Machaeroides

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Machaeroides
Temporal range: 54.9–46.2 Ma early to middle Eocene
Machaeroides eothen.JPG
Machaeroides eothen skull
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Oxyaenodonta
Family: Oxyaenidae
Subfamily: Machaeroidinae
Genus: Machaeroides
Matthew, 1909
Type species
Machaeroides eothen
Matthew, 1909
Species
  • M. eothen (Matthew, 1909)[1]
  • M. simpsoni (Dawson, 1986)[2]

Machaeroides ("dagger-like") is a genus of sabre-toothed predatory mammal that lived during the Eocene (54.9 to 46.2 mya). Its fossils were found in the U.S. state of Wyoming.

Description[]

Both species bore a passing or superficial resemblance to a very small, dog-sized saber-toothed cat. Machaeroides could be distinguished from actual saber-toothed cats by their more-elongated skulls, and their plantigrade stance. Machaeroides species are distinguished from the closely related by the fact that the former genus had smaller saber-teeth. Despite its small size, the genus Machairoides was well-equipped to hunt prey larger than itself, such as the small, primitive horses and rhinoceroses present at the time, as it was equipped with saber teeth and powerful forelimbs to subdue prey.[3]

M. eothen weighed an estimated 10–14 kg (22–31 lb), thus matching in size a small Staffordshire Terrier. M. simpsoni was probably smaller.[4]

Taxonomic placement[]

Its position within the mammals has been in dispute. Experts have been equally divided over whether Machaeroides and its sister-genus, , belong in Oxyaenidae or Hyaenodonta, though the most recent studies favor the former.[5]

Phylogeny[]

The phylogenetic relationships of the genus Machaeroides are shown in the following cladogram.[6][7][8][9]

 Pan-Carnivora 
 †Hyaenodonta 

Hyaenodonta (sensu stricto) Hyaenodon horridus by R. B. Horsfall (coloured).jpg

Altacreodus

Altacreodus/Tinerhodon clade
 sensu lato 

Carnivoramorpha Dogs, jackals, wolves, and foxes (Plate XI).jpg

 †Oxyaenodonta 
 †Oxyaenidae 

Tytthaeninae 

Oxyaeninae Patriofelis ferox by R. B. Horsfall (coloured).png

 †Machaeroidinae 

†Machaeroidinae sp. (CM 2386)

†Machaeroidinae sp. (FMNH PM 1506)

 ? 

 †Machaeroides 

Machaeroides eothen

Machaeroides simpsoni

References[]

  1. ^ W. D. Matthew. (1909.) "The Carnivora and Insectivora of the Bridger Basin, middle Eocene." Memoirs of the American Museum of Natural History 9:289-567
  2. ^ M. R. Dawson, R. K. Stucky, L. Krishtalka and C. C. Black. (1986.) "Machaeroides simpsoni, new species, oldest known sabertooth creodont (Mammalia), of the Lost Cabin Eocene." Contributions to Geology, University of Wyoming, Special Paper 3:177-182
  3. ^ Antón, Mauricio (2013). Sabertooth. Indiana University Press. ISBN 978-0-253-01049-0.
  4. ^ Egi, Naoko1 (2001). "Body Mass Estimates in Extinct Mammals from Limb Bone Dimensions: the Case of North American Hyaenodontids". Palaeontology. 44 (3): 497–528. doi:10.1111/1475-4983.00189.
  5. ^ Zack, S. (2014). "Saber-tooth origins: a new skeletal association and the affinities of Machaeroidinae (Mammalia, Creodonta)". Journal of Vertebrate Paleontology, Program and Abstracts: 259–260.
  6. ^ Gunnel, Gregg F.; Gingerich, Philip D. (1991). "Systematics and evolution of late Paleocene and early Eocene Oxyaenidae (Mammalia, Creodonta) in the Clarks Fork Basin, Wyoming" (PDF). Contributions from the Museum of Paleontology. The University of Michigan. 28 (7): 141–180.
  7. ^ Solé, Floréal; Ladevèze, Sandrine (2017). "Evolution of the hypercarnivorous dentition in mammals (Metatheria,Eutheria) and its bearing on the development of tribosphenic molars". Evolution & Development. 19 (2): 56–68. doi:10.1111/ede.12219. PMID 28181377.
  8. ^ Prevosti, F. J. & Forasiepi, A. M. (2018.) "Introduction. Evolution of South American Mammalian Predators During the Cenozoic: Paleobiogeographic and Paleoenvironmental Contingencies"
  9. ^ Shawn P. Zack (2019). "A skeleton of a Uintan machaeroidine 'creodont' and the phylogeny of carnivorous eutherian mammals". Journal of Systematic Palaeontology. 17 (8): 653–689. doi:10.1080/14772019.2018.1466374.
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