2020 in paleoichthyology

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List of years in paleoichthyology
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2020
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2018
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This list of fossil fishes described in 2020 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes of every kind that were described during the year 2020, as well as other significant discoveries and events related to paleoichthyology that occurred in 2020.

New taxa[]

Jawless vertebrates[]

Name Novelty Status Authors Age Type locality Country Notes Images
[1] Nom. nov Valid Elliott, Lassiter & Geyer Devonian (Givetian) Yahatinda Formation  Canada
( Alberta) 		A member of the family Pteraspididae; a replacement name for Helaspis Elliott et al. (2000).
[1] Gen. et comb. nov Valid Elliott, Lassiter & Geyer Devonian (Eifelian) Spring Mountain beds  United States
( Idaho) 		A member of the family Pteraspididae; a new genus for "" idahoensis Denison (1968).
[2] Sp. nov Valid Cui et al. Devonian (Lochkovian) Xishancun

Xitun

 China A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
[2] Sp. nov Valid Cui et al. Devonian (Lochkovian) Xishancun

Xitun

 China A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
[2] Sp. nov Valid Cui et al. Devonian (Lochkovian) Xitun  China A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.
[3] Gen. et sp. nov Valid Dec Devonian (Givetian)  Poland A member of the family Psammosteidae. The type species is P. mediocris.
[4] Gen. et comb. et sp. nov Valid Shan et al. Silurian (Telychian) Xikeng Formation  China A member of Galeaspida belonging to the group and the family . The type species is "" xikengensis Pan & Wang (1980); genus also includes new species R. shipanensis.
[1] Gen. et sp. nov Valid Elliott, Lassiter & Geyer Devonian (Eifelian) Spring Mountain beds  United States
( Idaho) 		A member of the family Pteraspididae. Genus includes new species S. wilsoni.

Placoderms[]

Name Novelty Status Authors Age Type locality Country Notes Images
Actinolepis zaikai[5] Sp. nov Valid Plax & Newman Devonian (Emsian) Lepel Beds  Belarus
[6] Gen. et sp. nov Valid Liu, Zong & Gong Late Devonian Zhulumute Formation  China A member of the family Selenosteidae. Genus includes new species B. liui.
[7] Gen. et comb. nov Valid Vaškaninová Early Devonian Prague Basin  Czech Republic A homostiid arthrodire. Genus includes "Asterolepis" bohemicus Barrande (1872).
Minjinia[8] Gen. et sp. nov Valid Brazeau et al. Devonian (Pragian)  Mongolia A placoderm closely related to the gnathostome crown group. Genus includes new species M. turgenensis.
[5] Gen. et sp. nov Valid Plax & Newman Devonian (Emsian) Lepel Beds  Belarus A phlyctaeniid arthrodire. Genus includes new species S. svidunovitchi.

Acanthodians[]

Name Novelty Status Authors Age Type locality Country Notes Images
[9] Gen. et sp. nov In press Newman, Den Blaauwen & Burrow Devonian (Givetian) Mey Flagstone Formation  United Kingdom A cheiracanthid acanthodian. Genus includes new species F. davidsoni.

Cartilaginous fishes[]

Name Novelty Status Authors Age Type locality Country Notes Images
Amamriabatis[10] Gen. et sp. nov Valid Adnet et al. Eocene  Tunisia
 Egypt? 		A ray belonging to the family Mobulidae. The type species is A. heni.

Asteracanthus dunaii[11]

Sp. nov

Valid

Szabó & Főzy

Jurassic

 Hungary

Atlantitrygon[12] Gen. et comb. et sp. nov Valid Sambou et al. Paleocene (Thanetian) and Eocene (Lutetian)

 Niger
 Senegal
 Togo

A putative marine potamotrygonid. The type species is "Dasyatis" sudrei Cappetta (1972); genus also includes new species A. senegalensis.
[13] Gen. et sp. nov Valid Popov, Johns & Suntok Late Oligocene Sooke  Canada
( British Columbia) 		A member of the family Chimaeridae. Genus includes new species C. suntoki.
[10] Sp. nov Valid Adnet et al. Eocene (Bartonian)  Tunisia
 Jamaica? 		A species of Carcharhinus.
Coupatezia cristata[10] Sp. nov Valid Adnet et al. Eocene  Egypt
 Tunisia 		A ray belonging to the superfamily .
[14] Gen. et sp. nov Valid Maisey, Ehret & Denton Late Cretaceous (Campanian) Mooreville Chalk
Navesink Formation? 		 United States
( Alabama
 New Jersey?) 		A member of the family Squatinidae. The type species is C. americana.
[15] Gen. et comb. nov Valid Marramà et al. Eocene Monte Bolca  Italy A member of the family Platyrhinidae. Genus includes "Platyrhina" bolcensis Heckel (1851).
[16] Gen. et comb. nov Valid Marramà et al. Eocene Monte Bolca  Italy A guitarfish; a new genus for "Rhinobatos" primaevus.
[17] Gen. et sp. nov Valid Frey et al. Devonian (Famennian)  Morocco A member of Symmoriiformes. The type species is F. oukherbouchi.
[10] Sp. nov Valid Adnet et al. Eocene (Bartonian)  Tunisia A whiptail stingray, a species of Himantura.
[18] Gen. et sp. nov Valid Ivanov in Ivanov et al. Permian (Guadalupian) Bell Canyon Formation  United States
( Texas) 		A member of Hybodontiformes belonging to the superfamily Hybodontoidea. Genus includes new species L. triangulus. Announced in 2018; the final version of the article naming it was published in 2020.
[10] Sp. nov Valid Adnet et al. Eocene  Tunisia
 Egypt?
 Jordan? 		A relative of the barbeled houndshark.
[10] Gen. et sp. nov Valid Adnet et al. Eocene  Egypt
 Tunisia 		A whiptail stingray belonging to the subfamily . The type species is M. asperodentulus.
[19] Sp. nov Valid Cicimurri, Ebersole & Martin Paleocene (Danian) Clayton  United States
( Alabama) 		A sand shark
Mennerotodus parmleyi[19] Sp. nov Valid Cicimurri, Ebersole & Martin Eocene (Bartonian) Clinchfield  United States
( Georgia (U.S. state)) 		A sand shark
[12] Gen. et sp. nov Valid Sambou et al. Eocene (Ypresian to Lutetian)  Senegal A of uncertain phylogenetic placement. Genus includes new species M. matami.
Nanocetorhinus zeitlingeri[20] Sp. nov Valid Feichtinger, Pollerspöck & Harzhauser Oligocene (Chattian)  Austria A member of Neoselachii of uncertain phylogenetic placement.

[12]

Sp. nov Valid Sambou et al. Paleocene (Thanetian) and Eocene (Ypresian)

 Mali
 Morocco
 Senegal
 Tunisia

A member of Neoselachii of uncertain phylogenetic placement
[10] Sp. nov Valid Adnet et al. Eocene (Bartonian)  Tunisia A butterfly ray.
[10] Gen. et comb. nov Valid Adnet et al. Eocene  Egypt
 Tunisia 		A butterfly ray. The type species is "Coupatezia" attiai Cook in Murray et al. (2010).
[15] Gen. et comb. nov Valid Marramà et al. Eocene]] Monte Bolca  Italy A relative of panray. Genus includes "Torpedo" egertoni De Zigno (1876).
[21] Sp. nov Valid Reinecke et al. Miocene (Aquitanian) Ebelsberg  Austria A species of Pristiophorus.
[21] Sp. nov Valid Reinecke et al. Oligocene (Chattian)  Germany A species of Pristiophorus.
[21] Sp. nov Valid Reinecke et al. Miocene (Tortonian)  Germany A species of Pristiophorus.
[21] Sp. nov Valid Reinecke et al. Miocene (Burdigalian)  Germany A species of Pristiophorus.
[16] Gen. et comb. nov Valid Marramà et al. Eocene Monte Bolca  Italy A guitarfish; a new genus for "Rhinobatos" dezignii.

[22]

Gen. et sp. nov

Valid

Lebedev in Lebedev, Ivanov & Linkevich

Devonian (Famennian)

 Russia
( Lipetsk Oblast)

A member of Ctenacanthiformes of uncertain phylogenetic placement. The type species is S. makhlaevi.

[10] Sp. nov Valid Adnet et al. Eocene  Tunisia
 Egypt? 		A species of Sphyrna.
[10] Sp. nov Valid Adnet et al. Eocene  Tunisia
 Egypt? 		A relative of the zebra shark.
[23] Gen. et sp. nov Andreev et al. Early Silurian  China A member of Mongolepidida. The type species is T. asiaticus.
[23] Gen. et sp. nov Andreev et al. Early Silurian

 China A member of Mongolepidida. The type species is T. sinensis.
[23] Gen. et sp. nov Andreev et al. Early Silurian

 China A member of Mongolepidida. The type species is X. liui.
[23] Gen. et sp. nov Andreev et al. Early Silurian  China A cartilaginous fish of uncertain phylogenetic placement. The type species is Y. bachunensis.

Ray-finned fishes[]

Name Novelty Status Authors Age Type locality Country Notes Images
Agassizilia[24] Gen. et sp. nov Valid Cooper & Martill Cenomanian Kem Kem  Morocco Possibly a member of the family Pycnodontidae. The type species is A. erfoudina.
[25] Gen. et sp. nov Valid Ebert Late Jurassic Solnhofen  Germany A member of Halecomorphi belonging to the group . Genus includes new species A. boomerang.
[26] Gen. et sp. nov Valid Yabumoto, Hirose & Brito Late Cretaceous (Santonian)  Japan A member of Ichthyodectiformes. Genus includes new species A. goshouraensis. Announced in 2018; the final version of the article naming it was published in 2020.
Ampheristus americanus[27] Sp. nov Valid Schwarzhans & Stringer Late Cretaceous (Maastrichtian) and Paleocene (Danian) Clayton
Kemp Clay 		 United States
( Arkansas
 Texas) 		A cusk-eel.
[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) and Paleocene (Danian)[27] Clayton[27]
Ripley 		 United States
( Arkansas[27]
 Mississippi) 		A member or a relative of the family Anguillidae
Anomoeodus wolfi[29] Sp. nov Valid Capasso Late Cretaceous (Cenomanian) Del Rio  United States
( Texas) 		A member of the family Pycnodontidae.
[30] Gen. et sp. nov. Valid Bannikov & Zorzin Eocene (Ypresian) Monte Bolca  Italy A member of Percomorpha of uncertain phylogenetic placement. The type species is A. lorenzonii.
Apateodus crenellatus[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Owl Creek

Ripley

 United States
( Mississippi)
[31] Sp. nov In press Aguilera & Rodriguez in Aguilera et al. Late Oligocene to early Miocene Castillo  Venezuela A species of Ariopsis.
Armigatus carrenoae[32] Sp. nov Valid Alvarado-Ortega, Than-Marchese & Melgarejo-Damián Early Cretaceous (Albian) Tlayúa  Mexico A member of Clupeomorpha belonging to the group and to the family .
[31] Sp. nov In press Aguilera & Rodriguez in Aguilera et al. Late Oligocene to early Miocene Castillo  Venezuela A species of Bagre.
[33] Gen. et 3 sp. nov Valid Altner & Reichenbacher Late Miocene  Kenya A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is B. senutae; genus also includes B. sonyii and B. tallamae. Announced in 2020; the final version of the article naming it was published in 2021.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy A tadpole goby.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy A tadpole goby.
[35] Gen. et sp. nov Valid Weems Eocene Nanjemoy  United States
( Virginia) 		A member of the family Latidae. The type species is B. debernardi.
[36] Sp. nov In press Agnolin & Bogan Late Miocene  Argentina A species of Brachyplatystoma
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Tortonian and Messinian)  Greece A species of Buenia.
[37] Sp. nov Valid Bakaev & Kogan Permian (Wordian–Capitanian)  Russia An early ray-finned fish. Announced in 2019; the final version of the article naming it was published in 2020.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy
 Romania 		A species of Caspiosoma.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy A species of Caspiosoma.
[38] Gen. et sp. nov Valid Bannikov Early Oligocene Abkhazia
 Russia 		A member of the family Scombridae. Genus includes new species C. histiopterygius.
[39] Gen. et sp. nov Valid Cantalice Severiano, Alvarado Ortega & Bellwood Paleocene  Mexico A member of the family Pomacentridae. Genus includes new species C. gonzalezorum. Announced in 2019; the final version of the article naming it was published in 2020.
[28] Gen. et comb. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Ripley  United States
( Mississippi) 		A member of Paracanthopterygii of uncertain phylogenetic placement. The type species is "genus Perciformorum" cepoloides Nolf & Dockery (1990).

[40]

Sp. nov

Valid

Schwarzhans, Brzobohatý & Radwańska

Miocene

Possibly a species of Chromogobius.

[41] Nom. nov Valid Yabumoto & Nazarkin Miocene  Japan A species of Clupea; a replacement name for Clupea macrocephala Yabumoto & Nazarkin (2018).
[27] Gen. et sp. nov Valid Schwarzhans & Stringer Late Cretaceous (late Campanian and Maastrichtian) and Paleocene (Danian) Clayton
Coffee Sand
Kemp Clay 		 United States
( Alabama
 Arkansas
 Mississippi
 Texas) 		A member of the family Congridae. The type species is C. transterminus.
[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Ripley  United States
( Mississippi) 		A beardfish
[42] Sp. nov Valid Nam & Nazarkin Miocene  South Korea A species of Cyclothone. Announced in 2020; the final version of the article naming it was published in 2021.
[27] Gen. et sp. nov Valid Schwarzhans & Stringer Late Cretaceous (Maastrichtian) Fox Hills
Kemp Clay 		 United States
( North Dakota
 Texas) 		Possibly a member of the family Gadidae. The type species is D. hogansoni.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Tortonian)  Italy A species of Eleotris.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Tortonian)  Italy A species of Eleotris.
[43] Sp. nov Valid Polck et al. Early Cretaceous (Aptian) Barra Velha Formation  Brazil A member of Clupeomorpha belonging to the group and to the family . Announced in 2019; the final version of the article naming it was published in 2020.
[44] Sp. nov In press De Figueiredo & Gallo Early Cretaceous Recôncavo Basin  Brazil A member of Clupeomorpha belonging to the group Ellimmichthyiformes.
[25] Gen. et sp. nov Valid Ebert Late Jurassic  Germany A member of Halecomorphi belonging to the group . Genus includes new species E. woelflei.
[27] Sp. nov Valid Schwarzhans & Stringer Paleocene (Danian) Clayton  United States
( Arkansas) 		A relative of the Japanese gissu.
[45] Gen. et sp. nov Valid Murray et al. Paleocene Fort Union  United States
( Montana) 		A sturgeon. Genus includes new species E. milviaegis.
[34] Gen. et sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy Possibly a member of the family Psychrolutidae. The type species is E. socialis.
[46] Gen. et sp. nov In press Alves, Alvarado Ortega & Brito Early Cretaceous (Albian) Tlayúa  Mexico A member of Elopiformes. Genus includes new species E. martinezi. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.
[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Kemp Clay[27]
Ripley 		 United States
( Mississippi
 Texas[27]) 		Probably a member of Beryciformes
[47] Gen. et sp. nov Valid Xu Middle Triassic (Anisian) Guanling  China A member of the family Colobodontidae. The type species is F. yunnanensis.
[29] Gen. et sp. nov Valid Capasso Late Cretaceous (Cenomanian) Del Rio  United States
( Texas) 		A member of the family Pycnodontidae. The type species is G. dentespassim.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Pliocene (Zanclean)  Greece A species of Gobius.

[40]

Sp. nov

Valid

Schwarzhans, Brzobohatý & Radwańska

Miocene

A species of Gobius.

[48] Gen. et comb. nov Valid Taverne, Capasso & Del Re Early Cretaceous (Albian)  Italy A member of the family Pycnodontidae. The type species is "Palaeobalistum" bassanii d'Erasmo (1914).
[49] Gen. et sp. nov Valid Murray Early Cenozoic (possibly Eocene)  Indonesia A member of Cyprinoidea. Genus includes new species H. sandersae.
[50] Sp. nov In press Alvarado-Ortega & Díaz-Cruz Late Cretaceous (Turonian) Agua Nueva  Mexico A member of Aulopiformes belonging to the family .
[51] Sp. nov In press Baños Rodríguez et al. Cretaceous (Albian–Cenomanian) El Doctor  Mexico A member of Ichthyodectiformes
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Tortonian and Messinian)  Italy A goby.

[40]

Sp. nov

Valid

Schwarzhans, Brzobohatý & Radwańska

Miocene

 Moldova

A goby. Originally described as a species of Hesperichthys, but subsequently transferred to the genus Sarmatigobius.[52]

[53] Gen. et sp. nov In press Kim et al. Late Triassic  South Korea A member of Redfieldiiformes. Genus includes new species H. boryeongensis. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

[40]

Gen. et comb. nov

Valid

Schwarzhans, Brzobohatý & Radwańska

Miocene

 Germany

A goby. Genus includes H. preaclarus (Procházka, 1893) and H. laevis (Weiler, 1942).

[54] Gen. et sp. nov Valid Ebert, Thies & Hauff Early Jurassic (Toarcian) Posidonia  Germany A member of Halecomorphi. Genus includes new species H. rebmanni.
Hoplopteryx langfordi[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Ripley  United States
( Mississippi)
Hypsocormus posterodorsalis[55] Sp. nov Valid Maxwell et al. Late Jurassic (Kimmeridgian) Nusplingen  Germany
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy A species of Knipowitschia.

[40]

Sp. nov

Valid

Schwarzhans, Brzobohatý & Radwańska

Miocene

 Poland

A species of Knipowitschia.

[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Owl Creek

Ripley

 United States
( Mississippi) 		A member of Osteoglossiformes of uncertain phylogenetic placement
[56] Gen. et sp. nov Valid Veysey, Brito & Martill Late Cretaceous (Turonian)  Morocco A member of Crossognathiformes. Genus includes new species K. asflaensis. Announced in 2019; the final version of the article naming it was published in 2020.
[57] Sp. nov Valid McClellan & Smith Late Miocene Salt Lake Formation  United States
( Utah) 		A species of Lavinia.
[58] Gen. et sp. nov Valid Taverne & Capasso Late Cretaceous (Cenomanian)  Lebanon A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is L. nammourensis.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Tortonian)  Italy A species of Lesueurigobius.
[59] Gen. et sp. nov. Valid Murray et al. Late Cretaceous (Maastrichtian) Scollard Formation  Canada
( Alberta) 		A member of Percopsiformes. Genus includes new species L. albertensis. Announced in 2019; the final version of the article naming it was published in 2020.
[60] Gen. et sp. nov Valid Xu Middle Triassic (Anisian)  China A member of the stem group of Neopterygii. Genus includes new species L. pusillus.
Luganoia fortuna[61] Sp. nov Valid Xu Middle Triassic (Ladinian)  China
[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Ripley  United States
( Mississippi) 		Probably member of the family Megalopidae
[62] Gen. et sp. nov Capobianco et al. Eocene (Lutetian)  Pakistan A stem-engraulid. The type species is M. chureloides.
[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Owl Creek

Ripley

 United States
( Mississippi) 		An eel of uncertain phylogenetic placement
[63] Gen. et sp. nov Valid Carnevale et al. Eocene (Ypresian) Monte Bolca  Italy A frogfish. Genus includes new species N. ceratoi.
[64] Gen. et sp. nov In press Cooper & Martill Turonian  Morocco A member of Pycnodontiformes. Genus includes new species N. asflaensis.
[24] Sp. nov Valid Cooper & Martill Cenomanian Kem Kem  Morocco A member of the family Pycnodontidae
[65] Gen. et sp. nov In press Cawley et al. Late Cretaceous (Turonian)  Germany A member of the family Pycnodontidae. The type species is N. dyckerhoffi.
[66] Gen. et sp. nov In press Hernández-Guerrero et al. Cretaceous (Albian-Cenomanian)  Mexico A bonefish belonging to the subfamily Pterothrissinae. Genus includes new species N. mexicanus.

[40]

Sp. nov

Valid

Schwarzhans, Brzobohatý & Radwańska

Miocene

A relative of the coralline goby.

[54] Gen. et sp. nov Valid Ebert, Thies & Hauff Early Jurassic (Toarcian) Posidonia  Germany A member of Halecomorphi. Genus includes new species O. bodmani.
Osmeroides mississippiensis[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Owl Creek

Ripley

 United States
( Mississippi)
[28] Gen. et sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Ripley  United States
( Mississippi) 		Probably a member of Beryciformes. The type species is O. labiatus.
[28] Sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Ripley  United States
( Mississippi) 		An otolith of a fish of uncertain phylogenetic placement.
Palaeogadus weltoni[27] Sp. nov Valid Schwarzhans & Stringer Late Cretaceous (Maastrichtian) Kemp Clay  United States
( Texas)
[67] Sp. nov Valid Schwarzhans & Mironenko Late Jurassic  Russia Possibly a stem-albuliform
[67] Sp. nov Valid Schwarzhans & Mironenko Late Jurassic  Russia Possibly a stem-albuliform.
[64] Sp. nov In press Cooper & Martill Turonian  Morocco A member of Pycnodontiformes.
[49] Gen. et comb. nov Valid Murray Early Cenozoic (possibly Eocene) Sangkarewang  Indonesia A member of Cyprinoidea. Genus includes "Puntius" bussyi.
[68] Gen. et sp. nov In press London & Shimada Late Cretaceous (Cenomanian) Tarrant  United States
( Texas) 		A member of the family Pachyrhizodontidae. Genus includes new species P. lloydhilli.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Pliocene (Zanclean)  Greece A species of Proterorhinus.
[27] Sp. nov Valid Schwarzhans & Stringer Late Cretaceous (Maastrichtian) and Paleocene (Danian) Clayton
Kemp Clay 		 United States
( Arkansas
 Texas) 		A species of Pythonichthys.
[69] Sp. nov Valid Godfrey & Carnevale Miocene (Tortonian) St. Marys  United States
( Maryland) 		A relative of the cobia.
[70] Sp. nov Valid Kevrekidis & Reichenbacher in Kevrekidis et al. Miocene  Kenya A cichlid belonging to the tribe Oreochromini.
Rebekkachromis vancouveringae[70] Sp. nov Valid Kevrekidis & Reichenbacher in Kevrekidis et al. Miocene Ngorora  Kenya A cichlid belonging to the tribe Oreochromini.
[27] Sp. nov Valid Schwarzhans & Stringer Late Cretaceous (Maastrichtian) Kemp Clay  United States
( Texas) 		A species of Rhynchoconger.
[71] Gen. et comb. nov Valid Baykina & Kovalchuk in Kovalchuk et al. Oligocene (Rupelian)  Russia A member of family Clupeidae. Genus includes R. rata (Daniltshenko, 1959).
[72] Gen. et sp. nov Valid Bannikov Early Oligocene Abkhazia
 Poland
 Russia
( Adygea) 		A member of Perciformes belonging to the new family . Genus includes new species S. paratethyensis.
[73] Gen. et sp. nov Valid López-Arbarello, Maxwell & Schweigert Late Jurassic (Kimmeridgian) Nusplingen Limestone  Germany A member of Halecomorphi. Genus includes new species S. rieteri.
[49] Gen. et sp. nov Valid Murray Early Cenozoic (possibly Eocene) Sangkarewang  Indonesia A member of Cyprinoidea. Genus includes new species S. sumatranus.
[74] Sp. nov Valid Zhang Early Eocene  China A species of Scleropages. Announced in 2019; the final version of the article naming it was published in 2020.
Scombroclupea javieri[75] Sp. nov In press Than Marchese et al. Late Cretaceous (Cenomanian)  Mexico A member of Clupeomorpha of uncertain phylogenetic placement
[76] Sp. nov Valid Yabumoto Miocene  Japan A species of Siniperca.
[55] Gen. et sp. nov Valid Maxwell et al. Late Jurassic (Kimmeridgian–Tithonian)  Germany A member of the family Pachycormidae. The type species is S. macrolepidotus.
[77] Gen. et sp. nov Valid Taverne & Capasso Late Cretaceous (Cenomanian)  Lebanon A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is S. libanicus.
[78] Gen. et sp. nov. Valid Bannikov & Zorzin Eocene]] (Ypresian) Monte Bolca  Italy A member of Percoidei of uncertain phylogenetic placement. The type species is S. mariannae.
[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy A species of Thorogobius.
[28] Gen. et sp. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Ripley  United States
( Mississippi) 		A member of Aulopiformes belonging to the family . The type species is T. acutus.
[28] Gen. et sp. et comb. nov Valid Schwarzhans & Stringer in Stringer et al. Late Cretaceous (Maastrichtian) Owl Creek

Ripley

 United States
( Mississippi) 		A member of Holocentriformes of uncertain phylogenetic placement. The type species is T. mythica; genus also includes "genus Ophidiidarum" cavatus Nolf & Stringer (1996).
[79] Gen. et sp. nov Valid Přikryl & Carnevale Early Oligocene  Czech Republic A member of the family Platytroctidae. Genus includes new species V. moraviensis.

[40]

Sp. nov

Valid

Schwarzhans, Brzobohatý & Radwańska

Miocene

A species of Vanderhorstia.

[80] Gen. et sp. nov Valid Díaz-Cruz, Alvarado-Ortega & Giles Late Cretaceous (Cenomanian)  Mexico A member of Aulopiformes belonging to the family . The type species is V. coitecus.

[81]

Gen. et sp. nov

Valid

Altner, Ruthensteiner & Reichenbacher

Late Miocene

 Kenya

A cichlid belonging to the subfamily Pseudocrenilabrinae. The type species is W. unicuspidatus.

[34] Sp. nov Valid Schwarzhans, Agiadi & Carnevale Miocene (Messinian)  Italy
 Romania 		A relative of the grass goby.

Lobe-finned fishes[]

Name Novelty Status Authors Age Type locality Country Notes Images
Ferganoceratodus annekempae[82] Sp. nov Valid Cavin, Deesri & Chanthasit Late Jurassic Phu Kradung  Thailand A lungfish.
Megalichthys mullisoni[83] Sp. nov Valid Downs & Daeschler Devonian (Famennian) Catskill  United States
( Pennsylvania)
Metaceratodus baibianorum[84] Sp. nov Valid Panzeri et al. Late Cretaceous La Colonia  Argentina A lungfish.
[85] Sp. nov Valid Kemp & Berrell Cretaceous  Australia A lungfish.
Ptychoceratodus oldhami[86] Sp. nov Valid Bhat & Ray Late Triassic (Carnian) Tiki  India A lungfish. Announced in 2018; the final version of the article naming it was published in 2020.
[87] Gen. et sp. nov Valid Johanson et al. Carboniferous (Tournaisian) Ballagan Formation  United Kingdom A member of Rhizodontida. The type species is R. clackae.

Research[]

  • A study on the morphology of the osteostracans, evaluating different methods used to determine the morphological variation within this group and its evolution, is published by Ferrón et al. (2020).[88]
  • A study on the morphological diversity of osteostracan headshields, aiming to determine the relationship between their morphological diversity and hydrodynamic performance and its implications for the knowledge of the ecological diversity of the osteostracans, is published by Ferrón et al. (2020).[89]
  • A study aiming to test the alternative hypotheses of placoderm jaw bone homologies, and evaluating their implications for the knowledge of evolution of jaw bones in early jawed vertebrates, is published by King & Rücklin (2020).[90]
  • A study on the fossil dentitions of acanthothoracids is published by Vaškaninová et al. (2020), who report that the teeth of acanthothoracids differed fundamentally from those of arthrodires, and argue that the characteristic traits of acanthothoracid dentition might be ancestral for all jawed vertebrates.[91]
  • Redescription of the anatomy of speciosa, based on data from new fossil material, and a study on the phylogenetic relationships of this species is published online by Lukševičs (2020).[92]
  • Description of the neurocranial anatomy of scheii is published by Castiello et al. (2020).[93]
  • A study aiming to determine whether Titanichthys was a suspension feeder, focusing on mechanical properties of its jaw, is published by Coatham et al. (2020).[94]
  • The earliest fossilized vertebrate embryos reported so far, preserved with an adult specimen of fletti from the Givetian Eday Flagstone Formation (Orcadian Basin; Scotland, United Kingdom), are described by Newman et al. (2020).[95]
  • Burrow, Newman & den Blaauwen (2020) describe external spiracular elements in Middle Devonian acanthodians from northern Scotland, differing from spiracles of all known extant and extinct fishes, and report the oldest record of elastic cartilage in the fossil record.[96]
  • A study examining the factors influencing the long-term variations of genus-level diversity of elasmobranchs and ray-finned fishes throughout their evolutionary history is published by Guinot & Cavin (2020).[97]
  • Carrillo-Briceño et al. (2020) describe a new elasmobranch assemblage from the Oligocene–Miocene boundary in the Dos Bocas Formation (Ecuador), and evaluate the implications of this assemblage for chronostratigraphic inferences and the knowledge of local paleoenvironment.[98]
  • Two large vertebrae of sharks belonging to the genus Ptychodus, providing new information on the life history and body size of members of the family Ptychodontidae, are described from the Santonian of Spain by Jambura & Kriwet (2020).[99]
  • A study on the evolution of body size in lamniform sharks, including the evolution of gigantism in the lineage of Otodus megalodon, is published by Shimada, Becker & Griffiths (2020).[100]
  • Taxonomic revision of the Oligocene and Miocene sand sharks is published by Hovestadt (2020).[101]
  • A study aiming to determine the linear body dimensions of Otodus megalodon at different life stages is published by Cooper et al. (2020).[102]
  • A study on the class structure of assemblages of specimens of Otodus megalodon in eight previously known formations and in a newly described Miocene locality from northeastern Spain is published by Herraiz et al. (2020), who interpret their findings as indicative of existence of five potential nurseries of these sharks ranging from the Langhian to the Zanclean.[103]
  • An isolated tooth of Cosmopolitodus hastalis from the Miocene of South Korea is described by Yun (2020).[104]
  • A study on the fossil record of the great white shark from the Pliocene of Peru and Chile is published by Villafaña et al. (2020), who interpret their findings as indicating that great white sharks used the Coquimbo locality in Chile as a nursery and Pisco (Peru) and Caldera (Chile) localities as feeding grounds during the Pliocene.[105]
  • A study on the anatomy and phylogenetic relationships of "Urolophus" crassicaudatus is published by Marramà et al. (2020), who transfer this species to the genus Arechia.[106]
  • Collareta et al. (2020) describe a fossil stinger a stingray from the Pliocene (Piacenzian) locality La Serra (Italy), twice as long as the longest caudal spines reported from any living stingray species of the Mediterranean Sea, and possibly representing the longest stingray stinger ever reported from both the fossil and the recent records.[107]
  • A study on the morphology of the marginal dentition of Lophosteus superbus is published by Chen et al. (2020), who reconstruct the dental ontogeny in this taxon, and evaluate its implications for the knowledge of the evolution of teeth of bony fishes.[108]
  • Redescription of the anatomy of mcallisteri is published by Stack et al. (2020).[109]
  • Fragmentary fossil material of Gyrosteus mirabilis is reported from the Toarcian of the Ahrensburg erratics assemblage (Schleswig-Holstein, Germany) by Hornung & Sachs (2020), expanding known geographic range of this species, and representing the first record of a chondrosteid species beyond its type area.[110]
  • Redescription of the skeletal anatomy of longidorsalis is published by Hilton, Grande & Jin (2020).[111]
  • Revision and a study on the phylogenetic relationships of members of the subfamily Pycnodontinae is published by Poyato-Ariza (2020).[112]
  • A study on the skeletal anatomy and phylogenetic relationships of decorata is published by Taverne (2020).[113]
  • A study on the degree of preservation of the skin of an aspidorhynchid specimen from the Barremian Paja Formation (Colombia), representing the first instance of soft tissue preservation in vertebrates from the Early Cretaceous of northern South America, is published by Alfonso-Rojas & Cadena (2020).[114]
  • A study on the diversity and distribution of non-marine teleost fishes in the Western Interior of North America during the late Maastrichtian, based on fossils from the Hell Creek Formation, the Lance Formation and the Scollard Formation, is published online by Brinkman et al. (2020).[115]
  • Description of new fossil material of libanicus and byblosi, providing new information on the skeletal anatomy of these taxa, is published by Taverne & Capasso (2020).[116]
  • Fossil material of Xiphactinus is described from the latest Maastrichtian Salamanca Formation (Chubut Province, Argentina) by De Pasqua, Agnolin & Bogan (2020), representing the first record of this genus from southern part of South America.[117]
  • A methodology for assessing locomotion energetics in extinct bony fishes is presented by Ferrón (2020), who interprets his findings as providing evidence of endothermy in Xiphactinus audax.[118]
  • Redescription and a study on the phylogenetic relationships of ancestralis is published by Brito, Figueiredo & Leal (2020).[119]
  • A study on the skeletal anatomy of Pirskenius, aiming to resolve whether can be sustained as a separate family, is published by Reichenbacher et al. (2020).[120]
  • A study aiming to infer the genetic basis of the reduction of pelvic skeleton in a Miocene stickleback fish is published by Stuart, Travis & Bell (2020).[121]
  • A fish larva sharing anatomical similarities with the so‐called tholichthys larval stage of butterflyfishes is described from the Eocene (Bartonian) locality of Gornyi Luch (Krasnodar Krai, Russia) by Carnevale & Bannikov (2020).[122]
  • New fossil material of Mawsonia gigas, including one of the anatomically most informative specimens referable to the genus Mawsonia, is described from the Mesozoic Tacuarembó Formation (Uruguay) by Toriño et al. (2020).[123]
  • New fossil material of Axelrodichthys megadromos is described from several Campanian and Maastrichtian sites in southern France by Cavin et al. (2020), who present a reconstruction of the skull of this species, and study its phylogenetic relationships and ecology.[124]
  • Redescription of the anatomy of the skull of edentatus is published by Mondéjar‐Fernández, Friedman & Giles (2020).[125]
  • Description of new material of tristichopterids from the Devonian (Famennian) locality of Strud (Belgium), and a study on the phylogenetic relationships of tristichopterids, is published by Olive et al. (2020).[126]
  • Description of a new, 1.57-metre-long articulated specimen of Elpistostege watsoni from the Upper Devonian of Canada, and a study on the implications of this specimen for the knowledge of the early evolution of the vertebrate hand, is published by Cloutier et al. (2020).[127]
  • A study aiming to determine the potential significance of tides for the evolution of bony fish and early tetrapods from the Late Silurian to early Late Devonian is published by Byrne et al. (2020).[128]
  • Evidence of enhanced fish production during the extreme global warmth of the early Paleogene is presented by Britten & Sibert (2020).[129]
  • A study aiming to determine the impact of changes in the Earth system during the Eocene–Oligocene transition on pelagic fish production and biodiversity is published by Sibert et al. (2020).[130]

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