Cheiracanthium
Yellow sac spider | |
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C. insulanum, adult male | |
Cheiracanthium sp., Pateros, Washington | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Class: | Arachnida |
Order: | Araneae |
Infraorder: | Araneomorphae |
Family: | Cheiracanthiidae |
Genus: | Cheiracanthium C. L. Koch, 1839[1] |
Type species | |
C. punctorium (Villers, 1789)
| |
Species | |
212, see text | |
Synonyms[1] | |
|
Cheiracanthium, commonly called yellow sac spiders, is a genus of araneomorph spiders in the family Cheiracanthiidae, and was first described by Carl Ludwig Koch in 1839.[4] They are usually pale in colour, and have an abdomen that can range from yellow to beige. Both sexes range in size from 5 to 10 millimetres (0.20 to 0.39 in). They are unique among common house spiders because their tarsi do not point either outward, like members of Tegenaria, or inward, like members of Araneus), making them easier to identify. The name is a reference to the backwardly directed process on the cymbium of the male palp.[5] The species epithet is derived from the Greek Ancient Greek: χείρ, romanized: cheir, meaning "hand", and Acanthium, a genus of thorny-stemmed plants.
Venom[]
Though they are beneficial predators in agricultural fields, they are also known to be mildly venomous to humans. Painful bites may be incurred from species such as C. punctorium in Europe, C. mildei in Europe and North America, C. inclusum in the Americas, C. lawrencei in South Africa and C. japonicum in Japan.[6] Cheiracanthium venom is purportedly necrotic, and can cause pain, swelling, and lesions in humans,[6] but the necrotic nature and severity of its bite has been disputed.[7] A study of twenty confirmed Cheiracanthium bites in the United States and Australia found that none resulted in necrosis.[7]
Some of these spiders are attracted to the smell of volatiles in gasoline.[8] An unspecified Cheiracanthium species is reportedly attracted to the smell of petroleum and has caused problems by weaving webs inside the of particular models of Mazda vehicles, resulting in blockages and build-up of pressure that could potentially cause fuel leakage from the fuel tank and an increased risk of fire. Mazda issued a voluntary recall of Mazda 6 models built between 2010 and 2012, to fit them with software which would alert drivers if problems were developing.[9]
Species[]
Cheiracanthium is primarily an Old World genus, with many species found from northern Europe to Japan, from Southern Africa to India and Australia. The only known species in the New World are C. inclusum and C. mildei. While the former also occurs in Africa and Réunion, the latter is found in the Holarctic region and Argentina. They can also be found in the lower mainland of British Columbia, Canada. The genus is quite diverse in Africa and at least three or four species are known to occur in Egyptian cotton fields alone.[1] As of September 2019 it contains 212 species, found in the Caribbean, South America, Oceania, Europe, Central America, Africa, Asia, North America, and on Saint Helena:[1]
- C. abbreviatum Simon, 1878 – France, Denmark
- Simon, 1884 – Africa
- Lotz, 2007 – Yemen
- O. Pickard-Cambridge, 1885 – China (Yarkand), Karakorum
- C. africanum Lessert, 1921 – Africa, Madagascar, Réunion
- Biswas & Biswas, 2007 – India
- Lotz, 2007 – Yemen
- (Blackwall, 1859) – Madeira
- C. algarvense Wunderlich, 2012 – Portugal, Spain
- Lotz, 2014 – Madagascar
- Lotz, 2014 – Madagascar
- Lotz, 2014 – Madagascar
- Lotz, 2007 – Angola, Zimbabwe, South Africa
- C. angulitarse Simon, 1878 – Spain, France (Corsica), Italy, Hungary, Romania
- Lotz, 2014 – Madagascar
- C. annulipes O. Pickard-Cambridge, 1872 – Spain, Egypt, Israel
- Chen & Huang, 2012 – Taiwan
- Platnick, 1998 – Samoa
- O. Pickard-Cambridge, 1885 – China (Yarkand)
- Lotz, 2014 – Madagascar
- Caporiacco, 1936 – Libya
- Zhang, Zhang & Yu, 2018 – China
- Merian, 1911 – Indonesia (Sulawesi)
- (Lucas, 1846) – Algeria
- Lotz, 2015 – Congo
- L. Koch, 1873 – Indonesia (Lombok), Western Australia
- Kroneberg, 1875 – Central Asia
- Song, Feng & Shang, 1982 – China, Korea
- C. campestre Lohmander, 1944 – Sweden, Denmark, [[Central Europe]], Romania, Ukraine, Russia (Europe)
- Wunderlich, 1987 – Canary Is., Turkey, Egypt
- Barrion & Litsinger, 1995 – Philippines
- (Thorell, 1887) – Myanmar
- Simon, 1906 – India
- (Thorell, 1891) – India (Nicobar Is.)
- Rainbow, 1920 – Australia (Norfolk Is.)
- Tikader, 1975 – India
- Barrion & Litsinger, 1995 – Indonesia, Philippines
- Simon, 1890 – Chad, Yemen
- Caporiacco, 1939 – Ethiopia, Congo
- Lotz, 2007 – South Africa
- Zhang, Zhang & Yu, 2018 – China
- C. effossum Herman, 1879 – Central to eastern Europe
- C. elegans Thorell, 1875 – Europe, Turkey, Caucasus, Russia (Europe to south Siberia), Kazakhstan, Central Asia
- O. Pickard-Cambridge, 1874 – Libya, Egypt
- C. erraticum (Walckenaer, 1802) – Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Korea, Japan
- Barrion, Barrion-Dupo & Heong, 2013 – China
- Bösenberg & Strand, 1906 – Taiwan, Japan, Korea?
- Rainbow, 1920 – Australia (Norfolk Is.)
- (Lucas, 1846) – Algeria
- Zhang & Zhu, 1993 – China
- Chen, Huang, Chen & Wang, 2006 – Taiwan
- Lotz, 2015 – Gabon
- Pavesi, 1895 – Israel
- Zhang, Hu & Zhu, 1994 – China
- Chen & Huang, 2012 – Taiwan
- Lotz, 2014 – Madagascar
- Wunderlich, 2008 – Azores
- Lotz, 2015 – South Africa
- Lotz, 2014 – Madagascar
- Gong, 1983 – China
- C. fulvotestaceum Simon, 1878 – France
- L. Koch, 1873 – Samoa
- C. furculatum Karsch, 1879 – Cape Verde Is., Africa, Madagascar, Comoros
- Lotz, 2015 – Ghana
- Schmidt & Barensteiner, 2000 – China
- L. Koch, 1873 – Australia (Queensland, New South Wales)
- C. gratum Kulczyński, 1897 – Germany, Hungary
- Lotz, 2014 – Madagascar
- Schmidt & Piepho, 1994 – Cape Verde Is.
- Lotz, 2007 – Zimbabwe
- Gravely, 1931 – India
- Zhang & Zhu, 1993 – China
- C. ienisteai Sterghiu, 1985 – Romania, Albania
- Morano & Bonal, 2016 – Spain
- Thorell, 1881 – Australia (Queensland)
- C. incertum O. Pickard-Cambridge, 1869 – Sri Lanka
- C. inclusum (Hentz, 1847) – North America, Central America, Caribbean, South America. Introduced to Réunion
- (Thorell, 1891) – India (Nicobar Is.)
- C. indicum O. Pickard-Cambridge, 1874 – India, Sri Lanka
- Wang & Zhang, 2013 – China
- O. Pickard-Cambridge, 1874 – India
- C. insigne O. Pickard-Cambridge, 1874 – India, Sri Lanka, Thailand, Myanmar, Laos, China
- (Thorell, 1878) – Myanmar, Laos to China, Taiwan, Indonesia (Moluccas), Philippines
- C. insulare L. Koch, 1866 – Samoa
- C. insulare (Vinson, 1863) – Madagascar, Réunion
- O. Pickard-Cambridge, 1874 – Libya, Egypt
- Barrion & Litsinger, 1995 – Philippines
- Majumder & Tikader, 1991 – India
- Bösenberg & Strand, 1906 – China, Korea, Japan
- Lotz, 2014 – Comoros, Madagascar
- Simon, 1910 – São Tomé and Príncipe
- Wunderlich, 2008 – Azores
- Denis, 1947 – Egypt
- Lotz, 2015 – Uganda
- Lotz, 2015 – Kenya
- Lotz, 2015 – Ivory Coast, Ghana, Congo
- Majumder & Tikader, 1991 – India
- Ponomarev, 2007 – Kazakhstan
- Lotz, 2007 – Africa
- Lessert, 1921 – Ethiopia, Congo, Kenya, Tanzania, Uganda
- Merian, 1911 – Indonesia (Sulawesi)
- Lotz, 2011 – Tanzania
- Lotz, 2007 – Cameroon
- Chrysanthus, 1967 – New Guinea
- Karsch, 1879 – Russia (Sakhalin), China, Korea, Japan
- Simon, 1897 – Madagascar
- Barrion & Litsinger, 1995 – Philippines
- Barrion & Litsinger, 1995 – Philippines
- Xie, Yin, Yan & Kim, 1996 – China
- Merian, 1911 – Indonesia (Sulawesi)
- L. Koch, 1873 – Australia (Queensland), Tonga, Fiji, Vanuatu, New Caledonia
- (Thorell, 1890) – Indonesia (Sumatra)
- Xu, 1993 – China
- Lessert, 1921 – Congo, Kenya, Tanzania, Madagascar
- Lotz, 2015 – Congo
- C. macedonicum Drensky, 1921 – Italy, Slovenia, Slovakia, Hungary, Romania, Macedonia, Bulgaria
- Lotz, 2014 – Comoros, Madagascar
- Lotz, 2014 – Madagascar
- Lotz, 2007 – Nigeria
- Workman, 1896 – Singapore, Indonesia (Sumatra)
- Lotz, 2007 – Namibia, Botswana
- C. margaritae Sterghiu, 1985 – Romania
- Chrysanthus, 1967 – New Guinea
- Lotz, 2015 – Congo
- (Thorell, 1895) – India, Bangladesh, Myanmar
- Strand, 1911 – Indonesia (Aru Is.)
- C. mildei L. Koch, 1864 – Europe, North AfricaEurope, North Africa, Turkey, Middle East, Caucasus, Russia (Europe) to Central Asia. Introduced to North America, Argentina
- Merian, 1911 – Indonesia (Sulawesi)
- Lotz, 2007 – Zimbabwe, Botswana, South Africa
- L. Koch, 1875 – Africa, Saudi Arabia
- Main, 1954 – Australia (Western Australia)
- Schenkel, 1963 – Mongolia
- L. Koch, 1877 – Europe, Turkey, Caucasus, Iran
- L. Koch, 1866 – China, Taiwan, Australia to Samoa, New Hebrides, Solomon Is., French Polynesia
- (Thorell, 1895) – India, Myanmar
- Majumder & Tikader, 1991 – India, Bangladesh
- Patel & Patel, 1973 – India
- Simon, 1909 – Australia (Western Australia)
- Lotz, 2011 – Mauritania
- Zhang & Yin, 1999 – China
- C. occidentale L. Koch, 1882 – Spain (Minorca), Italy
- Zhang & Zhu, 1993 – China
- C. oncognathum Thorell, 1871 – Europe
- Rainbow, 1920 – Australia (Lord Howe Is.)
- Majumder & Tikader, 1991 – India
- (C. L. Koch, 1837) – Southern and eastern Europe, Turkey, Caucasus, Russia (Europe) to Tajikistan
- C. pennatum Simon, 1878 – Southern Europe, Romania
- Simon, 1909 – Australia (Western Australia)
- O. Pickard-Cambridge, 1873 – Europe, Turkey, Caucasus, Russia (Europe to south Siberia), Iran, Central Asia, China
- Thorell, 1899 – Ivory Coast, Nigeria, Cameroon
- Schenkel, 1963 – China
- Majumder & Tikader, 1991 – India
- Schenkel, 1963 – China
- Caporiacco, 1949 – Congo, Rwanda, Kenya
- C. punctorium (Villers, 1789) (type) – Europe, Turkey, Caucasus, Russia (Europe to south Siberia), Iran, Central Asia
- Sadana & Bajaj, 1980 – India
- Lotz, 2014 – Madagascar
- Strand, 1915 – Israel
- Lotz, 2014 – Madagascar
- (Thorell, 1897) – Myanmar, China, Indonesia
- Lotz, 2007 – Ethiopia
- Lotz, 2011 – Rwanda
- Mukhtar, 2015 – Pakistan
- Roewer, 1938 – New Guinea
- C. salsicola Simon, 1932 – France
- Patel & Reddy, 1991 – India
- Strand, 1907 – Ivory Coast to Zanzibar
- Tikader, 1962 – India
- Caporiacco, 1949 – Kenya, Rwanda, Zimbabwe, Botswana, South Africa
- L. Koch, 1864 – Mediterranean to Central Asia
- Patel & Reddy, 1991 – India
- Lotz, 2015 – Congo, Kenya
- Lotz, 2007 – South Africa
- Majumder & Tikader, 1991 – India, Bangladesh
- Rainbow, 1897 – Australia (New South Wales)
- Zhang & Yin, 1999 – China
- Thorell, 1899 – Cameroon, Nigeria
- El-Hennawy, 2001 – Egypt
- Zhang, Zhu & Hu, 1993 – China
- Merian, 1911 – Indonesia (Sulawesi)
- (Thorell, 1887) – Myanmar
- Zhang, Zhu & Hu, 1993 – China
- L. Koch, 1873 – New Zealand, Australia (Tasmania)
- L. Koch, 1879 – Russia (Middle and south Siberia)
- Simon, 1878 – Western Mediterranean
- Paik, 1990 – China, Korea, Japan
- Biswas & Raychaudhuri, 2003 – Bangladesh
- Chen, Huang, Chen & Wang, 2006 – China, Taiwan
- Biswas & Roy, 2005 – India
- Lotz, 2015 – Tanzania
- C. taprobanense Strand, 1907 – Sri Lanka
- L. Koch, 1873 – Australia (Queensland)
- Caporiacco, 1949 – Kenya
- Strand, 1911 – New Guinea
- Chen & Huang, 2012 – Taiwan
- (Thorell, 1895) – India, Myanmar
- Simon, 1906 – India
- (Thorell, 1895) – Myanmar
- Kroneberg, 1875 – Uzbekistan, Tajikistan
- Strand, 1917 – Thailand to Australia (Queensland)
- Paik, 1985 – China, Korea
- Bösenberg & Strand, 1906 – Korea, Japan, China, Laos
- Marusik & Fomichev, 2016 – Mongolia
- Lawrence, 1936 – Southern Africa
- Lotz, 2011 – Cape Verde Is.
- (Sundevall, 1833) – Europe, Caucasus, Russia (Europe to Far East), Iran, China
- O. Pickard-Cambridge, 1874 – India
- Mukhtar, 2015 – Pakistan
- Chrysanthus, 1967 – New Guinea
- (Benoit, 1977) – St. Helena
- Savelyeva, 1972 – Russia (south Siberia), Kazakhstan
- Hu & Song, 1982 – China, Korea
See also[]
- List of Cheiracanthiidae species
- List of spiders associated with cutaneous reactions
References[]
- ^ Jump up to: a b c d Gloor, Daniel; Nentwig, Wolfgang; Blick, Theo; Kropf, Christian (2019). "Gen. Cheiracanthium C. L. Koch, 1839". World Spider Catalog Version 20.0. Natural History Museum Bern. doi:10.24436/2. Retrieved 2019-10-06.
- ^ Bonaldo, A. B.; Brescovit, A. D. (1992). "As aranhas do gênero Cheiracanthium C. L. Koch, 1839 na região neotropical (Araneae, Clubionidae)". Revista Brasileira de Entomologia. 36: 732.
- ^ Lotz, L. N. (2007). "The genus Cheiracanthium (Araneae: Miturgidae) in the Afrotropical region. 1. Revision of known species". Navorsinge van die Nasionale Museum Bloemfontein. 23: 4.
- ^ Koch, C. L. (1839). Die Arachniden. C. H. Zeh'sche Buchhandlung. pp. 125–158.
- ^ Ubick, D.; et al. (2005). Spiders of North America: An Identification Manual. American Arachnological Society.
- ^ Jump up to: a b Papini, R (2012). "Documented bites by a yellow sac spider (Cheiracanthium punctorium) in Italy: a case report". Journal of Venomous Animals and Toxins Including Tropical Diseases. 18 (3): 349–354. doi:10.1590/S1678-91992012000300014.
- ^ Jump up to: a b Vetter, RS; Isbister, GK; Bush, SP; Boutin, LJ (June 2006). "Verified bites by yellow sac spiders (genus Cheiracanthium) in the United States and Australia: where is the necrosis?". The American Journal of Tropical Medicine and Hygiene. 74 (6): 1043–8. doi:10.4269/ajtmh.2006.74.1043. PMID 16760517.
- ^ "Gas-loving spider prompts Mazda recall in U.S." Reuters. March 4, 2011. Archived from the original on October 6, 2015. Retrieved June 30, 2017.
- ^ "Spider invasion prompts Mazda software fix". BBC News. 7 April 2014. Retrieved 9 April 2014.
Further reading[]
- Howell, Mike; Jenkins, Ronald L. (2004). Spiders of the US: A photographic guide. ISBN 0-536-75853-0.
External links[]
- Media related to Cheiracanthium at Wikimedia Commons
- Cheiracanthium
- Araneomorphae genera
- Cosmopolitan spiders
- Taxa named by Carl Ludwig Koch