List of non-marine molluscs of Malaysia

Location of Malaysia

The non-marine mollusks of Malaysia are a part of the molluscan wildlife of Malaysia. A number of species of non-marine mollusks are found in the wild in Malaysia.

Freshwater gastropods[]

Freshwater nerites are used in Malaysian cuisine

Ampullariidae

Pila ampullacea (Linnaeus, 1758)^[1]
Pila scutata (Mousson, 1848)^[1]
Pomacea canaliculata (Lamarck, 1819)^[1]

Nassariidae

Clea bangueyensis EA Smith, 1895^[1]
Clea nigricans A Adams, 1885^[1]

Neritidae

Neritina pulligera (Linnaeus, 1767)^[1]
Septaria porcellana (Linnaeus, 1858)^[1]
Vittina coromandeliana (Sowerby, 1836)^[1]
(Lesson, 1831)^[1]

Pachychilidae

(Thiele, 1908)^[1]

Viviparidae

(Kobelt, 1906)^[1]

Paludomidae

EA Smith, 1894^[1]
Adams, 1874^[1]

Physidae

Physella acuta (Draparnaud, 1805)^[1]

Planorbidae

Indoplanorbis exustus (Deshayes, 1834)^[1]

Thiaridae

Mieniplotia scabra (OF Müller, 1774)^[1]
Melanoides tuberculata (OF Müller, 1774)^[1]
Tarebia granifera (Lamarck, 1822)^[1]

Land gastropods[]

Assimineidae

Marzuki, Liew & Mohd-Azlan, 2021^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
Acmella nana Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
(Heude, 1882)^[2]

Cyclophoridae

(Godwin-Austen, 1889)^[2]
(Sowerby, 1843)^[4]
Cyclophorus aurantiacus pernobilis Gould, 1844^[4]
Cyclophorus cantori (Benson, 1851)^[4]
Cyclophorus expansus (Pfeiffer, 1853)^[4]
Cyclophorus malayanus (Benson, 1852)^[4]
(Metcalfe, 1851)^[2]
Cyclophorus pfeifferi Reeve, 1861^[4]
Cyclophorus volvulus (O.F. Müller, 1774)^[4]
Vermeulen, Liew & Schilthuizen, 2015^[3]
(E. A. Smith, 1895)^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
(L. Pfeiffer, 1855)^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
(Issel, 1874)^[2]
(Godwin-Austen, 1889)^[2]
(E. A. Smith, 1895)^[2]
(L. Pfeiffer, 1851)^[2]
(Metcalfe, 1851)^[2]
(Mousson, 1849)^[2]
(L. Pfeiffer, 1854)^[2]
(Godwin-Austen, 1889)^[2]
(L. Pfeiffer, 1856)^[2]
(Godwin-Austen, 1889)^[2]
(Metcalfe, 1851)^[2]

Diplommatinidae

Maassen, 2008^[5]
Marzuki & Foon, 2016^[5]
(H. Adams & A. Adams, 1851)^[2]
Marzuki, 2019^[6]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Godwin-Austen, 1889^[2]^[6]
H. Adams, 1872^[2]
Vermeulen & Khalik, 2021^[7]
Godwin-Austen, 1889^[2]
Sykes, 1903^[6]
Laidlaw, 1949^[2]
E. A. Smith, 1893^[7]
Fulton, 1901^[2]
Godwin-Austen, 1889^[2]
Vermeulen, 1993^[2]
Laidlaw, 1949^[7]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, 1991^[2]
Thompson, 1978^[2]
(Vermeulen, 1991)^[2]
Vermeulen, 1991^[2]
Vermeulen, 1991^[2]
Vermeulen, 1994^[2]
Vermeulen, 1991^[2]
(E. A. Smith, 1894)^[2]
(E. A. Smith, 1893)^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
(E. A. Smith, 1893)^[2]
(Vermeulen, 1994)^[2]
Ancey, 1887^[2]

Pupinidae

Godwin-Austen, 1889^[2]
Godwin-Austen, 1889^[2]
Godwin-Austen, 1889^[2]
(L. Pfeiffer, 1855)^[2]
Issel, 1874^[2]

Hydrocenidae

Khalik, Hendriks, Vermeulen & Schilthuizen, 2018^[8]
Khalik, Hendriks, Vermeulen & Schilthuizen, 2018^[2]^[8]
E. A. Smith, 1895^[2]
Thompson & Dance, 1983^[8]
Godwin-Austen, 1889^[2]^[8]
Godwin-Austen, 1889^[2]
Khalik, Hendriks, Vermeulen & Schilthuizen, 2018^[8]
Gredler, 1902^[8]
Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
Khalik, Hendriks, Vermeulen & Schilthuizen, 2018^[8]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Godwin-Austen, 1889^[8]
Thompson & Dance, 1983^[8]
Georissa saulae (van Benthem-Jutting, 1966)^[8]
(van Benthem-Jutting, 1966)^[8]
Khalik, Hendriks, Vermeulen & Schilthuizen, 2018^[8]
Khalik, Hendriks, Vermeulen & Schilthuizen, 2018^[8]

Camaenidae

Fulton, 1896^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Amphidromus cf. similis Pilsbry, 1900^[2]
Bradybaena similaris (Férussac, 1822)^[2]
(L. Pfeiffer, 1854)^[2]
Tan, Chan & Foon, 2017^[9]
Foon, Tan & Clements, 2015^[9]
Clements & Tan, 2012^[9]
Landouria winteriana (L. Pfeiffer, 1842)^[2]
(Benson, 1859)^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]

Chronidae

Kaliella barrakporensis (L. Pfeiffer, 1852)^[2]
(E. A. Smith, 1895)^[2]
(Gould, 1852)^[2]
(L. Pfeiffer, 1846)^[2]
(Mousson, 1865)^[2]
(Mousson, 1857)^[2]
Möllendorff, 1897^[2]
(Cox, 1871)^[2]
Platymma tweediei (Tomlin, 1938)^[10]

Endodontidae

(C. Semper, 1874)^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
(Aldrich, 1889)^[2]
(E. A. Smith, 1893)^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]

Euconulidae

Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]

Punctidae

Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]

Charopidae

^[2]

Trochomorphidae

Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
(Metcalfe, 1851)^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
(Martens, 1864)^[2]
(Martens, 1867)^[2]

(A. Adams & Reeve, 1848)^[2]

Helicarionidae

Godwin-Austen, 1891^[2]

Euconulidae

(E. A. Smith, 1893)^[2]

Clausiliidae

(L. Pfeiffer, 1854)^[2]

Valloniidae

(C. B. Adams, 1845)^[2]
(Stoliczka, 1873)^[2]
(Van Benthem-Jutting, 1940)^[2]
(Issel, 1874)^[2]

Veronicellidae

Laevicaulis alte (Férussac, 1821)^[11]
(Issel, 1874)^[11]
(Heynemann, 1885)^[11]^[2]

Rathouisiidae

Collinge 1902^[11]

Ariophantidae

Collinge, 1901^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
(A. Adams & Reeve, 1850)^[2]
(L. Pfeiffer, 1854)^[2]
(Godwin-Austen, 1891)^[2]
(Issel, 1874)^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]
Vermeulen, Liew & Schilthuizen, 2015^[3]^[2]
Godwin-Austen, 1891^[2]
Ibycus rachelae Schilthuizen & Liew, 2008^[11]
Collinge & Godwin Austen, 1895^[11]
Collinge & Godwin Austen, 1895^[11]
Parmarion martensi Simroth, 1893^[11]
(Collinge, 1899)^[11]
(Metcalfe, 1851)^[2]

Limacidae

Deroceras laeve (Müller, 1774)^[11]

Philomycidae

(Van Hasselt, 1824)^[11]
Laidlaw 1937^[11]
(Stoliczka, 1872)^[11]

Subulinidae

Janssen & Witte, 2002^[12]
Vermeulen, Liew & Schilthuizen, 2015^[3]

Alycaeidae

Foon & Liew, 2017^[13]
Möllendorff, 1902^[13]
Alycaeus balingensis Tomlin, 1948^[13]^[9]
Maassen, 2006^[13]
Foon & Liew, 2017^[13]
Foon & Liew, 2017^[13]
Fulton, 1902^[13]
Foon & Liew, 2017^[13]
Foon & Liew, 2017^[13]
Stoliczka, 1872^[13]
Foon & Liew, 2017^[13]
Morgan, 1885^[13]
Morgan, 1885^[13]
Sykes, 1902^[13]
Foon & Liew, 2017^[13]
Preston, 1907^[13]
Crosse, 1879^[13]
Foon & Liew, 2017^[13]
Möllendorff, 1894^[13]
Foon & Liew, 2017^[13]
Foon & Liew, 2017^[13]
Morgan, 1885^[13]
Foon & Liew, 2017^[13]
(Godwin-Austen, 1889)^[2]
(H. Adams, 1870)^[2]
(Godwin-Austen, 1889)^[2]
(E. A. Smith, 1895)^[2]

Vertiginidae

F. G. Thompson & Dance, 1983^[2]
Hypselostoma megaphonum van Benthem-Jutting, 1950^[14]
Hypselostoma perigyra van Benthem-Jutting, 1950^[15]
Paraboysidia serpa van Benthem-Jutting, 1950^[9]

Dyakiidae

Godwin-Austen, 1891^[2]
E. A. Smith, 1895^[2]
(H. Adams, 1870)^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Marzuki, Liew & Mohd-Azlan, 2021^[2]
Rhinocochlis nasuta (Metcalfe, 1851)^[2]
Quantula striata (Gray, 1834)^[16]^[2]

Achatinellidae

(Möllendorff, 1897)^[2]

Achatinidae

Lissachatina fulica (Bowdich, 1822)^[2]
Allopeas clavulinum (Potiez & Michaud, 1838)^[2]
Allopeas gracile (T. Hutton, 1834)^[2]
(Westerlund, 1883)^[2]
Paropeas achatinaceum (L. Pfeiffer, 1846)^[2]

Diapheridae

Laidlaw, 1950^[2]
Sinoennea Kobelt, 1904^[17]

Streptaxidae

Gulella bicolor (T. Hutton, 1834)^[2]
Tomlin, 1948^[9]

Succineidae

(Martens, 1867)^[2]

Freshwater bivalves[]

References[]

^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r Ng TH, Dulipat J, Foon JK, Lopes-Lima M, Zieritz A, Liew T-S (2017) A preliminary checklist of the freshwater snails of Sabah (Malaysian Borneo) deposited in the BORNEENSIS collection, Universiti Malaysia Sabah. ZooKeys 673: 105–123. DOI:10.3897/zookeys.673.12544
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad ^ae ^af ^ag ^ah ^ai ^aj ^ak ^al ^am ^an ^ao ^ap ^aq ^ar ^as ^at ^au ^av ^aw ^ax ^ay ^az ^ba ^bb ^bc ^bd ^be ^bf ^bg ^bh ^bi ^bj ^bk ^bl ^bm ^bn ^bo ^bp ^bq ^br ^bs ^bt ^bu ^bv ^bw ^bx ^by ^bz ^ca ^cb ^cc ^cd ^ce ^cf ^cg ^ch ^ci ^cj ^ck ^cl ^cm ^cn ^co ^cp ^cq ^cr ^cs ^ct ^cu ^cv ^cw ^cx ^cy ^cz ^da ^db ^dc ^dd ^de ^df ^dg ^dh ^di ^dj ^dk ^dl ^dm ^dn ^do ^dp ^dq ^dr Marzuki ME, Liew T-S, Mohd-Azlan J (2021) Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1–113. https://doi.org/10.3897/zookeys.1035.60843
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad ^ae ^af ^ag ^ah ^ai ^aj ^ak ^al ^am ^an ^ao ^ap ^aq ^ar ^as ^at ^au ^av Vermeulen JJ, Liew TS, Schilthuizen M (2015) Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species. ZooKeys 531: 1–139. doi: 10.3897/zookeys.531.6097
^ ^a ^b ^c ^d ^e ^f ^g DO Duc Sang, DO Van Nhuong (2019). "Family Cyclophoridae in Vietnam (Gastropoda: Cyclophoroidea): the genus Cyclophorus Montfort, 1810". Ruthenica 29(1): 1-53.
^ ^a ^b Marzuki, M.E.b.; Foon, J.K. (2016). "A new land snail, Arinia (Notharinia) micro (Caenogastropoda: Cyclophoroidea: Diplommatinidae), from a limestone karst in Perak, Peninsular Malaysia". Raffles Bulletin of Zoology. 64: 313–318.
^ ^a ^b ^c Mohammad Effendi bin Marzuki (2019). "Diplommatina azlani, a new land snail species from Sarawak (Gastropoda: Cyclophoroidea: Diplommatinidae)". Raffles Bulletin of Zoology 67: 56–59. DOI: 10.26107/RBZ-2019-0004
^ ^a ^b ^c Jaap J. Vermeulen & Mohd Zacaery Khalik (2021). "Two new species of Diplommatina Benson, 1849 (Gastropoda, Caenogastropoda: Diplommatinidae) from Borneo". 29(3): 174–177. DOI: https://doi.org/10.12657/folmal.029.019
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m Mohd Zacaery Khalik, Kasper Hendriks, Jaap J. Vermeulen, Menno Schilthuizen (2018). "A molecular and conchological dissection of the “scaly” Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)". ZooKeys 773: 1-55. https://doi.org/10.3897/zookeys.773.24878
^ ^a ^b ^c ^d ^e ^f Tan SK, Chan SY & Foon JK (2017) Description of a new Kenyirus species (Pulmonata, Camaenidae) from Kedah, Peninsular Malaysia. Occasional Molluscan Papers, 6: 1–4.
^ Tomlin, J.R. le B. (1938). "A new Ariophantid from the Malay". Proceedings of the Malacological Society of London. 23: 116-117.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m M. Schilthuizen & T. S. Liew. (2008). "The slugs and semislugs of Sabah, Malaysian Borneo (Gastropoda, Pulmonata: Veronicellidae, Rathouisiidae, Ariophantidae, Limacidae, Philomycidae)". Basteria 72: 287-306.
^ Janssen, R. & Witte, V. (2002). "Allopeas myrmekophilos n. sp., the first snail reported as living in army ant colonies (Gastropoda: Pulmonata: Subulinidae)". Archiv für Molluskenkunde 131(1-2): 211-215. DOI: 10.1127/arch.moll/131/2002/211
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w Junn Kitt Foon, Thor-Seng Liew. (2017). "A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia". ZooKeys 692: 1-81. https://doi.org/10.3897/zookeys.692.14706
^ Clements R. (2007). Hypselostoma megaphonum. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.3. <www.iucnredlist.org>. Downloaded on 14 September 2010.
^ Clements R. (2007). Hypselostoma perigyra. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.3. <www.iucnredlist.org>. Downloaded on 14 September 2010.
^ Daston M. M. & Copeland J. (1993). "The luminescent organ and sexual maturity in Dyakia striata". Malacologia 35(1): 9–19. abstract
^ Sutcharit C., Naggs F., Wade C. M., Fontanilla I. & Panha S. (2010). "The new family Diapheridae, a new species of Diaphera Albers from Thailand, and the position of the Diapheridae within a molecular phylogeny of the Streptaxoidea (Pulmonata: Stylommatophora)". Zoological Journal of the Linnean Society 160: 1-16. doi:10.1111/j.1096-3642.2009.00598.x.

External links[]

The Terrestrial Gastropods of Sabah (North Borneo), Malaysia
Issel A. (1874). Molluschi Borneensi. Genova, Tipografia del R. Instituto Sordo-Muti.

[Ng,Dulipat,Foon,Lopes-Lima,Zieritz&Liew2017-1] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r Ng TH, Dulipat J, Foon JK, Lopes-Lima M, Zieritz A, Liew T-S (2017) A preliminary checklist of the freshwater snails of Sabah (Malaysian Borneo) deposited in the BORNEENSIS collection, Universiti Malaysia Sabah. ZooKeys 673: 105–123. DOI:10.3897/zookeys.673.12544

[Marzuki,Liew&Mohd-Azlan2021-2] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad ^ae ^af ^ag ^ah ^ai ^aj ^ak ^al ^am ^an ^ao ^ap ^aq ^ar ^as ^at ^au ^av ^aw ^ax ^ay ^az ^ba ^bb ^bc ^bd ^be ^bf ^bg ^bh ^bi ^bj ^bk ^bl ^bm ^bn ^bo ^bp ^bq ^br ^bs ^bt ^bu ^bv ^bw ^bx ^by ^bz ^ca ^cb ^cc ^cd ^ce ^cf ^cg ^ch ^ci ^cj ^ck ^cl ^cm ^cn ^co ^cp ^cq ^cr ^cs ^ct ^cu ^cv ^cw ^cx ^cy ^cz ^da ^db ^dc ^dd ^de ^df ^dg ^dh ^di ^dj ^dk ^dl ^dm ^dn ^do ^dp ^dq ^dr Marzuki ME, Liew T-S, Mohd-Azlan J (2021) Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species. ZooKeys 1035: 1–113. https://doi.org/10.3897/zookeys.1035.60843

[Vermeulen,Liew&Schilthuizen2015-3] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad ^ae ^af ^ag ^ah ^ai ^aj ^ak ^al ^am ^an ^ao ^ap ^aq ^ar ^as ^at ^au ^av Vermeulen JJ, Liew TS, Schilthuizen M (2015) Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species. ZooKeys 531: 1–139. doi: 10.3897/zookeys.531.6097

[Do&Do2019-4] ^ ^a ^b ^c ^d ^e ^f ^g DO Duc Sang, DO Van Nhuong (2019). "Family Cyclophoridae in Vietnam (Gastropoda: Cyclophoroidea): the genus Cyclophorus Montfort, 1810". Ruthenica 29(1): 1-53.

[Ariniamicro-5] Marzuki, M.E.b.; Foon, J.K. (2016). "A new land snail, Arinia (Notharinia) micro (Caenogastropoda: Cyclophoroidea: Diplommatinidae), from a limestone karst in Perak, Peninsular Malaysia". Raffles Bulletin of Zoology. 64: 313–318.

[Marzuki2019-6] Mohammad Effendi bin Marzuki (2019). "Diplommatina azlani, a new land snail species from Sarawak (Gastropoda: Cyclophoroidea: Diplommatinidae)". Raffles Bulletin of Zoology 67: 56–59. DOI: 10.26107/RBZ-2019-0004

[Vermeulen&Khalik2021-7] Jaap J. Vermeulen & Mohd Zacaery Khalik (2021). "Two new species of Diplommatina Benson, 1849 (Gastropoda, Caenogastropoda: Diplommatinidae) from Borneo". 29(3): 174–177. DOI: https://doi.org/10.12657/folmal.029.019

[Khalik,Hendriks,Vermeulen&Schilthuizen2018-8] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m Mohd Zacaery Khalik, Kasper Hendriks, Jaap J. Vermeulen, Menno Schilthuizen (2018). "A molecular and conchological dissection of the “scaly” Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)". ZooKeys 773: 1-55. https://doi.org/10.3897/zookeys.773.24878

[Tan,Chan&Foon2017-9] ^ ^a ^b ^c ^d ^e ^f Tan SK, Chan SY & Foon JK (2017) Description of a new Kenyirus species (Pulmonata, Camaenidae) from Kedah, Peninsular Malaysia. Occasional Molluscan Papers, 6: 1–4.

[10] Tomlin, J.R. le B. (1938). "A new Ariophantid from the Malay". Proceedings of the Malacological Society of London. 23: 116-117.

[Schilthuizen_&_Liew,_2008-11] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m M. Schilthuizen & T. S. Liew. (2008). "The slugs and semislugs of Sabah, Malaysian Borneo (Gastropoda, Pulmonata: Veronicellidae, Rathouisiidae, Ariophantidae, Limacidae, Philomycidae)". Basteria 72: 287-306.

[12] Janssen, R. & Witte, V. (2002). "Allopeas myrmekophilos n. sp., the first snail reported as living in army ant colonies (Gastropoda: Pulmonata: Subulinidae)". Archiv für Molluskenkunde 131(1-2): 211-215. DOI: 10.1127/arch.moll/131/2002/211

[Foon_&_Liew,_2017-13] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w Junn Kitt Foon, Thor-Seng Liew. (2017). "A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia". ZooKeys 692: 1-81. https://doi.org/10.3897/zookeys.692.14706

[14] Clements R. (2007). Hypselostoma megaphonum. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.3. <www.iucnredlist.org>. Downloaded on 14 September 2010.

[15] Clements R. (2007). Hypselostoma perigyra. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.3. <www.iucnredlist.org>. Downloaded on 14 September 2010.

[16] Daston M. M. & Copeland J. (1993). "The luminescent organ and sexual maturity in Dyakia striata". Malacologia 35(1): 9–19. abstract

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