2020 in arthropod paleontology

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2020 in arthropod palentology is a list of new arthropod fossil taxa, including arachnids, crustaceans, insects, trilobites, and other arthropods that were announced or described, as well as other significant arthropod paleontological discoveries and events which occurred in 2020.

Arachnids[]

Newly named taxa[]

Name Novelty Status Authors Age Type locality Country Notes Images

[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider of uncertain phylogenetic placement, possibly related to the . The type species is A. praecursor.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family Epedanidae. Genus includes new species B. xiai.
[3] Gen. et sp. nov In press Hu et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A whip spider. Genus includes new species B. dunlopi.

Burmesarchaea bilongapophyses[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Archaeidae.

[1]

Gen. et sp. nov

Junior homonym

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Nemesiidae. The type species is B. sordida. The generic name is preoccupied by Burmesia Healey (1908); Wunderlich (2021) coined a replacement name Burmesiana.[4]

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A tree trunk spider.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Oonopidae.

Burmorchestina prominens[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Oonopidae.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider, probably related to trogloraptorids.

Burmorsolus longembolus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider, probably related to trogloraptorids.

Burmorsolus longibulbus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider, probably related to trogloraptorids.

Consteniusi[5]

Gen. et sp. nov

Valid

Downen & Selden

Eocene

Kishenehn Formation

 United States

An orb-weaver spider. The type species is C. leonae.

[6]

Gen. et sp. nov

Valid

Whalen & Selden

Carboniferous (Pennsylvanian)

Energy Shale

 United States

A member of Ricinulei. Genus includes new species C. bohemondi.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Cymbioblemma hamoembolus[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Electroblemma caula[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

Electroblemma pinnae[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family . Genus includes new species E. tuberculatus.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae.

[7] Sp. nov In press Arillo, Subías & Peñalver Cretaceous  Spain A mite belonging to the group Oribatida and to the family Caleremaeidae
[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family Epedanidae. Genus includes new species G. nilsi.

[8]

Gen. et sp. nov

Valid

Jiang & Li in Jiang et al.

Late Cretaceous

Burmese amber

 Myanmar

A spider belonging to the group Araneomorphae and to the new family . The type species is G. bian.

[5]

Gen. et sp. nov

Valid

Downen & Selden

Eocene

Kishenehn Formation

 United States

An orb-weaver spider. The type species is G. pamelae.

[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the superfamily Scytodoidea and the family . The type species is H. opilionoides.

[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Probably a tube-dwelling spider. The type species is M. tuber.

[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the group Laniatores and to the new family . Genus includes new species M. oehmkuehnlei.
[9] Gen. et sp. nov Valid Müller et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Schizomida. Genus includes new species M. groehni.
[10] Sp. nov In press García-Villafuerte & Valdez-Mondragón Miocene (Aquitanian) Mexican amber  Mexico A species of Modisimus

[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Nemesiidae. The type species is M. glaber.

[11] Sp. nov Carbuccia et al. Eocene (Ypresian) Oise amber  France A member of the family Archaeidae.
[12] Sp. nov Valid Zmudzinski Eocene Baltic amber Europe (Baltic Sea region) A mite, a species of Neophyllobius.
[12] Sp. nov Valid Zmudzinski Eocene Baltic amber Europe (Baltic Sea region) A mite, a species of Neophyllobius.
[13] Gen. et sp. nov Valid Guo et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A spider belonging to the family Lagonomegopidae. Genus includes new species O. titan.
[14] Sp. nov Valid Porta et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A mite belonging to the family Bdellidae.
[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones belonging to the family . Genus includes new species P. thilolebi.

[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family . The type species is P. hamulus.

[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Ctenizidae. The type species is P. parvula.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to family Zarqaraneidae.

[8]

Gen. et sp. nov

Valid

Jiang & Li in Jiang et al.

Late Cretaceous

Burmese amber

 Myanmar

A spider belonging to the group Araneomorphae and to the new family . The type species is P. ju.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family .

Priscaleclercera furcata[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Psilodercidae.

Priscaleclercera hamo[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Psilodercidae.

Priscaleclercera liber[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Psilodercidae.

[15] Gen. et sp. nov Valid Porta et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A pseudoscorpion belonging to the family Cheiridiidae and the subfamily . Genus includes new species P. judsoni.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family .

Propterpsiloderces cymbioseta[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Eopsilodercidae.

Propterpsiloderces duplex[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Eopsilodercidae.

[1]

Gen. et sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A tarantula. The type species is P. spinipes.

[2] Gen. et sp. nov In press Bartel et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of Opiliones, possibly belonging to the family . Genus includes new species P. nalae.

[1]

Sp. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family .

[16] Gen. et sp. nov In press Martine et al. Permian (Guadalupian)  Brazil A scorpion. Genus includes new species S. itayma.

[1]

Gen. et 3 sp. et comb. nov

Valid

Wunderlich & Müller

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A spider belonging to the family Tetrablemmidae. The type species is U. gracilicornis; genus also includes new species U. brevicornis and U. longicornis, as well as "Eogamasomorpha" unicornis Wunderlich (2017).

Crustaceans[]

Newly named taxa[]

Malacostracans[]

Name Novelty Status Authors Age Type locality Country Notes Images
[17] Gen. et sp. nov Valid Pasini et al. Early Pleistocene  Italy A member of the family Goneplacidae. The type species is A. tyrsenorum.

[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family Pylochelidae.

[19] Gen. et 2 sp. nov Valid Nyborg et al. Eocene to Oligocene  Canada
 United States
A crab belonging to the family Raninidae. Genus includes new species A. blandi and A. multispinata.

[20]

Gen. et sp. nov

Valid

Pasini, Luque & Garassino

Pleistocene

 Italy

A member of Xanthoidea sensu lato, of uncertain phylogenetic placement within that group. Genus includes new species A. collinsi.

[21]

Gen. et sp. nov

Valid

Audo & Furrer

Early Jurassic

  Switzerland

A member of Polychelida, possibly belonging to the family Coleiidae. Genus includes new species A. oberlii.

[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family .

[22]

Sp. nov

Valid

Ando in Ando et al.

Late Cretaceous (Cenomanian)

 Japan

A raninoid crab.

[23]

Gen. et sp. nov

Valid

Hegna & Lazo-Wasem in Hegna et al.

Early Miocene

Mexican amber

 Mexico

A member of Amphipoda belonging to the family Talitridae. Genus includes new species C. bousfieldi.

[24]

Sp. nov

Valid

Wallaard et al.

Miocene (Burdigalian)

 Portugal

A species of Calappilia.

[25] Sp. nov Valid Hyžný Miocene  Hungary
 Poland
 Slovakia
A species of Callianassa.
[26] Sp. nov Valid Ando et al. Latest Pleistocene to Holocene Moeshima Silt Bed  Japan A member of Axiidea belonging to the family .

[27]

Sp. nov

In press

Hernáez, Buchmann & Santana

Late Pleistocene

 Brazil

A species of Callichirus.

[28] Sp. nov Valid Bahman, Feldmann & Schweitzer Miocene St. Marys  United States A species of Cancer
[28] Sp. nov Valid Bahman, Feldmann & Schweitzer Miocene St. Marys  United States A species of Cancer
[29] Gen. et sp. nov Valid Marangon & De Angeli Oligocene (Rupelian)  Italy A member of the family Homolidae. The type species is C. italica.

[30]

Gen. nov

Valid

Artal & van Bakel

Eocene (Ypresian)

 Spain

A crab belonging to the family Xanthidae.

[31] Gen. et sp. nov Valid Pasini, Poore & Garassino Late Cretaceous (Cenomanian)  Lebanon A ghost shrimp belonging to the family . Genus includes new species C. levantina.
[32] Gen. et sp. nov In press Ossó, van Bakel & Ferratges Early Cretaceous (Aptian)  Spain A crab belonging to the family . Genus includes new species C. jadeae.

[33]

Sp. nov

Valid

Ferratges, Zamora & Aurell

Late Eocene

 Spain

[34]

Sp. nov

Valid

Marangon & De Angeli

Oligocene (Rupelian)

Ligure-Piemontese Basin

 Italy

A species of Dardanus.

[35] Sp. nov Valid Jakobsen et al. Paleocene (Danian)  Denmark A species of Dardanus.
[36] Sp. nov Valid Ferratges, Zamora & Aurell Eocene (Bartonian)  Spain A hermit crab, a species of Diogenes.
[37] Gen. et sp. nov Valid Schädel et al. Late Triassic (Carnian)  Austria A member of Isopoda belonging to the group . The type species is D. aschauerorum.
[38] Sp. nov Valid Ossó et al. Eocene (Priabonian)  Spain A crab belonging to the family Dynomenidae.
[39] Sp. nov Valid Klompmaker et al. Late Jurassic (Oxfordian)  Poland A crab belonging to the family .
Eodromites cristinarobinsae[39] Sp. nov Valid Klompmaker et al. Early Cretaceous (Albian)  Spain A crab belonging to the family Goniodromitidae.

[40]

Gen. et sp. nov

Valid

Gašparič et al.

Late Triassic (Carnian)

 Slovenia

A member of the family Polychelidae. Genus includes new species E. cornuaureus.

Eryma moriedaorum[22]

Sp. nov

Valid

Ando et al.

Late Cretaceous (Cenomanian)

Enokuchi Formation

 Japan

A member of the family Erymidae.

[41]

Sp. nov

Valid

Ossó & Moreno-Bedmar

Early Cretaceous (Aptian)

 Spain

A crab belonging to the family Etyiidae.

[42]

Sp. nov

In press

Luque in Luque et al.

Miocene

 Mexico

A crab.

[39] Gen. et comb. nov Valid Klompmaker et al. Late Jurassic  Austria
 Czech Republic
 Germany
 Romania
A crab belonging to the family . The type species is "Prosopon" aculeatum von Meyer (1857); genus also includes "Prosopon" abbreviatum Schweitzer & Feldmann (2009).
[17] Sp. nov Valid Pasini et al. Early Pleistocene  Italy A species of Galathea.

[43]

Gen. et sp. nov

In press

Jarzembowski et al.

Early Cretaceous (Hauterivian)

Weald

 United Kingdom

A member of Amphipoda. Genus includes new species G. vonki.

[37] Gen. et sp. nov Valid Schädel et al. Middle Triassic (Anisian) Lower Muschelkalk Vossenveld Formation  Netherlands A member of Isopoda belonging to the group Cymothoida. The type species is G. winterswijkensis.

[33]

Sp. nov

Valid

Ferratges, Zamora & Aurell

Late Eocene

 Spain

Glebocarcinus doii[44]

Sp. nov

Valid

Karasawa & Takahashi

Early Miocene

 Japan

A member of the family Cancridae

[45]

Sp. nov

Valid

Vega in Vega et al.

Late Cretaceous (Maastrichtian)

Arroyo Grande Formation

 Cuba

A crab belonging to the family Dynomenidae.

Hefriga schlechtingerae[46]

Sp. nov

Valid

Winkler

Late Jurassic (Tithonian)

Solnhofen

 Germany

A caridean shrimp

[47]

Sp. nov

Valid

Van Bakel et al.

Early Cretaceous (Albian)

Gault

 United Kingdom

A member of Galatheoidea.

Hoploparia echinata[48]

Sp. nov

Valid

Pinheiro et al.

Late Cretaceous (SantonianCampanian)

Santa Marta Formation

Antarctica

Hoploparia hinokunika[22]

Sp. nov

Valid

Ando & Shimada in Ando et al.

Late Cretaceous (Cenomanian)

Enokuchi Formation

 Japan

A lobster.

Hoploparia tectumque[49]

Sp. nov

In press

Fraaije et al.

Late Cretaceous (Cenomanian)

 Kazakhstan

[50]

Sp. nov

In press

Vega in Vega et al.

Late Cretaceous (Maastrichtian)

 Cuba

A crab belonging to group Portunoidea and the family . Announced in 2020; the final version of the article naming it is scheduled to be published in 2021.

[51]

Gen. et sp. nov

Valid

Fraaije et al.

Eocene

 Italy

A hermit crab belonging to the family . Genus includes new species J. bolcensis.

[52]

Gen. et comb. nov

In press

Van Bakel et al.

Late Jurassic (Kimmeridgian)

 Germany

A crab belonging to the family Necrocarcinidae. The type species is "" angulosum Wehner (1988).

[53]

Sp. nov

Valid

Starzyk

Late Jurassic (Oxfordian)

 Poland

A crab belonging to the family .

[54]

Gen. et sp. nov

In press

Van Bakel et al.

Early Cretaceous (Aptian)

 Russia

A crab belonging to the family . The type species is L. cornutus.

[33]

Sp. nov

Valid

Ferratges, Zamora & Aurell

Late Eocene

 Spain

[55]

Sp. nov

Valid

Karasawa and Kobayashi

Pleistocene

 Japan

A member of the family Hexapodidae.

[56]

Gen. et sp. nov

Valid

Artüz & Sakınç

Late Pleistocene

 Turkey

A member of Cumacea belonging to the family Nannastacidae. Genus includes new species M. samimei.

[18]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

 Czech Republic

A hermit crab belonging to the family .

[57]

Sp. nov

Valid

Davis, Garassino & Weaver

Middle Eocene

Castle Hayne

 United States

A member of the family Matutidae.

[58] Gen. et sp. nov Valid Ossó, Gagnaison & Bailleul Eocene (Ypresian)  Morocco A member of the family Carpiliidae. The type species is M. binodosus.
[59] Sp. nov Valid Gašparič, Robins & Gale Late Jurassic (Kimmeridgian/Tithonian)  Slovenia A member of the family Galatheidae.

Mesoparapylocheles janetjacksonae[18]

Sp. nov

Valid

Fraaije et al.

Late Jurassic (Tithonian)

 Czech Republic

A hermit crab belonging to the family .

[44] Gen. et comb. nov Valid Karasawa in Karasawa & Takahashi EarlyMiddle Miocene  Japan A member of the family Cancridae. Monotypic with the type species, Imaizumi, 1962.

[52]

Gen. et sp. nov

In press

Van Bakel et al.

Early Cretaceous (Berriasian)

Štramberk Limestone

 Czech Republic

A crab belonging to the family Necrocarcinidae. The type species is M. stramberkensis.

Munidopsis kaedetatsuyai [60]

Sp. nov

Valid

Karasawa

Miocene

 Japan

A member of the family Munidopsidae.

[61] Sp. nov Valid De Angeli & Garassino Early Eocene  Italy A species of Neoliomera.
[37] Gen. et sp. nov Valid Schädel et al. Late Triassic (Carnian)  Austria A member of Isopoda belonging to the group Scutocoxifera. The type species is O. summesbergeri.

[62]

Sp. nov

Valid

Karasawa

Early Miocene

 Japan

A member of the family Orithyiidae.

[34]

Sp. nov

Valid

Marangon & De Angeli

Oligocene (Rupelian)

Ligure-Piemontese Basin

 Italy

A species of Paguristes.

[35] Sp. nov Valid Jakobsen et al. Paleocene (Danian)  Denmark A species of Paguristes.
[63] Sp. nov Valid Wallaard et al. Miocene (Tortonian)  Cyprus A species of Paguristes.

[34]

Sp. nov

Valid

Marangon & De Angeli

Oligocene (Rupelian)

Ligure-Piemontese Basin

 Italy

A species of Paguristes.

[64]

Gen. et 2 sp. et comb. nov

Valid

Fraaije et al.

Late Jurassic to Late Cretaceous (Cenomanian)[65]

 Czech Republic
 France
 Lebanon[65]

A furry lobster. Genus includes new species P. culocervus and P. jbeilensis,[65] as well as "Palinurus" strambergensis Bachmayer (1959).

[66] Sp. nov Jauvion et al. Middle Jurassic (Callovian) La Voulte-sur-Rhône  France A member of Polychelida

[54]

Gen. et 2 sp. et comb. nov

In press

Van Bakel et al.

Cretaceous (Aptian to Cenomanian)

Hudspeth Formation

 Russia
 United States

A crab belonging to the superfamily Dorippoidea and the family . The type species is P. kalashnikovi; genus also includes new species P. levashiensis, as well as "" connori Nyborg et al. (2019).

[67] Sp. nov Valid Garassino & Nyborg in Garassino et al. Late Cretaceous (Santonian) Nanaimo Group  Canada A species of Petrolisthes.
[68] Sp. nov Valid Audo, Schweigert & Charbonnier Late Jurassic (Oxfordian)  France
[68] Sp. nov Valid Audo, Schweigert & Charbonnier Middle Jurassic (Callovian) Antarctica
[39] Sp. nov Valid Klompmaker et al. Late Jurassic (Tithonian) Ernstbrunn Limestone  Austria A crab belonging to the family Prosopidae.
[69] Fam. et Gen. et sp. nov In press Heard et al. Early Cretaceous (Aptian) Sierra Madre  Mexico A member of Tanaidacea belonging to the group , in the new family . Genus includes new species P. espinalensis.

[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family Paguridae.

Protopagurus duopupae[18]

Sp. nov

Valid

Fraaije et al.

Early Cretaceous (Berriasian)

 Czech Republic

A hermit crab belonging to the family Paguridae.

[70] Sp. nov Valid Charbonnier & Audo Early Jurassic (Sinemurian)  France A member of Astacidea belonging to the family
[71] Sp. nov Valid Nyborg, Garassino & Ross Late Cretaceous (late Campanian)  Canada A dromioid crab
[72] Sp. nov Valid Feldmann, Schweitzer & Haggart Late Cretaceous Haslam  Canada A member of the family Erymidae.
Stenodactylina devillezi[73] Sp. nov Valid Schweigert & Härer Late Jurassic (Kimmeridgian) Nusplingen  Germany A member of the family Erymidae.
Stenodactylina geigerae[73] Sp. nov Valid Schweigert & Härer Late Jurassic (Tithonian) Usseltal  Germany A member of the family Erymidae.
[39] Sp. nov Valid Klompmaker et al. Late Jurassic (Tithonian) Ernstbrunn Limestone  Austria A crab belonging to the family .
Tanidromites weinschenki[39] Sp. nov Valid Klompmaker et al. Late Jurassic (Kimmeridgian) Oberjura-Massenkalk Formation  Germany A crab belonging to the family Tanidromitidae.

[74]

Sp. nov

Valid

Nyborg & Garassino

Miocene

Astoria

 United States

A crab belonging to the family Cyclodorippidae.

[74]

Sp. nov

Valid

Nyborg & Garassino

Miocene

 United States

A crab belonging to the family Cyclodorippidae.

[75]

Sp. nov

Valid

Hyžný in Hyžný et al.

Pliocene

 Azores

A species of Upogebia. Announced in 2020; the final version of the article naming it was published in 2021.

[76]

Sp. nov

Valid

Ando

Late Eocene

 Japan

A ghost shrimp belonging to the family .

[54]

Sp. nov

In press

Van Bakel et al.

Early Cretaceous (Aptian)

Kubina Formation

 Russia

A crab belonging to the family .

Ostracods[]

Name Novelty Status Authors Age Type locality Country Notes Images
[77] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
[78] Sp. nov Valid Nazik et al. Late Devonian  Mongolia A member of Podocopida belonging to the superfamily and the family .
[79] Gen. et 2 sp. nov Valid Melnikova Ordovician  Russia Genus includes new species A. demissa and A. aenigma.
[80] Gen. et sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Podocopida belonging to the family Bairdiidae. The type species is A. formosa.
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
[82] Sp. nov Valid Cabral, Lord & Pinto in Cabral et al. Early Jurassic (Pliensbachian and Toarcian)  Portugal A member of the family Bairdiidae.
[83] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian) Kössen Formation  Austria
 Romania
A member of Podocopida belonging to the family Bairdiidae.
[80] Sp. nov Valid Gliwa in Gliwa et al. Late Permian and Early Triassic (Changhsingian–Griesbachian)  Iran A member of Podocopida belonging to the family Bairdiidae.
[78] Sp. nov Valid Nazik et al. Late Devonian  Mongolia A member of Platycopida belonging to the superfamily and the family .
[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian)  Tunisia
Bythocypris? multagracilis[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
Bythocypris? pythagorasi[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Aptian) Orbata  Tunisia
Candonopsis carthaginensis[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere fekihi[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere giovannae[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Capsacythere salaji[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
[80] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Palaeocopida belonging to the family .
[80] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Platycopida belonging to the family .
Cavellina hairapetiani[80] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Platycopida belonging to the family Cavellinidae.
[83] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Podocopida belonging to the family Bairdiidae.
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Chrysocythere gliozziae[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Chrysocythere russoi[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
[86] Sp. nov Valid Yasuhara et al. Early Miocene  India
[87] Sp. nov Valid Gonta Ordovician  Russia
[87] Sp. nov Valid Gonta Ordovician  Russia
[88] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous  Brazil
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A species of Cytherella.
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family .
[89] Sp. nov Valid Forel in Forel, Kolar-Jurkovšek & Jurkovšek Late Triassic (Carnian)  Slovenia A member of Podocopida belonging to the family Bairdiidae.
[79] Gen. et sp. nov Valid Melnikova Ordovician  Russia Genus includes new species D. nana.
[79] Sp. nov Valid Melnikova Ordovician  Russia A member of the family .
[90] Sp. nov Valid Lord, Cabral & Danielopol Early Cretaceous (Albian)  Germany A member of Podocopida belonging to the family Cytheridae
[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
[87] Sp. nov Valid Gonta Ordovician  Russia
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Dorukella nevioi[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Dorukella razgallahae[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
Dorukella ruggeroi[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
[80] Sp. nov Valid Gliwa in Gliwa et al. Permian (Wuchiapingian–Changhsingian)  Iran A member of Podocopida belonging to the superfamily .
[80] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Podocopida belonging to the family Bairdiidae.
[79] Sp. nov Valid Melnikova Ordovician  Russia
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
[79] Sp. nov Valid Melnikova Ordovician  Russia A member of the group belonging to the family .
Hastacypris[88] Gen. et sp. nov Junior homonym Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil Genus includes new species H. adamantem. The generic name is preoccupied by Hastacypris Croneis & Gutke (1939).
[91] Sp. nov Valid Durak, Akkiraz & Nazik Early Miocene Aspiras Basin  Turkey A species of Heterocypris. Announced in 2020; the final version of the article naming it was published in 2021.
[83] Gen. et sp. et comb. nov Valid Forel in Forel & Grădinaru Late Triassic (Carnian) to Middle Jurassic (Bathonian)  Austria
 France
 Germany
 India
 Italy
 Romania
  Switzerland
A member of Podocopida belonging to the family Bairdiidae. The type species is H. pontuseuxinusensis; genus also includes "Anchistrocheles"? spinosa Sheppard in Brand (1990), "Anchistrocheles"? tuningensis Beher (2004) and "Anchistrocheles" gemmellaroi Crasquin in Crasquin et al. (2018).
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family .
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Healdiidae.
[80] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Myodocopida belonging to the family Cylindroleberididae.
[77] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
Ilyocypris? arca[88] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil
Ilyocypris? latanodi[88] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil
[83] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Podocopida belonging to the family Bairdiidae.
[87] Sp. nov Valid Gonta Ordovician  Russia
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Cytheruridae.
[79] Sp. nov Valid Melnikova Ordovician  Russia A member of the family .
[82] Sp. nov Valid Cabral, Pinto & Lord in Cabral et al. Early Jurassic (Toarcian)  Portugal
 United Kingdom
A member of the family Protocytheridae.
[92] Sp. nov Valid Hearing et al. Ordovician (Katian)  Vietnam
[79] Gen. et 2 sp. nov Valid Melnikova Ordovician  Russia A member of the family . Genus includes new species K. conspicua and K. taimyrica.
[79] Sp. nov Valid Melnikova Ordovician  Russia A member of the family .
Laccochilina (Prochilina) yanpingensis[93] Sp. nov In press Zhang Ordovician  China
[94] Sp. nov Valid Alivernini in Alivernini et al. Miocene (Messinian) and Pliocene Zhada Basin  China A member of the family Limnocytheridae.
[80] Sp. nov Valid Gliwa in Gliwa et al. Late Permian and Early Triassic (Changhsingian–Griesbachian)  Iran A member of Podocopida belonging to the superfamily Bairdioidea.
[87] Gen. et sp. nov Valid Gonta Ordovician  Russia Genus includes new species L. risus
[90] Gen. et 2 sp. et comb. nov Valid Lord, Cabral & Danielopol Middle Jurassic (Aalenian and Bajocian)  Germany

 Netherlands
 United Kingdom

A cytherid Podocopida. The type species is M. macroporosa;
genus also includes new species M. angulata,
as well as "" maculosa Bate (1963) and "Dolocythere" tuberculata Luppold (2012).
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Limnocytheridae.
[95] Sp. nov In press Choi et al. Cretaceous  China
[96] Gen. et sp. nov Valid Wang et al. Late Cretaceous (Cenomanian) Burmese amber  Myanmar A member of the family Candonidae belonging to the subfamily . Genus includes new species M. hui.
[87] Sp. nov Valid Gonta Ordovician  Russia
[87] Sp. nov Valid Gonta Ordovician  Russia
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis decimai[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis guerneti[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis hendae[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis khayatii[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
Okadaleberis memmiae[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Trachyleberididae.
[80] Sp. nov Valid Gliwa in Gliwa et al. Permian (Changhsingian)  Iran A member of Podocopida belonging to the family Bairdiidae.
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Cytheridae.
[86] Gen. et comb. nov Valid Yasuhara et al. Early Miocene  India A new genus for "" gujaratensis Tewari & Tandon (1960)
[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Aptian) Orbata  Tunisia
[83] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Cypridoidea belonging to the family .
[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Aptian) Orbata  Tunisia
Perissocytheridea kouminiensis[84] Sp. nov In press Trabelsi et al. Early Cretaceous (Albian) Orbata  Tunisia
[83] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of the family Bairdiidae.
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
[89] Sp. nov Valid Forel in Forel, Kolar-Jurkovšek & Jurkovšek Late Triassic (Carnian)  Slovenia A member of Cypridoidea belonging to the family Pontocyprididae.
[87] Gen. et sp. nov Valid Gonta Ordovician  Russia Genus includes new species P. guttaformis.
[97] Sp. nov Valid Carignano, Echevarría & Zavattieri Middle Triassic Cuyo Basin  Argentina A member of .
[77] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
[83] Sp. nov Valid Forel in Forel & Grădinaru Late Triassic (Rhaetian)  Romania A member of Cypridoidea belonging to the family Pontocyprididae.
[79] Sp. nov Valid Melnikova Ordovician  Russia A member of the family .
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
[81] Sp. nov Crasquin et al. Late Triassic  Italy A member of the family Bairdiidae.
[88] Gen. et sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil Genus includes new species P. latavectis.
[88] Sp. nov In press Santos Filho, Fauth & Sames Early Cretaceous Orós Formation  Brazil
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Cytherettidae.
[87] Sp. nov Valid Gonta Ordovician  Russia
[77] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
[98] Sp. nov In press Warne Late Miocene and Pliocene  Australia
[79] Gen. et sp. nov Valid Melnikova Ordovician  Russia A member of the family . Genus includes new species T. rotundatus.
[77] Sp. nov Valid Becker, Braun & Pratt Devonian  Canada
[85] Sp. nov In press Sciuto & Temani in Sciuto, Temani & Ammar Miocene (Messinian)  Tunisia A member of Podocopa belonging to the family Hemicytheridae.
[99] Sp. nov Valid Bergue, Maranhao & Ng Permian  Brazil

Other crustaceans[]

Name Novelty Status Authors Age Type locality Country Notes Images

[100]

Sp. nov

Valid

Buckeridge et al.

Oligocene and Miocene

 Slovenia

A barnacle.

[101]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[101]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[102]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Ireland
 Russia
 United States?

A member of Cyclida. Genus includes A. capidulum (Chernyschev, 1933), A. simulans (Reed, 1908) and possibly A. ? minutus (Rogers, 1902).

[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
Angulotergum milviformis[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
Angulotergum robustum[103] Sp. nov In press Gale Cretaceous  United States A barnacle.

[104]

Sp. nov

Valid

Karasawa

Miocene

 Japan

A member of the family Scalpellidae.

[104]

Sp. nov

Valid

Karasawa

Miocene

 Japan

A member of the family Scalpellidae.

[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
[105] Sp. nov Valid Gale in Gale et al. Late Pleistocene  India A barnacle

[106]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[107] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
[107] Sp. nov Valid Gale Cretaceous A barnacle

[102]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 United Kingdom

A member of Cyclida belonging to the family . Genus includes B. rankini (Woodward, 1868), B. scotti (Woodward, 1894) and B. testudo (Peach, 1882).

[102]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Ireland
 United Kingdom

A member of Cyclida. Genus includes C. wrighti (Woodward, 1870).

[101]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[102]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Tajikistan

A member of Cyclida. Genus includes C. spinosus (Chernyschev, 1933).

[107] Gen. et 3 sp. nov Valid Gale Cretaceous  Morocco
 Tunisia
A barnacle. Genus includes new species C. aitlaminensis, C. robustus and C. tunisiensis.

[101]

Gen. et sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle. Genus includes new species C. buckeridgei.

[101]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[106]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[106]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[108] Sp. nov In press Gale, Jagt & Goolaerts Late Cretaceous (Maastrichtian)  Tunisia A barnacle.
[109] Gen. et sp. nov Valid Luo et al. Middle Jurassic Daohugou  China A member of Anostraca. Genus includes new species D. cheni.
[103] Sp. nov In press Gale Cretaceous  United States A barnacle.

[102]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

Mazon Creek fossil beds

 United States

A member of Cyclida belonging to the family Americlidae. Genus includes D. obesus (Schram, Vonk & Hof, 1997).

[107] Sp. nov Valid Gale Early Cretaceous (Albian)  France A barnacle
[107] Sp. nov Valid Gale Late Cretaceous (Campanian)  United Kingdom A barnacle
[110] Sp. nov Valid Gale TithonianBerriasian Portland Group  United Kingdom A barnacle
[107] Gen. et comb. nov Valid Gale Cretaceous  United Kingdom A brachylepadid barnacle. The type species is "Pollicipes" fallax Darwin (1851)

[101]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[111] Sp. nov Valid Teng & Li Early Jurassic (PliensbachianToarcian)  China A clam shrimp. Announced in 2020; the final version of the article naming was published in 2021.
[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
[107] Gen. et comb. et 3 sp. nov Valid Gale Cretaceous  United Kingdom
 United States
A barnacle. The type species is "Pollicipes" striatus Darwin (1851); genus also includes "Pollicipes" filosus Withers (1911), as well as new species J. africanum, J. spinosum and J. luteorum.[103]
[112] Gen. et sp. nov In press Gallego et al. Early Cretaceous La Huérguina  Spain An afrograptid clam shrimp. Genus includes new species L. multicostata.

[102]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Ireland
 United Kingdom
 United States?

A member of Cyclida. Genus includes L. bilobatus (Woodward, 1870), L. jonesianus (Woodward, 1870), L. torosus (Woodward, 1870) and possibly also L. ? communis (Rogers, 1902) and L. ? permarginatus (Rogers, 1902).

[113] Sp. nov Valid Gale Eocene (Lutetian)  France A barnacle. Announced in 2020; the final version of the article naming it was published in 2021.
[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
Loriculina rubrifluvia[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
[110] Sp. nov Valid Gale TithonianBerriasian  United Kingdom A barnacle

[101]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[106]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[107] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
[107] Sp. nov Valid Gale Cretaceous A barnacle

[101]

Sp. nov

Valid

Gale

Indian Ocean (Mascarene Plateau)

A barnacle.

[107] Gen. et 3 sp. et comb. nov Valid Gale Cretaceous  United Kingdom A barnacle. The type species is S. elegans; genus also includes new species S. bromleyi and S. barringtonensis, as well as "Scalpellum (Titanolepas)" martini Withers (1935).
[107] Gen. et sp. nov Valid Gale Cretaceous A barnacle. The type species is S. holtwilsoni.

[102]

Gen. et comb. nov

Valid

Schweitzer, Mychko & Feldmann

Carboniferous

 Japan

A member of Cyclida. Genus includes T. tazawai (Niko & Ibaraki, 2011).

[103] Gen. et sp. nov In press Gale Cretaceous  United States A barnacle. Genus includes new species T. holterhoffi.
[113] Gen. et comb. et sp. nov Valid Gale Eocene and Oligocene Barton
Bouldnor
Headon Hill
Selsey
 United Kingdom A barnacle. The type species is "Balanus" unguiformis Sowerby (1846); genus also includes "Balanus" erisma Sowerby (1846) and a new species V. mortoni. Announced in 2020; the final version of the article naming it was published in 2021.
[107] Gen. et 3 sp. nov Valid Gale Cretaceous  Morocco A barnacle. Genus includes new species V. aboudaensis, V. peakei and V. hancocki.
[107] Sp. nov Valid Gale Cretaceous A barnacle
[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
[107] Sp. nov Valid Gale Cretaceous  Tunisia A barnacle
[107] Sp. nov Valid Gale Cretaceous A barnacle
[103] Sp. nov In press Gale Cretaceous  United States A barnacle.
[107] Sp. nov Valid Gale Cretaceous  United Kingdom A barnacle
[107] Sp. nov Valid Gale Cretaceous A barnacle
[107] Gen. et comb. nov Valid Gale Cretaceous  France

 United Kingdom

A barnacle. The type species is " aptiensis Withers (1935); genus also includes "Cretiscalpellum" matrioni Gale (2019).

General research[]

  • Redescription and a study on the phylogenetic relationships of grimmi, based on data from new fossil material from the Solnhofen Limestone (Germany), is published by Pazinato et al. (2020).[114]
  • A hermit crab belonging to the family Pylochelidae preserved in a shell of an ammonite belonging to the species Craspedites nekrassovi will be described from the Upper Jurassic of Moscow, Russia by Mironenko (2020).[115]
  • A putative lobster Tricarina gadvanensis from the Early Cretaceous of Iran is reinterpreted as a seroloid isopod by Hyžný et al. (2020).[116]
  • A study on the phylogenetic relationships of Eocarcinus praecursor is published by Scholtz (2020).[117]
  • A study on anatomical adaptations that enabled an ecological transition into the water column in Silurian myodocopes, focusing on myodocopids from the Herefordshire Lagerstätte (United Kingdom), is published by Siveter, Perrier & Williams (2020).[118]
  • A study comparing sexual dimorphism in ostracods from the Gulf and Atlantic coastal plain before and after the Cretaceous–Paleogene extinction event is published by Martins et al. (2020).[119]
  • Redescription of Americlus rankini and a study on the paleoecology of this taxon is published by Clark et al. (2020).[120]

Insects[]

Trilobites[]

New taxa[]

Name Novelty Status Authors Age Type locality Country Notes Images

[121]

Sp. nov

Valid

Chatterton

Cambrian (Jiangshanian)

 Canada

Arduennella hainauensis[122]

Sp. nov

Valid

Van Viersen & Alberti

Devonian

[123] Gen. et sp. nov Valid Adrain, Karim & McAdams Ordovician (Darriwilian) Table Cove Formation  Canada A calymenine trilobite. The type species is A. bardensis.
[124] Gen. et sp. nov Valid Basse & Lemke Devonian (Famennian) Dasberg Limestone  Germany A cyrtosymboline trilobite. Genus includes new species B. kaufmanni.
[125] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Nolichucky Formation  United States A member of the family .
Blountia morgancreekensis[125] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Riley Formation  United States A member of the family Kingstoniidae.
Blountia nevadensis[125] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian  United States A member of the family Kingstoniidae.
Blountia newfoundlandensis[125] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Shallow Bay Formation  Canada A member of the family Kingstoniidae.
Blountia tennesseensis[125] Sp. nov Valid Armstrong, Westrop & Eoff Cambrian Nolichucky Formation  United States A member of the family Kingstoniidae.

[126]

Gen. et comb. nov

Valid

Geyer

Cambrian

 Italy

A new genus for "" courtessolei Pillola (1991).

[127]

Sp. nov

Valid

Peel

Cambrian (Miaolingian)

 Greenland

A member of the family Alokistocaridae.

[121]

Sp. nov

Valid

Chatterton

Cambrian (Jiangshanian)

 Canada

[128]

Sp. nov

Valid

Alberti

Devonian

 Germany

A member of Proetida belonging to the family .

Cyphaspides speculator[128]

Sp. nov

Valid

Alberti

Devonian

 Germany

A member of Proetida belonging to the family Aulacopleuridae.

Dicranurus webbyi[129]

Sp. nov

Valid

Holloway, Smith & Thomas

Ordovician (Katian)

 Australia

An odontopleurid trilobite.

Ehmaniella sermersuaqensis[127]

Sp. nov

Valid

Peel

Cambrian (Miaolingian)

Telt Bugt Formation

 Greenland

A member of the family Alokistocaridae.

Elrathia hensonensis[130]

Nom. nov

Valid

Geyer & Peel

Cambrian

Ekspedition Bræ Formation

 Greenland

A replacement name for Elrathia groenlandica Geyer & Peel (2017).

[124] Sp. nov Valid Basse & Lemke Devonian (Famennian) Wocklum Limestone  Germany

[126]

Gen. et sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of the family . Genus includes new species E. pustulata.

[131]

Gen. et 2 sp. nov

Valid

Crônier et al.

Devonian (Famennian)

 Mongolia

A phacopid trilobite. Genus includes new species F. mongoliensis and F. khovdensis. Announced in 2020; the final version of the article naming was published in 2021.

Gravicalymene bakeri[132]

Sp. nov

In press

Smith & Ebach

Ordovician (Katian)

Gordon Group

 Australia

A calymenid trilobite.

[131]

Sp. nov

Valid

Crônier et al.

Devonian (Famennian)

 Mongolia

A phacopid trilobite. Announced in 2020; the final version of the article naming was published in 2021.

[133]

Sp. nov

Valid

Geyer, Pais & Wotte

Cambrian

Jbel Wawrmast Formation

 Morocco

A member of Redlichiida belonging to the family .

[126]

Sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of the family Despujolsiidae.

Marsaisia? lemdadensis[126]

Sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of the family Despujolsiidae.

[134]

Gen. et sp. nov

Valid

Sun, Zeng & Zhao

Cambrian (Wuliuan)

 China

A member of Ptychopariacea of uncertain phylogenetic placement. The type species is P. auritus.

[135]

Gen. et 2 sp. nov

Valid

Liñán et al.

Cambrian Stage 3

Pusa Formation

 Spain

Genus includes new species P. toletana and P. vidalii.

[129]

Gen. et sp. nov

Valid

Holloway, Smith & Thomas

Ordovician (Katian)

 Australia

A calymenid trilobite. Genus includes new species P. jonesi.

[136]

Sp. nov

Valid

Wernette et al.

Cambrian (Furongian)

 Thailand

[137]

Gen. et sp. nov

Valid

Wernette et al.

Late Cambrian

Ao Mo Lae Formation

 Thailand

A kaolishaniid trilobite belonging to the new subfamily . Genus includes new species S. molaensis.

Saukianda? dresnayi[138]

Sp. nov

In press

Geyer & Landing

Cambrian

MoroccanAlgerian border region

[139]

Sp. nov

Valid

Van Viersen & Taghon

Devonian (Emsian)

Our Formation

 Belgium

A member of the family Acastidae.

[140]

Sp. nov

Valid

Sundberg

Cambrian (Wuliuan)

Lakeview Limestone
Spence Shale

 United States

A member of Ptychopariida belonging to the family .

General research[]

  • A study on the timing of the trilobite extinctions at the early to middle Cambrian transition is published by Sundberg et al. (2020).[141]
  • Evidence of gregarious moulting of Cambrian (Stage 4) trilobites from the "Tsinghsutung" Formation (China), representing one of the earliest records of synchronized moulting behaviours in the fossil record, is presented by Corrales-García et al. (2020).[142]
  • A study on trilobite size variations in the Ordovician Fezouata Formation (Morocco) deposited in a cold-water environment is published by Saleh et al. (2020).[143]
  • A study on the nature of variation among oryctocephalid specimens from the Combined Metals Member of the Pioche Formation (Nevada; Cambrian Stage 4) will be published by Webster & Sundberg (2020).[144]
  • A study on the ontogeny of the olenid trilobite salteri is published by Månsson & Clarkson (2020).[145]
  • A study on the anatomy of the internal structures of the compound eye of Aulacopleura koninckii is published by Schoenemann & Clarkson (2020).[146]
  • A study on exuviae of tetrasulcatus from the Ordovician (Hubei, China) is published by Zong (2020), who reports exuviae arranged with two or three together end to end or superimposed, and evaluates the implications of these findings for the knowledge of the behavior of trilobites.[147]

Other arthropods[]

Newly named taxa[]

Name Novelty Status Authors Age Type locality Country Notes Images

Adelophthalmus pyrrhae[148]

Sp. nov

Valid

Lamsdell et al.

Carboniferous (Tournaisian)

St. Nazaire Group (possibly Lydiennes Formation)

 France

A eurypterid.

[149]

Gen. et comb. nov

Disputed

Lamsdell

Jurassic

 United Kingdom

A horseshoe crab. The type species is "Limulus" woodwardi Watson (1909). However, Bicknell et al. (2021) transferred "L." woodwardi to the genus Mesolimulus.[150]

[149]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 Germany
 United States

A member of the family Belinuridae. The type species is "Euproops" longispina Packard (1885).

[149]

Gen. et comb. nov

Valid

Lamsdell

Triassic

 Germany

A member of the family Austrolimulidae. The type species is "Paleolimulus" fuchsbergensis Hauschke & Wilde (1987).

[149]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 United States

A member of the family Austrolimulidae. The type species is "Paleolimulus" longispinus Schram (1979).

[151] Gen. et sp. nov Valid O’Flynn & Liu in O’Flynn et al. Cambrian Stage 3 Chiungchussu  China An early euarthropod of uncertain phylogenetic placement. The type species is B. yangi.

[152]

Sp. nov

Valid

Broda, Rak & Hegna

Devonian (Famennian)

 Czech Republic

A thylacocephalan.

[152]

Sp. nov

Valid

Broda, Rak & Hegna

Devonian (Famennian)

 Poland

A thylacocephalan.

[153]

Sp. nov

Jobbins, Haug & Klug

Devonian (Famennian)

 Morocco

A thylacocephalan.

[152]

Sp. nov

Valid

Broda, Rak & Hegna

Devonian (Famennian)

 United States

A thylacocephalan.

[154]

Gen. et sp. nov

Valid

Tihelka, Tian & Cai

Devonian (Pragian)

Rhynie chert

 United Kingdom

Originally described as a member of Opiliones, but this interpretation was subsequently questioned by Pérez-González & Shultz (2021), who were unsure about the phylogenetic placement of this arthropod, but noted anatomical similarities with the euthycarcinoid species .[155] The type species is D. hutchinsoni.

[156]

Gen. et sp. nov

Valid

Ji et al.

Early Triassic

 China

A thylacocephalan. Genus includes new species D. chaohuensis.

[157]

Gen. et sp. nov

Valid

Hannibal & May

Permian (Cisuralian)

Richards Spur locality

 United States
( Oklahoma)

A millipede belonging to the group Helminthomorpha. The type species is D. subtila.

[158] Gen. et sp. nov Gueriau et al. Devonian (Famennian)  Belgium A member of Euthycarcinoidea. The type species is E. potii.
[159] Sp. nov Valid Skovsted et al. Cambrian Mural  Canada A member of Bradoriida belonging to the family .

Itagnostus idahoensis[140]

Sp. nov

Valid

Sundberg

Cambrian (Wuliuan)

Emigrant Formation
Lakeview Limestone

 United States

A member of Agnostida belonging to the family Peronopsidae.

Kanoshoia[160]

Gen. et sp. nov

Valid

Lerosey‐Aubril et al.

Cambrian (Drumian)

Wheeler Shale

 United States

A megacheiran. Genus includes new species K. rectifrons.

[157]

Gen. et sp. nov

Valid

Hannibal & May

Permian (Cisuralian)

Richards Spur locality

 United States
( Oklahoma)

A millipede belonging to the group Juliformia, possibly a member of the superfamily Xyloiuloidea. The type species is K. fortsillensis.

[149]

Gen. et comb. nov

Valid

Lamsdell

Triassic

 France

A horseshoe crab. The type species is "Limulus" vicensis Bleicher (1897).

[149]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 United Kingdom

A member of the family Belinuridae. The type species is "" arcuatus Baily (1859).

Mesolimulus tafraoutensis[161]

Sp. nov

In press

Lamsdell et al.

Late Cretaceous (CenomanianTuronian)

Gara Sbaa Lagerstätte

 Morocco

A horseshoe crab.

[156]

Sp. nov

Valid

Ji et al.

Early Triassic

 China

A thylacocephalan.

[149]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous (Tournaisian)

Horton Bluff Formation

 Canada

A member of Xiphosura belonging to the family . The type species is "Paleolimulus" woodae Lerner, Lucas & Mansky (2016).

[157]

Gen. et sp. nov

Valid

Hannibal & May

Permian (Cisuralian)

Richards Spur locality

 United States
( Oklahoma)

A millipede belonging to the group Helminthomorpha. The type species is O. richardsspurense.

[149]

Gen. et comb. nov

Valid

Lamsdell

Carboniferous

 Russia
 Ukraine
 United States

A member of the family Belinuridae. The type species is "" lunatus Martin (1809); genus also includes P. iswariensis (Chernyshev, 1928), P. lacoei (Packard, 1885), P. metschetensis.. (Chernyshev, 1928) and P. stepanovi (Chernyshev, 1928).

[152]

Gen. et comb. nov

Valid

Broda, Rak & Hegna

Carboniferous (Tournaisian)

Líšeň Formation

 Czech Republic

A thylacocephalan; a new genus for "Concavicaris" viktoryni Rak, Broda & Kumpan (2018).

Parioscorpio[162]

Gen. et sp. nov

Wendruff et al.

Silurian (Telychian)

Brandon Bridge

 United States

An arthropod of uncertain phylogenetic placement. Originally classified as an early scorpion, but subsequently argued to be basal to crown group Mandibulata and Chelicerata.[163] The type species is P. venator.

[164]

Gen. et comb. nov

Valid

Bicknell & Smith

Late Devonian

 United States

A stem xiphosurid; a new genus for "Kasibelinurus" randalli. Announced in 2020; the final version of the article naming it was published in 2021.

[165]

Sp. nov

In press

Su, Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A millipede belonging to the family Polyxenidae.

[165]

Sp. nov

In press

Su, Cai & Huang

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A millipede belonging to the family Polyxenidae.

Pruemopterus[166]

Gen. et sp. nov

Valid

Poschmann

Early Devonian

 Germany

A eurypterid. Genus includes new species P. salgadoi. Announced in 2020; the final version of the article naming it was published in 2021.

[159] Sp. nov Valid Skovsted et al. Cambrian Mural  Canada A member of Bradoriida belonging to the family Hipponicharionidae.

[167]

Gen. et comb. nov

Bicknell, Naugolnykh & Brougham

Permian (Asselian)

Araukaritovaya Formation

 Ukraine

A member of Xiphosura belonging to the group ; a new genus for "Paleolimulus" jakovlevi Glushenko & Ivanov (1961).

[168]

Gen. et sp. nov

Aria et al.

Cambrian Stage 3

Maotianshan Shale Member of the Yu’anshan Formation

 China

A megacheiran related to Jianfengia and . The type species is S. maomima.

Soligorskopterus shpinevi[169]

Sp. nov

Valid

Naugolnykh & Areshin

Late Devonian

 Russia

An eurypterid. Announced in 2019; the final version of the article naming it was published in 2020.

Tariccoia tazagurtensis[170] Sp. nov Valid Pérez-Peris et al. Early Ordovician Fezouata  Morocco A nektaspid arthropod. Announced in 2020; the final version of the article naming it was published in 2021.

Thelxiope holmani[171]

Sp. nov

Valid

Lerosey-Aubril, Skabelund & Ortega-Hernández

Cambrian (Drumian)

Wheeler Shale

 United States

A possible member of Chelicerata.

[149]

Gen. et sp. nov

Valid

Lamsdell

Early Triassic

Ankitokazo Basin

 Madagascar

A horseshoe crab. The type species is V. madagascarensis.

[172]

Gen. et sp. nov

Liu et al.

Cambrian Stage 3

Chiungchussu Formation

 China

A member of the group Fuxianhuiida. The type species is X. luoi.

[173]

Gen. et sp. nov

Valid

Zeng et al.

Cambrian Stage 3

 China

A member of Hymenocarina. Genus includes new species X. chenggongensis. Announced in 2020; the final version of the article naming it was published in 2021.

General research[]

  • Exceptionally preserved specimens of Chuandianella ovata carrying a large number of small eggs are described from the Cambrian Chengjiang biota (China) by Ou et al. (2020), who compare the reproductive strategies of C. ovata and closely related Waptia fieldensis.[174]
  • Sun, Zeng & Zhao (2020) report the discovery of a member of the genus Sidneyia belonging or related to the species S. inexpectans from the Wuliuan Mantou Formation of North China, expanding known distribution of this genus.[175]
  • A putative xiphosuran Kiaeria is reinterpreted as a member of Chasmataspidida by Lamsdell (2020a).[176]
  • A study on the anatomy of head structures of , evaluating its implications for the knowledge of the phylogenetic relationships of euthycarcinoids, is published by Edgecombe et al. (2020).[177]
  • A study on heterochrony in Xiphosura, showing that clades that occupy non-marine environments show elevated rates of peramorphosis or paedomorphosis is published by Lamsdell (2020b).[178]
  • A study on the morphology of the feeding apparatuses of Eurypterida and Trigonotarbida is published by Haug (2020).[179]
  • A study aiming to determine the range of prey sizes that could be captured by sweep-feeding eurypterids is published by Hughes & Lamsdell (2020).[180]
  • A study on the anatomy and phylogenetic relationships of handlirschi is published by Schädel & Haug (2020), who interpret this taxon as a probable member of Cyclida rather than an isopod.[181]
  • Liu et al. (2020) report evidence of the presence of a three-dimensionally preserved labrum associated with the mouth opening in juvenile specimens of Leanchoilia illecebrosa.[182]

References[]

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