings of the Malacological Society of London 11(2): 122.
^ Pilsbry H. A. (1919). Bulletin of the American Museum of Natural History 40: 245.
^ Jump up to: ^a ^b Van Mol J. J. (1970). "Révision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie - systématique - zoogéographie". Annales Musée Royal de l’Afrique Centrale, Sciences Zoologiques 180: 1-231.
^ Herbert D. G. (2004). Sheldonia puzeyi. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.
^ de Winter A. J. (1987). "Verrucarion demeryi, a new semi-slug from Sierra Leone (Mollusca: Pulmonata: Urocyclidae)". Basteria 51(4-6): 87-93. abstract.
^ Simroth H. (1910). In: Voeltzkow Reise O.-Afr., Wiss. Ergebn. 2(5): 611.

External links[]

Media related to Urocyclidae at Wikimedia Commons

[1] Simroth H. (1899). "Beiträge zur Kenntniss der Nacktschnecken". Nova Acta, Kaiserlich Leopoldinish-Carolinische Deutsche Akademie der Naturforscher (= Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum) 54(1): 1-91, plates 1-4. p. 62.

[Bouchet_2005-2] Jump up to: ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.

[Hausdorf_2000-3] Jump up to: ^a ^b Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.

[Herbert&Kilburn2004-4] Herbert, Dai; Kilburn, Dick (2004). Field guide to the land snails and slugs of eastern south Africa (1st ed.). Pietermaritzburg :Natal Museum: Natal Museum. ISBN 0-620-32415-5.

[Van_Goethem_1977-5] Jump up to: ^a ^b ^c ^d Van Goethem J. (1977). "Révision systématique des Urocyclinae (Mollusca, Pulmonata, Urocyclidae)". Musée Royal de l'Afrique Centrale , Annales, Sciences zoologiques 218: 355 pp., 4 plates.

[6] Connolly (1925). Annals and Magazine of Natural History (9)15: 467.

[7] Godwin-Austen H. H. (1912). "A review of South-African Land-Mollusca belonging to the Family Zonitidae". Annals and Magazine of Natural History (8)9: 122-139. Plate 1-7. p. 124.

[8] Connolly (1912). Annals of the South African Museum 11(3): 101.

[9] Iredale (1914). Proceedings of the Malacological Society of London 11(2): 122.

[10] Pilsbry H. A. (1919). Bulletin of the American Museum of Natural History 40: 245.

[Van_Mol_1970-11] Jump up to: ^a ^b Van Mol J. J. (1970). "Révision des Urocyclidae (Mollusca, Gastropoda, Pulmonata). Anatomie - systématique - zoogéographie". Annales Musée Royal de l’Afrique Centrale, Sciences Zoologiques 180: 1-231.

[12] Herbert D. G. (2004). Sheldonia puzeyi. 2006 IUCN Red List of Threatened Species. Downloaded on 7 August 2007.

[13] Winter A. J. (1987). "Verrucarion demeryi, a new semi-slug from Sierra Leone (Mollusca: Pulmonata: Urocyclidae)". Basteria 51(4-6): 87-93. abstract.

[14] Simroth H. (1910). In: Voeltzkow Reise O.-Afr., Wiss. Ergebn. 2(5): 611.

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