Rotzo Formation
Rotzo Formation Stratigraphic range: Early-Late Pliensbachian ~ | |
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Type | Geological formation |
Unit of | |
Sub-units | Tovel Member[1] |
Underlies | Calcari Grigi di Noriglio Formation |
Overlies | |
Lithology | |
Primary | Lagoonal or restricted shallow subtidal; lithified, gray, silty marl. Paralic; ooidal, gray grainstone and bioturbated, intraclastic, ooidal, gray wackestone. Subtidal flat with mud banks and sand deposits.[2] |
Other | Light-grey to yellowish-grey packstone, with oolites, bioclasts, algal lumps, pellets, dasycladacean algae, foraminifera, lituolids, and miliolids |
Location | |
Coordinates | 45°42′N 11°06′E / 45.7°N 11.1°ECoordinates: 45°42′N 11°06′E / 45.7°N 11.1°E |
Approximate paleocoordinates | 32°06′S 16°42′E / 32.1°S 16.7°E |
Region | Veneto |
Country | Italy |
Type section | |
Named for | Rotzo |
Rotzo Formation (Italy) |
The Rotzo Formation is a geological formation in Italy, dating to roughly between 189 and 183 million years ago and covering the Pliensbachian stage of the Jurassic Period in the Mesozoic Era.[3] Has been traditionally classified as a Sinemurian-Pliensbachian Formation, but a large and detailed dataset of isotopic 13C and 87Sr/86Sr data, estimated the Rotzo Formation to span only over the whole Pliensbachian.[4] The Rotzo Formation represented the , being located over the Trento Platform and surrounded by the (marginal calcarenitic bodies), the (condensed deposits and emerged lands), the and (open marine), and finally towards the south, deep water deposits of the Adriatic Basin.[5]
Fossil prosauropod tracks have been reported from the formation.[6] This formation was deposited within a tropical lagoon environment which was protected by oolitic shoals and bars from the open deep sea located to the east () and towards the west (). It is characterized by a rich paleontological content. It is notable mostly thanks to its great amount of big aberrant bivalves, among which is the genus , described in the second half of the nineteenth century. The unusual shape of Lithiotis and shells, extremely elongated and narrow, characterized by a spoon-like body space placed in a high position, rarely preserved, seems to suggest their adaptation to soft and muddy bottoms with a high sedimentation rate.[7] The Bellori outcrop displays about 20 m of limestones with intercalated clays and marls rich in organic matter and sometimes fossil wood (coal) and amber. The limestones are well stratified, with beds 10 cm to more than one metre thick, whereas the clayey levels range between 3 and 40 cm in thickness.[8][9]
Invertebrata[]
Microfossils of the Rotzo Formation consist of benthic foraminifera, , Ostracoda and coprolites. Foraminifera are mainly benthic agglutinated species belonging to the superfamily (suborder Textulariina), while lamellar and porcellaneous-walled species are very rare.[10] The bivalve Opisoma excavatum is very common.[11]
Ichnofossils[]
In the Western Venetian Prealps a shallow-water, oceanic carbonate platform system, the Trento platform, developed on the Early Jurassic, producing a large succession of massive to well-bedded white Limestones, several 100 m thick that are part of the Calcari Grigi Group, where the Rotzo Formation is the Upper Member.[12] On the local limestone of the Rotzo Formation deep burrowing is a very common type of biogenic activity, as is shown due to the presence of a large characteristic network of burrows which reach down to the lagoonal, marly-clayey assigned strata, suggesting intense bioturbation by large unknown organisms, perhaps giant decapod crustaceans (Probably members of the family Erymidae), although, the burrows found are not closely related to the ones of Shrimps or other decapods, but resemble those of Stomatopoda and Malacostraca.[12] Other includes abandoned burrows, vertical biogenic action and infilling on the sea substrate.[12]
Genus | Species | Stratigraphic position | Material | Notes | Images |
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Campomolon, Valbona |
Burrowing and track Ichnofossils |
Thalassinoides suevicus has been found on mostly of the middle-upper part of the Rotzo Formation associated with muddy deposits. It ranges from 2–5 cm to 6–10 cm and the larger ones from 10 to 16 cm.[13] Y-shaped tunnels that seen in cross-section reveal circular walls made of pelletoidal grainstone, being more probably a fodichnia of a burrowing animal.[15] A few ichnofossils include simple cylindrical tubes up to 80 cm in length, that resemble crustacean described in Seychelles.[15] |
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Campomolon, Valbona |
Burrowing and track Ichnofossils |
Two major types of Ophiomorpha where recovered, a smaller one from 2–4 cm in size and the larger one from 5–15 cm in diameter.[15] They are complex burrow systems lined with pelletoidal sediments generally infilled by coarse-grained detritus.[13] Specimens Seems partly destroyed by weathering.[14] |
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Campomolon, Valbona |
Burrowing and track Ichnofossils |
In the Rotzo Formation Ophiomorpha irregulaire local specimens the walls are extensively reworked by small, secondary burrowers assigned to the ichnogenus Chondrites.[14] Interpreted as the feeding burrow of a sediment-ingesting animal. |
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Campomolon, Valbona |
Infilled abandoned burrows by coarse-grained skeletal debris |
Ichnofossils done by organisms advancing along the bottom surface. Very narrow, vertical or subvertical, slightly winding unlined shafts filled with mud. Locally, post hurricane burrows are found in fine-grained tempestite beds and muddy layers and they are Domichnia, Fodinichnia and .[12] |
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Campomolon, Valbona |
Infilled abandoned burrows by coarse-grained skeletal debris |
On the local waters during the Lower Jurassic, water motion due to the hurricane action truncated many mounds causing changes on the deposition on the sea-floor and inducing various phases of substrate infillings with carbonate mud, fine-to coarse-grained skeletal debris and fecal pellets.[12] They are assigned to Priapulida, Serpulidae, Siboglinidae, Sabellidae or even Oweniidae. |
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Campomolon, Valbona |
Vertical burrows with preserved entrances |
It is difficult to suggest this ichnogenus because on the Formation the vertical and lined burrow with a deep central crater typical of Chomatichnus is never preserved.[12] It resemble described burrows of endobenthic thalassinidean decapods, specially Callianassa subterranea of modern North Sea, Callianassa major, Callianassa californensis or Upogebia pugettensis.[12] It can be also Serpulidae Polychaetan burrows. |
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Coste dell’Anglone dinosaur ichnosite |
Star-shaped impressions |
A ichnogenus that represents the resting trace resting activity of sea stars (Asteroidea) and brittle stars (Ophiuroidea).[16] The recovered from the Rotzo formation are probably from specimens trapped on tidal changes.[16] |
Bivalves[]
The Rotzo Formation is know mostly due to its massive bivalve associations of the genera , and that extended all along the Pliensbachian Trento Platform forming mass accumulations of specimens that formed Reef-Like structures.[17] This fauna appeared after the early Pliensbachian C-cycle perturbation, that triggered the diffusion of the Lithiotis Fauna, noted on the rapid widespread of this biota after the event layers.[17] All of the genera related with this fauna appeared on the lower Jurassic, and all but one became extinct before the Middle Jurassic.[18] This "Reefs" had an strong zonation, starting with the bivalves and , restricted to intertidal and shallow-subtidal facies. is limited to lagoonal subtidal facies and even in some low-oxygen environments. Finally and are found in subtidal facies, constructing buildups.[18] This sections formed various kinds of ecosystems on the Trento platform, where it appeared in branched corals filled with (), Domal corals (), tubular corals, corals, unidentified colonial corals, regular echinoid debris, sponges, and the solitary coral sp., with also aggregated snail shells.[18]
Genus | Species | Stratigraphic position | Material | Notes | Images |
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Isolated Shells |
A clam, member of inside . The so-called Eomiodon horizon represents the lower Rotzo Formation, composed of organic-rich marlstones with abundant specimens of this genus, typical of stressed environment with low salinity.[19] This genus considered an opportunistic shallow infaunal suspension feeder, and the marker genus for brackish environments.[21] |
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Isolated Shells |
A clam, member of Astartidae inside Carditida. Is considered a genus that evolved from shallow burrowing ancestors, becoming a secondarily semi-infaunal edgewise recliner adapted to photosymbiosis.[11] |
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Isolated Shells |
A clam, Incertade sedis inside Pterioida. On the Rotzo formation this byssate bivalves indicate a shallow subtidal or intertidal environment.[22] |
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A clam, Incertade sedis inside Pterioida. A large bivalve, with a subequivalved shell, up to 60–70 cm high. It is one of the Three main bivalves recovered on the Lithiotis Facies, with its accumulations generally overlying megalodontid coquinas.[20] |
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A clam, Incertade sedis inside Pterioida. This genus was fund to be a bivalve with a byssate juvenile stage that developed different modes of life on the adulthood depending on the individual density and bottom firmness.[22] |
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An oyster, member of Malleidae inside Ostreida. It is the major Bivalve identified on the formation, and the genus that gives the name to the Lithiotis fauna.[20] Large, large and aberrant bivalves present on mostly of the Trento Platform.[22] Its accumulation have had different denominations on literature, such as banks, bioherms, biostromes, bivalve reefs or bivalve mounds.[20] |
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Isolated Shells |
A Oyster, member of Malleidae inside Ostreida. On the Rotzo formation this genus become abundant along rootlets, indicative of a very shallow and restricted lagoon or marsh environment.[20] |
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Isolated Shells |
Ammonoidea[]
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Juraphyllites libertus |
Contrada Ronchi (Recoaro Terme, Vicenza) |
Shells of different sizes.[23] |
Type member of the family Juraphyllitidae. It is the most abundant Ammonite found on the Rotzo Formation |
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Shells of different sizes.[23] |
An Ammonite of the Family Hildoceratidae |
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Shells of different sizes. |
An Ammonite of the family Hildoceratidae. |
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Ugdulenaia cf. ugdulenai |
|
Shells of different sizes. |
An Ammonite of the family Hildoceratidae. |
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Partschiceras anonimum |
|
Shells of different sizes. |
An Ammonite of the family Phylloceratidae. |
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Charmasseiceras[23] |
Charmasseiceras sp. |
Shells of different sizes.[23] |
An Ammonite of the family Schlotheimiidae. A very rare genus on the layers of the formation, being found only a few specimens. |
Gasteropoda[]
Genus | Species | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|
Neritopsis fabianii |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail), type genus of the Family Neritopsidae inside Cycloneritimorpha. |
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Guidonia pseudorotula |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail) of the Family Trochonematidae inside . |
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Pseudorhytidopilus detonii |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Limpet) of the Family Acmaeidae inside Patellogastropoda. |
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Proacirsa () crenata |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail) of the Family Gordenellidae inside Allogastropoda. |
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Discohelix excavata |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail), type genus of the Family Discohelicidae inside Vetigastropoda. |
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Eucyclidae Indeterminate |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail) of the Family Eucyclidae inside Seguenzioidea. |
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Eucyclus () kericserensis |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail), type genus of the Family Eucyclidae inside Seguenzioidea. |
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Austriacopsis austriaca |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail) of the Family Fissurellidae inside Fissurelloidea. |
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Emarginula (Emarginula) vadanaei |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail) of the Family Fissurellidae inside Fissurelloidea. |
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Anticonulus acutus |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Top Snail) of the Family Trochidae inside Trochoidea. |
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Plectotrochus sp. |
Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Top Snail) of the Family Trochidae inside Trochoidea. |
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Certosa di Vedana |
Shells of different sizes.[27] |
A Marine Gasteropod (Snail), type genus of the Family Ataphridae inside Trochoidea. |
Echinoidea[]
Genus | Species | Stratigraphic position | Material | Notes | Images |
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Two specimens (MCV.20/02 and MCV.20/03) |
A Phymosomatoidan. This Echinoids are recovered from a marginal marine layer, with abundant bivalves, gastropods, small corals, often found in concentrations due to tempestites.[28] |
Thylacocephala[]
Genus | Species | Stratigraphic position | Material | Notes | Images |
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Medium-sized bivalved carapace |
A Concavicaridan Thylacocephalan. This specimen is a rathin rare case where there was the discover of a Thylacocephalan specimen in a rock deposed in a well oxigenated environment, while other finds come mostly from were from anaerobic environments.[29] Rugocaris lived in an epibathyal environment.[29] |
Crustacea[]
Genus | Species | Stratigraphic position | Material | Notes | Images |
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Pustulina sp. |
Valbona Area |
Chelae |
An Erymidae Decapodan. There is a frequent presence of Thalassinoides burrows associated with Pustulina body fossils.[14] |
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Phlyctisoma sinemuriana |
Valbona Area.[14] |
Slightly deformed Exuvia |
An Erymid Decapodan Crustacean common on In mediterranean rocks. With a rostrum about 1.3 cm long and the cephalic part of carapace about 2.5 cm the specimen probably reached a total length between 9 and 10n cm, being one of the largest specimens belonging to this genus. Frequent association with Thalassinoides burrows. A complete seafloor section was fossilized.[14][13] |
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Phraterfabanella tridentinensis |
Tonezza del Cimone.[30] |
Valves |
An Ostracodan of the family Cytherideidae inside . The assemblage is dominated (>95%) by this taxon.[30] It is a rather Medium-sized Ostracodan and markedly sexually dimorphic (males more elongate and more subrectangular versus shorter, more inflated and more subtriangular females).[30] it is likely that the palaeoenvironment was somewhat "stressed" and probably influenced by Salinity, where this genus would adapt better that Other Ostracodans (is related to the modern euryhaline species, torosa).[30] |
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Klieana sp. |
Tonezza del Cimone.[30] |
Valves |
An Ostracodan of the family Cytherideidae inside . The earliest record of the genus, the next youngest records of the genus are from Middle Jurassic sequences of France and Great Britain.[30] |
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Limnocythere sp. |
Tonezza del Cimone.[30] |
Valves |
An Ostracodan of the family inside . High probability to be a new species of Limnocythere since the authors know of no other with similar posterolateral sulcation.[30] |
Vertebrata[]
Chondrichthyes[]
Episodic surficial bioturbation is common on the Rotzo Formation, due to invertebrates or fishes which alter intensely but rapidly the substrate for many cm in depth.[12] It this case the Bioturbation is assigned to mollusc predatory Chondrichthyes, such as Hybodontidae and Heterodontidae.[12] It also resembles present day flat angel sharks or Squatinidae and Guitarfish such as Rhinobatos.[12]
Genus | Species | Stratigraphic position | Material | Notes | Images |
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Orthacodus sp |
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Teeth |
A Shark, type genus of the family Orthacodontidae inside Synechodontiformes. The teeth recovered resemble Orthacodus longidens, and are related to an epibathyal environment, near to a major carbonate platform shelf. |
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Hybodus sp. |
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A Shark, type genus of the family Hybodontidae inside Hybodontiformes. A very prolific genus, found mostly on open marine units. |
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Chimaeriformes [32] |
Chimaeriformes Indeterminate |
Campiluzzi Tunnel, west of Monte Buso. |
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Uncertain remains |
Actinopterygii[]
Unidentified fish scales are known from the formation.[33]
Genus | Species | Stratigraphic position | Material | Notes | Images |
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Semionotiformes Indeterminate |
Campiluzzi Tunnel, west of Monte Buso. |
The assigned teeth where found on a layer referred to a Carbonate Platform nearshore section, probably a Lagoonar Environment, where fhis and marine crocodrylomorphs live. |
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Lepidotes sp. |
Campiluzzi Tunnel, west of Monte Buso. |
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A member of the family Semionotidae inside Neopterygii. |
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Pycnodontiformes indeterminate |
Campiluzzi Tunnel, west of Monte Buso. |
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Teleostei Fishes of small size, related to lagoonar environments |
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Pholidophoriformes Indeterminate |
Campiluzzi Tunnel, west of Monte Buso. |
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Teleostei fishes, with genera know to form large Fish schools. |
Crocodyliformes[]
Genus | Species | Location | Material | Notes | Images |
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Teleosauridae? Indeterminate |
Monte Pasubio |
A Thalattosuchian Mesoeucrocodylian. It was cited the presence of fragmentary and poorly preserved remains of “Teleosauridae?”. There are at least two morphotypes, implying two genera or two species. The fossils were found on lagoonal deposits.[33] |
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aff. Eopneumatosuchus sp. |
Monte Pasubio |
Partial skull and teeth.[34] |
A basal crocodyliform. Was originally thought to be Thalattosuchian remains. Has been compared recently with Eupneumatosuchus, is a member of the genus or a closely related species. |
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Metasuchia Indeterminate |
Monte Pasubio |
Upper jaw with rounded teeth.[34] |
A basal crocodyliform. It has rouded teeth that suggest a Duriphagous Diet. |
Dinosaurs[]
On the Inter-supratidal levels show that on the Rotzo Formation the Tracksites were rarely hit by Storm Waves.[36] Bella Lastra Tracksite recovers this environment, where the shales present (Where Fish & Crocodrylomorph Remains where found) are filled with plant roots, pollen grains, spores, freshwater ostracodes and the bivalve .[36] This was deposited mostly on a Lagoonar environment with abundant shed vegetation.[36] The main local Track record recovers specially Theropoda and Sauropoda, where the Sauropods are the most abundant tracks present (70%), moving the -like Sauropodomorphs of lower levels, with the climate changing from arid to humid.[36] The Coste dell’Anglone ichnosite is considered as derived from semi-arid tidal flat deposits, due to the abundance of Cheirolepidiaceae Pollen.[37] As the Pliensbachian Trento Platform is considered to be formed by a channelized barrier formed by sand, with reiterate tide emersions. The dinosaurs living here probably trampled on the subtidal flats looking for fishes trapped on tidal-derived ponds.[37]
Color key
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Notes Uncertain or tentative taxa are in small text; |
Dinosaurs of the Rotzo Formation | ||||||
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Genus | Species | Location | Member | Material | Notes | Images |
Moyenisauropus sp. |
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Tovel Member |
Footprints |
Thyreophoran tracks, type member of the ichnofamily , incertade sedis inside Neornithischia. Is considered by some authors synonymous with the ichnogenus Anomoepus. The tracks adscribed share some morphological affinity with those referred to the Ankylosauridae, such as the ichnogenera and Tetrapodosaurus, and probably belonged to medium-sized Scelidosaurs or other kind of Thyreophorans. Include Specimens of up to 30 cm, suggesting +4 m long scelidosauroids.[38] |
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Tovel Member |
Footprints |
Theropod tracks, type member of the ichnofamily , incertade sedis inside Neotheropoda. Probably related to Coelophysidae, such as Procompsognathus and Panguraptor or Coelophysoidea, such as Lophostropheus. All tracks were probably produced by individuals with the same functional anatomy of the hind foot.[36] |
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Tovel Member |
Footprints |
Theropod tracks, member of the ichnofamily , incertade sedis inside Neotheropoda. Includes Kayentapus sp. assigned to Sinosaurus-alike Theropods, but on the Rotzo Formation include also Abelisauroid-like tracks, similar to the foot of the genus Velocisaurus.[33] The tracks measure 30 cm long and have a distinctive robust digit III.[36] The Coste dell´Anglone tracksite had a pes with the metatarsal III elongated, as found on Dilophosaurus.[37] |
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Tovel Member |
Footprints |
Sauropodomorph tracks, member of the ichnofamily , incertade sedis inside Sauropodomorpha. A single trackway that strongly differs from the others found on the same tracksite. It wears morphological and morphometrical appearance that suggests relationships with a prosauropod trackmaker.[38] |
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Parabrontopodus sp. A Parabrontopodus sp. B |
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Tovel Member |
Footprints |
Sauropod tracks, type member of the ichnofamily , incertade sedis inside Sauropodomorpha. Tracks from large basal members of Sauropoda. The larger tracks comprise elliptic pes (L=70 cm; W=50 cm) and subcirluar manus prints (L=33 cm; W=30 cm), what are among the largest known dinosaur tracks of the lower jurassic.[36] While nearly destroyed, the Tracks resemble the foot of the genus Barapasaurus. There is a type B of Parabrontopodus slightly smaller that resemble the genus Vulcanodon. |
Flora[]
The Rotzo Formation was deposited on a Lagoon on the emerged Trento Platform, leading to a well preserved fossil flora record, collected and studied since the XIX century.[42] The great level of floral fossilization has even allow to discovery fossil amber on the Bellori section. This amber has allow to determine that the environment was a shallow tropical lagoon, only a few metres deep, closed seawards by oolitic shoals and bars.[42] This levels are dominated by a high abundance of sp. (Cheirolepidiaceae), associated with dry and wet climates in coastal areas. The abundance of this group of conifers is also proven by the high presence of cuticles of Pagiophyllum cf. rotzoanum.[43] Beyond this genera, spores are highly diversified, including from Sphenophyta, Selaginellales to Ferns, with abundance (more than 50%) of trilete spores (), what suggest a good freshwater availability corresponding to a wet climate, proven also by the presence of aquatic miospores of algae such as Botryococcus and .[42] The climate was arid on some seasons with monsoon months. The abundance of marine fauna on this sediments, including fragments of corals, bryozoans, bivalves, echinoids, and foraminifera, suggest transport from brackish lagoons and marshes, probably occurred during storm events.[42] Overall data points to a marshy and/or submerged paleoenvironment, comparable to the present-day Taxodium swamp or cypress swamp and a Bahamian-type marine environment in a rather wet monsoonal climate as in the modern southeastern Asia.[42][43]
Amber[]
Type | Location | Stratigraphic position | Material | Notes |
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Bellori village |
Amber Fragments |
The Lessini Mountains Amber represents the first report of Jurassic amber from Italy and one of the very few in the world. Due to being composed by drops of less than 1 mm with preserved exceptionally intact morphologies the Bellori amber was probably neither environmentally stressed nor affected by diseases.[44] While no animal remains were found inside the Bellori Amber so far there are various traces of very small (< 1 mm) vegetal fragments, here identified as tissue remains of wood and “mummified wood”.[44] It has also a large amount of Circumpolles (Cheirolepidiaceae), and in some fragments there are traces of the freshwater algae .[44] Although several cuticles found in Bellori could be attributed to Pagiophyllum (Araucariaceae).[44] Those lived on a coastal and wet palaeoenvironment similar to the present-day Taxodium swamps with monsoonal seasons as in the modern southern Asia.[44] |
Palynology[]
Genus | Species | Location | Stratigraphic position | Material | Notes |
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Accincitisporites sp. |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Unknown affinities | ||
Alisporites sp. |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with (), Corystospermales, Ginkgoopsida (Pelataspermales), Coniferopsida (Podocarpaceae, Ulmanniaceae, Voltziales).[44] | ||
Aratrisporites sp |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycophytes, in situ in , and zeiller.[44] | ||
Auritulinasporites scanicus |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Pteridophyta.[44] | ||
Baculatisporites sp |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Pteridopsida.[44] | ||
Calamospora sp |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Sphenopsida.[44] | ||
Calamospora sp |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopsida.[44] | ||
cf. Cabochonicus carbunculus |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[46] |
Affinities with Selaginellaceae | ||
Chasmatosporites sp |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopsida.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Pollen.[44] |
Affinities with Cheirolepidiaceae.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Pteridophyta.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Cycadophyta.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycophyta.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycophyta.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Selaginellaceae.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Pollen.[44] |
Affinities with Selaginellaceae.[44] | ||
Horstisporites harrisii |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[46] |
Affinities with Selaginella-like | ||
cf. Hughesisporites orlowskae |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[46] |
Affinities with Lycophyta | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Pteridopsida.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopsida.[44] | ||
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Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopsida.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopsida.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopsida.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Pteridopsida.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Pollen.[44] |
Affinities with Gymnospermophyta.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Pollen.[44] |
|||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopodiaceae.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Lycopodiaceae.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
|||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Chlorophyta.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Pollen.[44] |
Affinities with Chlorophyta.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Filicales.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Pteridopsida.[44] | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Pteridopsida.[44] | ||
cf. Trileites murrayi |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[46] |
Affinities with Selaginellaceae | ||
cf. Verrutriletes compostipunctatus |
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[46] |
Affinities with Selaginellaceae | ||
|
Bellori, Ponte Basaginocchi, Vajo dell’Anguilla |
Spores.[44] |
Affinities with Gymnospermophyta.[44] |
Plant remains[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
---|---|---|---|---|---|---|
|
|
Stems |
Affinities with Equisetaceae inside Polypodiopsida. Related to humid environments, the stems of local Equisetopsids show a rather large grown cycle, like the Bamboo on the modern Southern Asia, implicating tall Plants influenced by a Tropical Climate. |
|||
|
|
Leaf Whorl |
Affinities with Phyllothecaceae inside Equisetales |
| ||
|
|
Fronds |
Affinities with Dipteridaceae inside Gleicheniales. A rather lower Fern, with great resemblance with the modern genus Dipteris |
|||
|
|
Fronds |
Affinities with Matoniaceae inside Gleicheniales. |
|||
|
|
Fronds |
Affinities with Matoniaceae inside Gleicheniales. |
| ||
|
|
Fronds |
Affinities with Matoniaceae inside Gleicheniales. |
|||
|
|
Fronds |
Affinities with Dipteridaceae inside Gleicheniales. |
|||
|
|
Fronds |
Affinities with Polypodiales inside Polypodiopsida |
|||
|
|
Fronds |
Affinities with Lyginopteridopsida inside Lyginopteridales |
|||
|
|
Fronds |
Affinities with Cyclopteridaceae inside Pteridospermatophyta. |
| ||
|
|
Fronds |
Affinities with Cyclopteridaceae inside Pteridospermatophyta. |
|||
|
|
Fronds |
Affinities with inside . On the Roverè di Velo collection, C. brauniana is the most common Frond found. The Fronds belong to medium to large Arboreal Ferns. |
| ||
|
|
Fronds |
Affinities with inside . This frons genus has been Synonymized with Pachypteris , but it clearli differs due to the presence of odontopteridian pinnules, while Pachypteris has pinnules of the sphenopteridian type. Related to Arboreal Ferns. |
| ||
|
|
Leaflets |
Affinities with Caytoniales inside . |
|||
|
|
Leaflets |
Affinities with Caytoniaceae inside Caytoniales. There is a superficial doubt with the assigantion to S. goeppertiana, and du to that Roverè di Velo specimen may be confirmed by comparing them with original Zigno's Material. |
| ||
|
Grey limestones of Veneto |
Leaflets |
Affinities with Bennettitales inside . Related with Cycad-like trees. |
|||
|
Grey limestones of Veneto. |
Reproductive structure |
Affinities with Bennettitales inside . Weltrichia is considered by some authors some kind of Bennetitalean Flower, putting that group on relationships with the Angiosperms. |
|||
|
Grey limestones of Veneto. |
Leaflets |
Affinities with Bennettitales inside . This genus has been related with the more arboreal family Williamsoniaceae, although is more probably from a low arboreal to arbustive Bennetite. |
|||
|
|
Pinnate leaf fragments |
Affinities with Bennettitales inside . Overall, the genus Otozamites is among the most abundant flora genus recovered on some of the levels of the Rotzo Formation, and also one of the most diversified. It belongs to arbustive Bennetites. |
| ||
|
|
Leaves |
Affinities with Bennettitales. Was previously ascribed by Guiscardi (Director of the Geology Department of the Napoles University between 1861 al 1885) to Pachypteris visianica and Cycadopteris brauniana. |
| ||
|
|
Leaves |
Affinities with Williamsoniaceae. |
| ||
|
|
Leaves |
Affinities with the genus , as probably a member of Ginkgoales inside Ginkgoopsida. |
|||
|
Roverè di Velo |
Incomplete leaves |
Affinities with Ginkgoaceae inside Ginkgoales. Was assigned the Podozamites genus and named them Podozamites zeillerianus. |
|||
|
Grey limestones of Veneto. |
Pollen Organ |
Incertade sedis inside Coniferales. Some Specimens are difficult to identify. |
|||
|
Grey limestones of Veneto. |
Branched shoots |
Affinities with Cupressaceae inside Coniferales. Some Specimens are difficult to identify and where mistaken as Bennetite Fronds. |
| ||
|
Grey limestones of Veneto. |
Branched shoots |
Affinities with inside Coniferales, a genus that has been related to the fossil wood , being probably Fronds of the Podocarpaceae family. |
|||
|
Grey limestones of Veneto. |
Branched shoots |
Affinities with Podocarpaceae inside Coniferales. |
| ||
|
|
Branched shoots |
Affinities with Cheirolepidiaceae inside Coniferales. |
|||
|
|
Branched shoots |
Affinities with Araucariaceae or Cheirolepidiaceae inside Coniferales. Brachyphyllum tropidimorphyrn shows close resemblance between African and Venetian conifers and its distribution suggests a lowland araucarian forest.[52] |
| ||
|
Roverè di Velo |
|
Affinities with Araucariaceae or Cheirolepidiaceae inside Coniferales. One of the specimens was assigned to Otozamites massalongianus, due to confusing the overlapping appearance and the Otozamites-like shape of the leaves of the apical portion of the main shoot. |
| ||
|
Roverè di Velo |
Nearly complete pinna |
Due to the bad preservation of specimen no systematic attribution is possible. |
See also[]
- List of dinosaur-bearing rock formations
- List of stratigraphic units with sauropodomorph tracks
- Prosauropod tracks
- List of stratigraphic units with sauropodomorph tracks
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- Geologic formations of Italy
- Jurassic System of Europe
- Jurassic Italy
- Pliensbachian Stage
- Limestone formations
- Lagoonal deposits
- Ichnofossiliferous formations
- Paleontology in Italy