2021 in paleoentomology

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This paleoentomology list records new fossil insect taxa that are to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

Coleoptera[]

Adephaga[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[1]	Gen. et sp. nov	Valid	Kirichenko-Babko, Perkovsky & Vasilenko	Eocene	Rovno amber	Ukraine	A ground beetle belonging to the tribe Lebiini. Genus includes new species A. pripiatiensis.
^[2]	Gen. et sp. nov	Valid	Schmidt & Maddison in Schmidt, Scholz & Maddison	Eocene	Baltic amber	Europe (Baltic Sea region)	A ground beetle belonging to the subfamily Trechinae. The type species is B. kerneggeri.
^[3]	Sp. nov	Valid	Lee & Nam	Early Miocene		South Korea	A species of Calosoma.

Archostemata[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[4]	Sp. nov	Valid	Yan, Strelnikova & Cai	Early Cretaceous (Berriasian-Barremian)	Gusinoe Ozero Group	Russia ( Buryatia)	A Jurodid beetle.
Kirejtomma^[5]	Gen. et comb. nov	Valid	Li & Cai	Cretaceous	Burmese amber	Myanmar	An Ommatid beetle The type species is "Clessidromma" zengi (2020).
Limnomma^[6]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Middle Jurassic	Haifanggou Formation	China	An Ommatid beetle. The type species is L. daohugouense.
^[7]	Sp. nov	In press	Jarzembowski, Zheng & Zhao	Cretaceous	Burmese amber	Myanmar	An Omma species Ommatid beetle.
^[8]	Sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	An Omma species Ommatid beetle.
Paraodontomma leptocristatum^[9]	Sp. nov	Valid	Li, Yamamoto & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	An Ommatid beetle.
^[10]	Gen. et comb. et sp. nov	Valid	Ponomarenko	Late Permian		Russia	A . The type species is "" sibiricus (2013), as well as new species T. minimus.
^[10]	Sp. nov	Valid	Ponomarenko	Late Permian		Russia	A .
Zygadenia alexrasnitsyni^[11]	Sp. nov	Valid	Strelnikova & Yan	Early Cretaceous	Zaza Formation	Russia	An Ommatid beetle.

Myxophaga[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
^[10]	Gen. et sp. nov	Valid	Ponomarenko	Late Permian		Russia	A Coleocatiniid beetle. The type species is P. tomiensis.
Triamyxa^[12]	Fam., Gen. et sp. nov	Valid	Qvarnström et al.	Late Triassic (Carnian)	Keuper succession (Drawno Beds)	Poland	A Triamyxid beetle in the suborder Myxophaga. The type species is T. coprolithica.

Polyphaga[]

Bostrichiformia[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[13]	Gen. comb. et 3 sp. nov	Valid	Li & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	A nosodendrid beetle. The type species is "Nosodendron" cretaceum; genus also includes new species A. angulare, A. explanatum and A. remotidens.
^[14]	Gen. et comb. nov	Valid	Háva	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A carpet beetle. The type species is "Megatoma" atypica (2017).
^[15]	Gen. et sp. nov	Valid	Tihelka et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A wounded-tree beetle. The type species is M. scandens.
^[16]	Sp. nov	Valid	Alekseev & Bukejs	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A Petalium species death watch beetle.
^[16]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Petalium species death watch beetle.
^[17]	Sp. nov	Valid	Perkovsky, Háva & Zaitsev	Middle Eocene		Russia	A Trogoderma species carpet beetle.
^[14]	Gen. et sp. nov	Valid	Háva	Late Cretaceous Cenomanian)	Burmese amber	Myanmar	A carpet beetle. The type species is T. burmitica.
^[18]	Sp. nov	Valid	Bukejs, Alekseev & Háva	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Xyletinusspecies death watch beetle.

Cucujiformia[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[19]	Sp. nov	In press	Kolibáč et al.	Eocene	Baltic amber	Europe	A checkered beetle.
^[20]	Gen. et sp. nov	Valid	Tshernyshev	Late Eocene	Baltic amber	Europe	A Malachiinae malachite beetle. The type species is A. intercalaris.
^[21]	Sp. nov	In press	Legalov & Poinar	Eocene	Baltic amber	Europe	A Polydrusini Entiminae broad nosed weevil.
^[22]	Gen. et sp. nov	In press	Legalov	Late Cretaceous (Santonian-Campanian)	Arkagala Formation	Russia	A ithycerid weevil. The type species is A. zherikhini.
^[23]	Sp. nov	Valid	Nabozhenko & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	An Alleculinae darkling beetle.
^[24]	Sp. nov	In press	Legalov	Early Cretaceous (Aptian-Albian)	Santana Group	Brazil	An orsodacnid leaf beetle.
^[25]	Sp. nov	Valid	Bukejs & Legalov	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A tooth-nosed snout weevil.
^[26]	Gen. et 3 sp. nov	Valid	Szawaryn	Eocene	Baltic amber	Europe	A Microweiseini Microweiseinae lady beetle. The type species is B. damzeni; genus also includes B. damgaardi and B. szadziewskii.
^[27]	Gen. et sp. nov	In press	Poinar & Legalov	Miocene (Burdigalian)	Dominican amber	Dominican Republic	A Molytinae weevil. The type species is B. adunatus.
^[28]	Sp. nov	Valid	Alekseev & Bukejs	Late Eocene	Rovno amber	Ukraine	A silvanid beetle.
^[29]	Sp. nov	Valid	Legalov et al.	Late Eocene	Rovno amber	Ukraine	A weevil.
^[30]	Gen. et sp. nov	Valid	Li & Cai	Cenomanian	Burmese amber	Myanmar	A Colydiinae? Zopheridae tenebrionoid beetle. The type species is C. rasnitsyni.
^[31]	Gen. et sp. nov	In press	Zhao, Liu & Yu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A prionocerid beetle. The type species is C. burmensis.
^[32]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene		Ethiopia	A Platypodinae weevil.
^[32]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene		China	A Platypodinae weevil.
^[33]	Sp. nov	Valid	Legalov, Nazarenko & Perkovsky	Late Eocene	Rovno amber	Ukraine	A species of Dorytomus Curculioninae weevil.
^[34]	Gen. et sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle. The type species is E. chronographica.
^[35]	Gen. et sp. nov	Valid	Vitali	Eocene	Baltic amber	Europe (Baltic Sea region)	A longhorn beetle belonging to the tribe Crossotini. The type species is E. alekseevi.
^[36]	Sp. nov	Valid	Legalov, Nazarenko & Perkovsky	Eocene (Priabonian)	Rovno amber	Ukraine	A zygaenodinin anthribine anthribid fungus weevil.
^[37]	Sp. nov	Valid	Telnov et al.	Late Eocene Priabonian	Rovno amber	Ukraine	A mycterid palm beetle.
^[38]	Gen. et sp. nov	Valid	Yu, Kolibáč & Ślipiński in Yu et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A lophocaterid beetle. The type species is G. burmiticus.
^[39]	Sp. nov	Valid	Kupryjanowicz, Lyubarsky & Perkovsky	Eocene	Rovno amber	Ukraine	A Heterhelus species Kateretid short-winged flower beetle.
^[40]	Sp. nov	Valid	Nabozhenko & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	An Isomira species claw comb darkling beetle.
^[41]	Gen. et sp. nov	Valid	Legalov, Kupryjanowicz & Perkovsky	Late Eocene Priabonian	Baltic amber	Poland	A cossonine weevil. The type species is K. wanati.
^[42]	Gen. et 2 sp. nov	Valid	Tihelka et al.	Late Cretaceous Cenomanian	Burmese amber	Myanmar	A tenebrionoid beetle of uncertain phylogenetic placement. Genus includes new species K. enigmaticus and K. magnus.
^[43]	Gen. et sp. nov	In press	Kirejtshuk & Reid	Eocene	Baltic amber	Europe	A palophagine megalopodid leaf beetle. The type species is L. andreyi.
^[34]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
^[44]	Gen. et sp. nov	Valid	Batelka & Prokop	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A wedge-shaped beetle. The type species is M. serraticornis.
^[45]	Sp. nov	Valid	Nabozhenko, McKellar & Bukejs	Late Eocene Priabonian	Baltic amber	Europe	AMetaclisa species metaclisine darkling beetle.
^[34]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
^[46]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	A Trogossitid bark-gnawing beetle. The type species is M. qizhihaoi.
^[47]	Sp. nov	Valid	Bukejs et al.	Late Eocene Priabonian	Bitterfeld amber	Germany	A Monolepta species skeletonizing leaf beetle.
^[48]	Nom. nov	Valid	Zhang	Jurassic	Jiulongshan Formation	China	A Trogossitid bark-gnawing beetle. A replacement name for "" Yu et al. (2014).
^[49]	Sp. nov	Valid	Szawaryn, Sontag & Kubisz	Late Eocene Priabonian	Baltic amber	Europe	An Oedemera species false blister beetle.
^[50]	Gen. et sp. nov	Valid	Tihelka, Huang & Cai	Late Cretaceous Cenomanian	Burmese amber	Myanmar	A sooty mould beetle. The type species is P. azari.
^[51]	Gen. et sp. nov	Valid	Legalov	Eocene (Ypresian)	Green River Formation	United States ( Utah)	A Sagrinae leaf beetle. The type species is P. monrosi.
^[52]	Trib., Gen., et sp. nov	Valid	Legalov	Late Eocene Priabonian	Baltic amber	Europe	A Curculioninae weevil. The type species is P. horridus.
^[53]	Gen. et sp. nov	Valid	Poinar, Brown & Legalov	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A new york weevil. The type species is P. micrommatus.
^[54]	Gen. et sp. nov	Valid	Powell & Cline	Miocene ()	Dominican amber	Dominican Republic	A Cillaeinae sap beetle. The type species is P. cascus.
^[55]^[56]^[57]	Gen. et sp. nov	Valid	Tihelka et al.	Cretaceous	Burmese amber	Myanmar	A short-winged flower beetle. The type species is P. vivificus .
^[34]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A Phloiotrya species false darkling beetle.
^[58]	Gen. et sp. nov	Valid	Tihelka et al.	Late Cretaceous (Cenomanian)		France	A soft-wing flower beetle. The type species is P. cretaceus.
^[59]	Gen. et 2 sp. nov	Valid	Tshernyshev in Tshernyshev & Shcherbakov	Late Eocene	Baltic amber Rovno amber	Baltic Sea region Ukraine	A Dasytinae malachite beetle. The type species P. chaetophorata Genus also includes P. mikhailovi.^[60]
^[61]	Gen. et sp. nov	Valid	Bukejs, Ślipiński & Alekseev in Bukejs et al.	Eocene	Baltic amber	Europe	A cerylonid beetle. The type species is P. pandema.
^[25]	Sp. nov	Valid	Bukejs & Legalov	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Rhynchitinae tooth-nosed snout weevil.
^[62]	Gen. et sp. nov	Valid	Krell & Vitali	Eocene (Ypresian)	Green River Formation	United States ( Colorado)	A Sagrinae leaf beetle. The type species is P. attenboroughi.
^[63]	Trib., gen. et sp. nov	Valid	Poinar & Brown	Cretaceous Cenomanian	Burmese amber	Myanmar	A pine flower weevil. The type species is R. legalovii.
^[44]	Gen. et comb. nov	Valid	Batelka & Prokop	Late Cretaceous (Cenomanian)		France	A wedge shaped beetle. The type species is "Macrosiagon" ebboi (2004).
^[34]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
^[34]	Sp. nov	Valid	Alekseev & Bukejs	Middle Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A false darkling beetle.
^[64]	Sp. nov	In press	Alekseev et al.	Late Eocene Priabonian	Baltic amber	Europe	A teredid beetle.
^[65]	Sp. nov	Valid	Legalov	Late Eocene Priabonian	Baltic amber	Europe	An Apioninae straight-snouted weevil.
^[66]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A slime mold beetle. The type species is T. newtoni
^[67]	Gen. et sp. nov	Valid	Lyubarsky et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	A lophocaterid beetle. The type species is Z. pankowskiorum.
^[65]	Gen. et sp. nov	Valid	Legalov	late Eocene Priabonian	Baltic amber	Europe	A straight-snouted weevil. The type species is Z. rarus.

Elateriformia[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[68]	Sp. nov	Valid	Poinar, Vega & Legalov	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cerophytid rare click beetle.
^[69]	Gen. et sp. nov	Valid	Li, Kundrata & Cai in Li et al.	Cretaceous (Albian to Cenomanian)	Burmese amber	Myanmar	An elateroid click beetle of uncertain phylogenetic placement. The type species is A. lawrencei.
^[70]	Gen. et sp. nov	Valid	Fanti	Eocene	Prussian Formation (Baltic amber)	Poland	A soldier beetle belonging to the tribe Cantharini. The type species is A. pankowskiarum.
^[71]	Sp. nov	Valid	Muona	Eocene	Baltic amber	Europe	A Melasinae false click beetle.
^[72]	Gen. et sp. nov	Valid	Kundrata et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A dascillid beetle. The type species is B. serraticornis.
^[73]	Gen. et sp. nov	Valid	Jiang, Song & Wang in Jiang et al.	Cretaceous	Burmese amber	Myanmar	A metallic wood-boring beetle. The type species is B. cretacus.
^[74]	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber	Myanmar	A soldier beetle.
^[74]	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber	Myanmar	A soldier beetle.
^[74]	Sp. nov	In press	Yang et al.	Cretaceous	Burmese amber	Myanmar	A soldier beetle.
^[75]	Gen. et sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A throscid beetle. The type species is C. depressiceps.
^[76]	Sp. nov	Valid	Alekseev & Bukejs in Alekseev et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A turtle beetle.
^[76]	Sp. nov	Valid	Alekseev & Bukejs in Alekseev et al.	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A turtle beetle.
^[77]	Sp. nov	Valid	Poinar & Háva	Burdigalian	Dominican amber	Dominican Republic	A turtle beetle.
^[71]	Sp. nov	Valid	Muona	Eocene	Baltic amber	Europe	A false click beetle.
^[78]	Gen. et sp. nov	Valid	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A soldier beetle. The type species is C. veda.
^[79]	Gen. et sp. nov	Valid	Kundrata et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	An Agrypninae click beetle. The type species is C. burmitinus.
^[80]	Fam. nov., Gen. et sp. nov	Valid	Li, Kundrata, Tihelka & Cai in Li et al.	Cretaceous (Albian or Cenomanian)	Burmese amber	Myanmar	A elateroid beetle. The type species is C. azari.
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe (Baltic Sea region)	A false click beetle.
^[75]	Gen. et sp. nov	Valid	Li & Cai	Cretaceous Cenomanian	Burmese amber	Myanmar	A throscid false click beetle. The type species is E. yanpingae.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe (Baltic Sea region)	A Melasinae false click beetle.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An ? false click beetle.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An ? false click beetle.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An ? false click beetle.
^[71]	Gen. et sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An false click beetle. The type species is E. magnus.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An false click beetle.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An false click beetle.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An false click beetle.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An false click beetle.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	An false click beetle.
^[78]	Sp. nov	Valid	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A soldier beetle.
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	An Isorhipis species melasine false click beetle
^[81]	Sp. nov	Valid	Kazantsev	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A soldier beetle.
^[81]	Sp. nov	Valid	Kazantsev	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	A soldier beetle
^[82]	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber	Poland	A soldier beetle.
^[82]	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber	Poland	A soldier beetle, a species of Malthodes.
^[83]	Sp. nov	Valid	Pintilioaie, Fanti & Ionesi	Middle Miocene		Romania	A soldier beetle.
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Microrhagusspecies melasine false click beetle
^[84]	Gen. et sp. nov	Valid	Li & Cai in Li et al.	Cretaceous Cenomanian	Burmese amber	Myanmar	A net-winged beetle. The type species is M. longiantennus.
^[71]	Sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	A false click beetle.
^[71]	Gen. et sp. nov	Valid	Muona	Middle Eocene Priabonian	Baltic amber	Europe	A ? false click beetle. The type species is N. balticus.
^[78]	Gen. et sp. nov	Valid	Hsiao et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A soldier beetle. The type species is P. panna.
^[82]	Sp. nov	Valid	Fanti	Late Eocene Priabonian	Baltic amber	Poland	A soldier beetle.
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A false click beetle.
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A false click beetle.
^[85]	Gen. et sp. nov	In press	Li, Huang & Cai	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A water-penny beetle. The type species is P. burmensis.
^[75]	Gen. et sp. nov	Valid	Li & Cai in Li, Huang & Cai	Cretaceous ( Cenomanian)	Burmese amber	Myanmar	A throscid false click beetle. The type species is P. robustus.
^[86]	Sp. nov	Valid	Kundrata, Bukejs & Blank in Kundrata et al.	Late Eocene Priabonian	Baltic amber	Russia ( Kaliningrad Oblast)	APtilodactyla species toe winged beetle.
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Rhacopus species eucnemid false click beetle
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Melasinae false click beetle.
^[87]	Gen. et sp. nov		Li, Chang & Muona	Early Cretaceous (Hauterivian to Aptian)	Yixian Formation	China	An false click beetle. The type species is R. brevicornis.
^[88]	Sp. nov	Valid	Fanti & Pankowski	Burdigalian	Dominican amber	Dominican Republic	A species of Silis.
^[88]	Sp. nov	Valid	Fanti & Pankowski	Burdigalian	Dominican amber	Dominican Republic	A species of Silis.
^[71]	Sp. nov	Valid	Muona	Late Eocene Priabonian	Baltic amber	Europe	A Xylophilus species eucnemid false click beetle.

Scarabaeiformia[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
^[89]	Gen. et sp. nov	Valid	Matalin, Perkovsky & Vasilenko	Late Eocene Priabonian	Rovno amber	Ukraine	A tiger beetle. The type species is G. fungifora.
^[90]	Sp. nov	Valid	Tello et al.	Late Pleistocene	Pilauco Bajo	Chile	A scarab beetle.

Staphyliniformia[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[91]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Acritus.
^[92]	Sp. nov	Valid	Alekseev & Bukejs	Eocene	Baltic amber	Poland	A species of Bacanius.
^[93]	Gen. et sp. nov	Valid	Yin & Cai	Cretaceous	Burmese amber	Myanmar	A Pselaphinae rove beetle. The type species is B. rougemonti.
^[94]	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber	Myanmar	An Onthophilinae clown beetle. The type species is C. maddisoni.
^[95]	Sp. nov	In press	Yamamoto & Shavrin	Eocene	Baltic amber	Europe	A Phloeocharinae rove beetle.
^[96]	Gen. et sp. nov	In press	Sohn & Nam	Early Cretaceous	Jinju Formation	South Korea	A carrion beetle. The type species is C. jinjuensis.
^[94]	Gen. et sp. nov	Valid	Caterino	Cenomianian	Burmese amber	Myanmar	A Dendrophilinae clown beetle. The type species is D. aeterna.
^[97]	Sp. nov	Valid	Jałoszyński & Perkovsky	Eocene	Rovno amber	Ukraine	A Euconnus species Scydmaeninae rove beetle.
^[98]	Sp. nov	Valid	Shavrin in Shavrin & Kairišs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Eusphalerum species Omaliinae rove beetle.
^[99]	Gen. et sp. nov	Valid	Yin, Zhao & Cai	Cretaceous Cenomanian	Burmese amber	Myanmar	A Pselaphinae rove beetle. The type species is M. spinitibialis.
^[100]	Gen. et sp. nov	Valid	Cai	Early Cretaceous	Yixian Formation	China	A rove beetle. The type species is P. fossilis.
^[94]	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber	Myanmar	An Onthophilinae clown beetle. The type species is P. cristatus.
^[101]	Gen. et sp. nov	Valid	Prokin & Strelnikova	Early Cretaceous (Hauterivian)		Russia	A water scavenger beetle. The type species is P. buryaticus.
^[102]	Sp. nov	Valid	Perreau & Haelewaters in Perreau, Haelewaters & Tafforeau	Miocene Burdigalian	Dominican amber	Dominican Republic	A Ptomaphagini round fungus beetle.
^[103]	Sp. nov	Valid	Yamamoto & Newton	Cretaceous (Cenomanian)	Burmese amber	Myanmar	A Trichophya species rove beetle.
^[94]	Gen. et sp. nov	Valid	Caterino	Cretaceous	Burmese amber	Myanmar	A Dendrophilinae clown beetle. The type species is Y. peregrina.

Coleopteran research[]

A study on the evolution of beetles from the Early Permian to Middle Triassic, evaluating the impact of the Permian–Triassic extinction event on beetle diversity, is published by Zhao et al. (2021).^[104]
A study on the anatomy and phylogenetic relationships of funestus is published by Li et al. (2021).^[105]
Revision of Notocupes undatabdominus and N. ? multituberatus from the Lower Cretaceous of South China is published by Li, Huang & Cai (2021), who argue that both species do not belong to the genus , transfer N. ? multituberatus to the genus , and consider "N." undatabdominus to be likely a member of the family Artematopodidae.^[106]
A piece of Burmese amber preserving the complete series of free-living stages (except of the last larval instar) of a Cretaceous parasitoid beetle is described by Batelka, Engel & Prokop (2021).^[107]
The oldest solid wood-borer larvae reported to date (belonging to the families Buprestidae and possibly Cerambycidae) are described from the Cretaceous amber from Myanmar by Haug et al. (2021).^[108]
A study on the morphological diversity of larvae of extant and fossil members of the family Brachypsectridae is published by Haug et al. (2021).^[109]
Mitochondrial genomes and nuclear ribosomal DNA of beetles, in some cases identifiable to species, are recovered from pack rat middens from California (United States) and Baja California (Mexico) dating up to ~ 34,355 years before present by Smith et al. (2021).^[110]

Dermapterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[111]	Gen. et 2 sp. nov	Valid	Xiong et al.	Middle Jurassic	Jiulongshan Formation	China	A semenoviolid eodermapteran earwig. Genus includes the new species A. elongatum and A. cylindratum.
^[112]	Gen. et sp. nov	Valid	Xiong, Engel & Ren in Xiong et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Protodiplatyidae. The type species is A. oiodes.
^[113]	Gen. et comb. nov	Valid	Engel	Early Jurassic (Sinemurian)	Charmouth Mudstone Formation	United Kingdom	A dermapterid earwig. The type species is "Brevicula" teres Tihelka (2019).
^[114]	Gen. et sp. nov	Valid	Chen & Zhang	Cenomanian	Burmese amber	Myanmar	A pygidicranid earwig. Genus includes new species E. myanmarensis.
^[112]	Sp. nov	Valid	Xiong, Engel & Ren in Xiong et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Dermapteridae.

Dictyopterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[115]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A cockroach belonging to the family Corydiidae. Genus includes new species A. extincta.
^[116]	Sp. nov	In press	Sendi in Vršanský et al.	Cretaceous	Burmese amber	Myanmar	An alienopterid.
Alienopterix smidovae^[116]	Sp. nov	In press	Hinkelman in Vršanský et al.	Cretaceous	Burmese amber	Myanmar	An alienopterid.
^[117]	Fam. et gen. et sp. nov	Valid	Engel & Jiang in Jiang et al.	Cretaceous	Burmese amber	Myanmar	A termite belonging to the new family . Genus includes new species A. hospitis.
^[118]	Gen. et sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Ectobiidae. Genus includes new species C. hinkelmani.
^[119]	Gen. et 2 sp. nov	In press	Liang et al.	Middle Jurassic	Jiulongshan Formation	China	A cockroach belonging to the family . Genus includes new species C. huntianlingensis and C. lata.
^[120]	Gen. et sp. nov	In press	Hinkelman	Cretaceous	Burmese amber	Myanmar	A cockroach. Genus includes new species C. brevialata .
^[118]	Gen. et sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Ectobiidae. Genus includes new species E. protecta.
^[115]	Sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Blattulidae.
^[121]	Gen. et sp. nov	Valid	Chen, Zhang Shi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An alienopterid. Genus includes new species E. burma.
^[122]	Fam. et gen. et 2 sp. nov	Valid	Hinkelman & Vršanský in Vršanský et al.	Late Jurassic–Late Cretaceous (Kimmeridgian–Cenomanian)	Burmese amber	Germany Kazakhstan Myanmar	A member of Corydioidea, the type genus of the new family Vršanský & Hinkelman. The type species is F. articulata Hinkelman; genus also includes new species F. aristovi Vršanský.
^[123]	Gen. et sp. nov	Valid	Chen, Shih & Ren in Chen et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Corydiidae. The type species is F. pectinata.
^[124]	Sp. nov	Valid	Bouju, Jouault & Perrichot	Early Miocene	Ethiopian amber	Ethiopia	A termite belonging to the family Kalotermitidae.
^[125]	Gen. et sp. nov	Valid	Schneider et al.	Carboniferous (Kasimovian)	Atrasado Formation	United States ( New Mexico)	A member of Blattodea belonging to the family . The type species is K. huberi.
^[115]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Liberiblattinidae. Genus includes new species K. gondwanensis.
^[115]	Sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Liberiblattinidae.
^[126]	Gen. et sp. nov	Valid	Demers‐Potvin et al.	Late Cretaceous (Cenomanian)	Redmond Formation	Canada	A praying mantis belonging to the family Baissomantidae. Genus includes new species L. guilbaulti.
^[127]	Sp. nov	In press	Vršanský, Poschmann & Vidlička	Late Oligocene		Germany	A species of Latiblattella.
^[118]	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar
^[128]	Gen. et sp. nov	Valid	Luo et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Umenocoleidae. Genus includes new species L. liuyani.
^[129]	Gen. et sp. nov	Valid	Perkovsky & Nel	Eocene (Priabonian)	Rovno amber	Ukraine	A termite belonging to the family Rhinotermitidae. Genus includes new species L. milescaput.
Meiatermes cretacicus^[130]	Sp. nov	In press	Bezerra et al.	Early Cretaceous	Crato Formation	Brazil	A termite.
^[115]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Mesoblattinidae. Genus includes new species M. brezinica.
^[131]	Gen. et sp. nov	Valid	Jouault et al.	Cretaceous	Burmese amber	Myanmar	A termite belonging to the family Mastotermitidae. Genus includes new species M. engeli.
^[132]	Gen. et sp. nov	In press	Sendi	Early Cretaceous (Barremian)	Lebanese amber	Lebanon	A cockroach belonging to the family Liberiblattinidae. Genus includes new species M. libera.
^[133]	Sp. nov	In press	Hinkelman	Cretaceous	Burmese amber	Myanmar	A member of the family Mesoblattinidae.
^[134]	Sp. nov	In press	Šmídová, Vidlička & Wedmann	Eocene	Messel pit	Germany	A cockroach belonging to the family Blaberidae.
^[134]	Sp. nov	In press	Šmídová, Vidlička & Wedmann	Cretaceous	Burmese amber	Myanmar	A cockroach belonging to the family Blaberidae.
^[116]	Gen. et sp. nov	In press	Vršanský, Hinkelman & Sendi in Vršanský et al.	Cretaceous	Burmese amber	Myanmar	An alienopterid. Genus includes new species N. postava.
^[135]	Sp. nov	In press	Šmídová	Cretaceous	Burmese amber	Myanmar	A cockroach belonging to the family Corydiidae.
^[122]	Sp. nov	Valid	Koubová & Vršanský in Vršanský et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Corydioidea.
^[135]	Gen. et sp. nov	In press	Šmídová	Cretaceous	Burmese amber	Myanmar	A cockroach belonging to the family . Genus includes new species O. vrsanskyi.
^[115]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of the family Mesoblattinidae. Genus includes new species O. systematicka.
^[136]	Sp. nov	In press	Song et al.	Cretaceous	Burmese amber	Myanmar	A member of belonging to the family .
^[137]	Gen. et sp. nov	Valid	So et al.	Early Cretaceous	Sinuiju Formation	North Korea	A member of the family . Genus includes new species P. coreanica.
^[138]	Sp. nov	Valid	Anisyutkin & Perkovsky	Eocene	Rovno amber	Ukraine	A cockroach, possibly a species of Periplaneta.
^[139]	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation	Japan	A member of the family Umenocoleidae.
^[139]	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation	Japan	A member of the family Mesoblattinidae.
Praeblattella inexpecta^[139]	Sp. nov	In press	Oyama, Yukawa & Imai	Early Cretaceous	Kitadani Formation	Japan	A member of the family Mesoblattinidae.
^[122]	Gen. et 2 sp. nov	Valid	Sendi & Vršanský in Vršanský et al.	Late Jurassic–Early Cretaceous (Kimmeridgian–Barremian)	Lebanese amber	Kazakhstan Lebanon	A member of the family Liberiblattinidae. The type species is P. occisor Sendi; genus also includes new species P. nigroalba Vršanský.
^[131]	Sp. nov	Valid	Jouault et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A termite, a species of Reticulitermes.
^[115]	Gen. et sp. nov	In press	Vršanský et al.	Cretaceous (Hauterivian?)		Algeria	A member of Holocompsinae. Genus includes new species S. equatorialis.
^[140]	Sp. nov	Valid	So et al.	Early Cretaceous	Sinuiju Formation	North Korea	A member of the family Mesoblattinidae.
^[132]	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Liberiblattinidae.
^[132]	Sp. nov	In press	Sendi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A cockroach belonging to the family Liberiblattinidae.
^[117]	Gen. et sp. nov	Valid	Engel & Cai in Jiang et al.	Cretaceous	Burmese amber	Myanmar	A termite belonging to the family . Genus includes new species T. philetaerus.

Dictyopteran research[]

A study on the phylogenetic placement of the family Umenocoleidae, based on data from a new specimen of sinuatus from the Lower Cretaceous Zhonggou Formation (China), is published by Liuo et al. (2021).^[141]
Description of new alienopterid nymphs from the Cretaceous Burmese amber, and a study on life history and affinities of alienopterids, is published by Luo et al. (2021).^[142]
A study on the sensory organs of hui, and on their implications for the knowledge of the ecology of this cockroach, is published by Taniguchi et al. (2021).^[143]

Dipterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[144]	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea belonging to the new family .
Acartophthalmites luridus^[144]	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea belonging to the new family Clusiomitidae.
Acartophthalmites rugosus^[144]	Sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea belonging to the new family Clusiomitidae.
^[145]	Gen. et sp. et comb. nov	Valid	Krzemiński et al.	Cretaceous	Burmese amber	Myanmar	A member of the crane fly subfamily Limoniinae. The type species is A. rasnitsyni; genus also includes "Antocha" lapra Podenas & Poinar (2009).
^[146]	Sp. nov	Valid	Nel	Early Eocene	Oise amber	France	A member of the family Cecidomyiidae.
^[147]	Sp. nov	Valid	Bouju et al.	Early Miocene	Ethiopian amber	Ethiopia	A species of Allodia.
^[148]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Mexican amber	Mexico	A species of Apocephalus.
^[148]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Dominican amber	Dominican Republic	A species of Apocephalus.
^[148]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene			A species of Apocephalus.
^[149]	Sp. nov	In press	Lamas, Sampronha & Ribeiro	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Asilidae.
^[150]	Gen. et sp. nov	Valid	Lukashevich	Middle Triassic	Röt Formation	Germany	A member of the family . Genus includes new species B. franconica.
^[151]	Sp. nov	Valid	Skartveit & Wedmann	Oligocene		Germany	A species of Bibio.
^[152]	Sp. nov	Valid	Skartveit	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Bibio.
^[153]	Gen. et 3 sp. nov	Valid	Ševčík et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Bibionidae. The type species is B. antennatus; genus also includes B. conicus and B. pedicellatus.
^[154]	Gen. et 2 sp. nov	Valid	Ševčík, Hippa & Burdíková	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Ditomyiidae. The type species is B. korneliae; genus also includes B. wieslawi.
^[154]	Gen. et sp. nov	Valid	Ševčík, Hippa & Burdíková	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Sciaroidea of uncertain phylogenetic placement. The type species is B. mesozoica.
^[155]	Gen. et sp. nov	Valid	Poinar & Vega	Cretaceous (Albian/Cenomanian)	Burmese amber	Myanmar	A member of the family Apsilocephalidae. The type species is C. tubuliferous.
^[144]	Fam. et gen. et comb. et sp. nov	Valid	Roháček & Hoffeins	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A member of Opomyzoidea, the type genus of the new family Clusiomitidae. The type species is "" clusioides Roháček (2016); genus also includes new species C. ornatus.
^[156]	Sp. nov	Valid	Liu & Huang	Middle–Late Jurassic	Jiyuan Basin	China	A member of the family Ptychopteridae.
^[157]	Sp. nov	Valid	Lukashevich, Amorim & Ribeiro	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Bibionidae.
^[158]	Sp. nov		Kopeć in Kopeć et al.	Jurassic/Cretaceous boundary	Glushkovo Formation	Russia ( Zabaykalsky Krai)	A member of the subfamily .
Cretolimonia lukashevichae^[158]	Sp. nov		Kopeć & Krzemiński in Kopeć et al.	Jurassic/Cretaceous boundary	Dain Formation	Russia ( Zabaykalsky Krai)	A member of the subfamily Architipulinae.
Cretolimonia mikolajczyki^[158]	Sp. nov		Kopeć, Krzemiński, Soszyńska-Maj in Kopeć et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the subfamily Architipulinae.
Cretolimonia pseudojurassica^[158]	Sp. nov		Krzemiński in Kopeć et al.	Jurassic/Cretaceous boundary	Dain Formation	Russia ( Zabaykalsky Krai)	A member of the subfamily Architipulinae.
^[159]	Gen. et sp. nov	In press	Carmo, Lamas & Ribeiro	Early Cretaceous	Crato Formation	Brazil	A member of the family Therevidae. Genus includes new species C. antiqua.
^[160]	Gen. et sp. nov	Valid	Li, Zhang & Xiao	Cretaceous	Burmese amber	Myanmar	A member of the family Bibionidae. Genus includes new species C. burmensis.
^[161]	Sp. nov	Valid	Giłka, Harbach & Perkovsky	Eocene	Rovno amber	Ukraine	A mosquito, a species of Culex.
^[162]	Sp. nov	In press	Peñalver et al.	Eocene	Anglesea amber	Australia	A species of Culicoides.
^[163]	Sp. nov	Valid	Kania-Kłosok et al.	Eocene	Green River Formation	United States	A member of the family Cylindrotomidae.
^[163]	Sp. nov	Valid	Kania-Kłosok et al.	Eocene	Green River Formation	United States	A member of the family Cylindrotomidae.
^[164]	Gen. et sp. nov	Valid	Krzemiński et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the crane fly subfamily Limoniinae. The type species is D. podenasi.
^[165]	Sp. nov		Kania-Kłosok & Krzemiński	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Elephantomyia.
^[165]	Sp. nov		Kania-Kłosok & Krzemiński	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Elephantomyia.
^[166]	Sp. nov	Valid	Szadziewski, Sontag & Pankowski	Miocene	Ethiopian amber	Ethiopia	A species of Forcipomyia.
^[167]	Sp. nov		Kania-Kłosok, Krzemiński & Arillo	Early Cretaceous (Albian)	Utrillas Group (Álava amber)	Spain	A species of Helius.
^[167]	Sp. nov		Kania-Kłosok, Krzemiński & Arillo	Early Cretaceous (Albian)	Utrillas Group (San Just amber)	Spain	A species of Helius.
^[168]	Sp. nov	Valid	Wedmann et al.	Eocene (Lutetian)	Messel Formation	Germany	A member of the family Nemestrinidae.
^[169]	Sp. nov	Valid	Badano, Zhang, Yeates & Cerretti in Badano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Phoroidea.
^[170]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
^[170]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
^[170]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
^[170]	Sp. nov	Valid	Ribeiro, Santos & Santos	Early Cretaceous	Crato Formation	Brazil	A species of Leptotarsus.
^[171]	Gen. et sp. nov	Valid	Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.	Late Cretaceous (Santonian)	(Taimyr amber)	Russia	A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is M. kaluginae.
^[162]	Sp. nov	In press	Peñalver et al.	Eocene	Anglesea amber	Australia	A member of the family Ceratopogonidae.
^[172]	Sp. nov	In press	Pielowska-Ceranowska, Azar & Szwedo	Early Cretaceous (Barremian)	Lebanese amber	Lebanon	A member of the family Ceratopogonidae.
^[148]	Sp. nov	In press	Solórzano-Kraemer et al.	Miocene	Zhangpu amber	China	A species of Myriophora.
^[173]	Sp. nov	Valid	Wagner	Eocene		Ukraine	A member of the family Psychodidae.
^[174]	Gen. et sp. nov	Valid	Men & Podenas	Cretaceous	Burmese amber	Myanmar	A member of the crane fly subfamily Limoniinae. Genus includes new species O. simplex.
^[171]	Gen. et sp. nov	Valid	Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.	Cretaceous (probably Albian–Cenomanian)	Burmese amber	Myanmar	A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is P. krzeminskii.
^[175]	Sp. nov	Valid	Skibińska, Krzemiński & Zhang in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Palaeoglaesum stebneri^[175]	Sp. nov	Valid	Skibińska & Krzemiński in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
Palaeoglaesum teres^[175]	Sp. nov	Valid	Skibińska & Albrycht in Skibińska et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.
^[176]	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber	Myanmar	A species of Paleoplatyura.
^[176]	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber	Myanmar	A species of Paleoplatyura.
^[176]	Sp. nov	Valid	Ševčík, Krzemiński & Skibińska	Cretaceous	Burmese amber	Myanmar	A species of Paleoplatyura.
^[177]	Gen. et comb. et sp. nov	Valid	Curler & Skibińska	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Psychodidae. The type species is "Pericoma" formosa Meunier (1905); genus also includes new species P. madrizi.
^[178]	Gen. et sp. nov	Valid	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Bibionidae. Genus includes new species P. skartveiti.
^[179]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
^[179]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
^[179]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
^[179]	Sp. nov	Valid	Kania-Kłosok et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Rhabdomastix.
^[146]	Sp. nov	Valid	Nel	Early Eocene	Oise amber	France	A member of the family Cecidomyiidae.
^[180]	Sp. nov	Valid	Zakrzewska & Jankowska	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Stempellinella.
^[181]	Sp. nov	Valid	Kania-Kłosok et al.	Early Cretaceous (Albian)	Utrillas Group	Spain	A species of Trichoneura.
^[182]	Sp. nov	In press	Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Zhangsolvidae.

Dipteran research[]

Baranov et al. (2021) document a high diversity of fly larvae from the Eocene Baltic amber, preserving taxonomically, morphologically, and ecologically important characters, and evaluate the implications of this finding for the knowledge of the functional ecology of the Eocene fly larvae.^[183]

Ephemeroptera[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[184]	Gen. et sp. nov	Valid	Brandão et al.	Early Cretaceous	Crato Formation	Brazil	A mayfly of uncertain phylogenetic placement. Genus includes new species A. cretacica.
^[185]	Gen. et 2 sp. nov	Valid	Godunko, Martynov & Staniczek	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A mayfly. The type species is B. paucivenosa; genus also includes B. clypeata.
^[186]	Sp. nov	In press	Storari et al.	Early Cretaceous (Aptian)	Crato Formation	Brazil	A mayfly.
^[187]	Sp. nov	Valid	Sinitshenkova	Triassic		Russia	A mayfly.

Ephemeropteran research[]

Sroka et al. (2021) provide new information on the morphology of Permian stem-mayflies Misthodotes sharovi and M. zalesskyi from the Tshekarda locality (Kungurian Koshelevka Formation; Perm Krai, Russia), and infer the life history traits of both the adult and larval stages of these insects.^[188]
Possible underwater mayfly burrows are described from the Turonian Ferron Sandstone (Utah, United States) by King et al. (2021), who interpret this finding as evidence that burrowing mayflies adapted to domicile filter-feeding during or prior to the Turonian, and name a new ichnospecies gingrasi.^[189]

Hemipterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[190]	Trib. et gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cixiidae; the type genus of the new tribe . The type species is A. multigranulata Luo & Bourgoin.
^[191]	Sp. nov	Valid	Heiss in Heiss, Kairišs & Bukejs	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Aneurus.
^[192]	Gen. et sp. nov	Valid	Swanson et al.	Eocene	Green River Formation	United States	A member of the family Reduviidae belonging to the subfamily Harpactorinae. Genus includes new species A. danjuddi.
^[193]	Gen. et sp. nov	Valid	Kim, Chérot & Jung	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Miridae. The type species is B. herczeki.
^[190]	Gen. et comb. nov	In press	Bourgoin & Szwedo in Luo et al.	Early Cretaceous	Weald Clay	United Kingdom	A member of the family Cixiidae; a new genus for "Cixius" petrinus Fennah.
^[194]	Sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation	Kyrgyzstan	A member of Cicadomorpha belonging to the superfamily and the family .
Beaconiella pulchra^[194]	Sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation	Kyrgyzstan	A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.
^[195]	Sp. nov	Valid	Souma, Yamamoto & Takahashi	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Tingidae.
Burmacader grandis^[196]	Sp. nov	Valid	Heiss & Guilbert	Cretaceous	Burmese amber	Myanmar	A member of the family Tingidae.
^[197]	Gen. et sp. nov	Valid	Liu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family . Genus includes new species B. liaoyaoi.
^[198]	Gen. et sp. nov	Valid	Qiao et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Tettigarctidae. Genus includes new species B. nervosa.
^[199]	Gen. et sp. nov	Valid	Souma, Yamamoto & Takahashi	Cretaceous (Albian or Cenomanian)	Burmese amber	Myanmar	A member of the family Tingidae. The type species is B. anomalocapitata.
Cretamystilus^[200]	Gen. et sp. nov	Valid	Kim, Lim & Jung	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Miridae belonging to the tribe . Genus includes new species C. herczeki.
^[201]	Sp. nov	In press	Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Mimarachnidae.
^[202]	Gen. et sp. nov	Valid	Fu & Huang	Cretaceous	Burmese amber	Myanmar	A member of the family Sinoalidae. Genus includes new species C. burmensis.
^[203]	Sp. nov	Valid	Zhang, Yao & Pang in Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Mimarachnidae.
^[190]	Gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is D. tibiocoronata Luo & Bourgoin.
^[204]	Sp. nov	Valid	Song, Szwedo & Bourgoin in Song et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A planthopper.
^[205]	Gen. et sp. nov	Valid	Chen et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A leafhopper belonging to the subfamily Ledrinae. Genus includes new species D. electronica.
^[206]	Sp. nov	Valid	Kim & Jung	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Miridae.
^[207]	Gen. et 2 sp. nov	Valid	Wedmann et al.	Eocene	Messel pit	Germany United States ( Colorado)	A member of the family Pentatomidae. The type species is E. peterkulkai; genus also includes E. greenriverensis.
^[208]	Gen. et sp. nov	Valid	Cumming & Le Tirant	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Coreidae belonging to the subfamily Coreinae. The type species is F. excalibur.
^[209]	Sp. nov	In press	Monferran et al.	Early Cretaceous	Crato Formation	Brazil	A leafhopper.
^[210]	Gen. et sp. nov	Valid	Cifuentes-Ruiz & Brailovsky in Cifuentes-Ruiz et al.	Early Miocene	La Quinta Formation (Mexican amber)	Mexico	A member of the family Schizopteridae. The type species is H. bretoni.
^[211]	Fam. et gen. et sp. nov	Valid	Shcherbakov & Emeljanov	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A planthopper belonging to the new family . The type species is I. rapunzel.
^[212]	Sp. nov	Valid	Fu & Huang	Late Jurassic	Daohugou Beds	China	A member of the family Sinoalidae.
^[213]	Nom. nov	Valid	Doweld	Jurassic (Bathonian-Oxfordian)		Kazakhstan	A member of the family ; a replacement name for Carpenterella Becker-Migdisova (1968).
^[214]	Gen. et sp. nov		Wang, Tang & Yao in Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Tingidae. The type species is L. carinbifarium.
Microtingis^[215]	Gen. et sp. nov	Junior synonym	Heiss & Golub	Cretaceous	Burmese amber	Myanmar	A member of the family Tingidae. Genus includes new species M. leptosoma. Subsequently Heiss & Golub (2021) considered this species to be a junior synonym of Burmavianaida anomalocapitata.^[216]
^[203]	Gen. et sp. nov	Valid	Zhang, Yao & Pang in Zhang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Mimarachnidae. The type species is M. orthotropa.
^[217]	Sp. nov	Valid	Heiss & Golub	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Tingidae.
Paleocader pulchellus^[217]	Sp. nov	Valid	Heiss & Golub	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Tingidae.
Paleocader rovnensis^[217]	Sp. nov	Valid	Heiss & Golub	Eocene	Rovno amber	Ukraine	A member of the family Tingidae.
^[218]	Sp. nov	Valid	Golub, Perkovsky & Vasilenko	Late Eocene	Rovno amber	Ukraine	A member of the family Tingidae.
^[190]	Gen. et sp. nov	In press	Bourgoin & Luo in Luo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is P. kachinensis Luo & Bourgoin.
^[219]	Gen. et sp. nov	Valid	Kóbor & Roca-Cusachs	Cretaceous	Burmese amber	Myanmar	A member of the family Yuripopovinidae. Genus includes new species P. longicornis.
^[220]	Gen. et sp. nov	In press	Tang, Wang & Yao in Tang et al.	Cretaceous	Burmese amber	Myanmar	A member of the family . Genus includes new species P. carinalis.
^[221]	Nom. nov	Valid	Kment	Late Oligocene		Germany	A member of Lygaeoidea of uncertain phylogenetic placement; a replacement name for Pyrrhocoris tibialis Statz & Wagner (1950).
^[222]	Gen. et sp. nov	Valid	Dietrich, Dmitriev & Perkovsky	Late Eocene	Rovno amber	Ukraine	A leafhopper. Genus includes new species R. rasnitsyni.
^[223]	Sp. nov	In press	Luo, Liu & Jarzembowski in Luo et al.	Early Cretaceous	Weald Clay	United Kingdom	A member of the family Mimarachnidae.
^[224]	Sp. nov	In press	Zhang, Zheng & Zhang in Zhang et al.	Middle Triassic	Tongchuan Formation	China	A member of the family .
^[225]	Gen. et sp. nov	In press	Zhang, Chen & Zhang	Middle Triassic		China	A member of belonging to the family . Genus includes new species S. qishuiheensis.
^[226]	Gen. et sp. nov	Valid	Luo, Wang & Jarzembowski	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A planthopper of uncertain phylogenetic placement. The type species is S. shcherbakovi.
^[227]	Fam. nov., Gen. et sp. nov	Valid	Zhang et al.	Late Triassic		China	A planthopper in the new family . Genus includes new species S. huanglongensis
^[228]	Sp. nov	In press	Fu, Azar & Huang	Middle Triassic		China	A member of Cicadomorpha belonging to the family .
^[194]	Gen. et sp. nov	Valid	Shcherbakov	Triassic (Ladinian–Carnian)	Madygen Formation	Kyrgyzstan	A member of Cicadomorpha belonging to the superfamily and the family . The type species is T. mira.
^[229]	Sp. nov	Valid	Poinar & Brown		Dominican amber	Dominican Republic	A member of Psylloidea.
^[230]	Gen. et sp. nov	Valid	Li, Chen & Jarzembowski	Early Cretaceous		United Kingdom	A member of the family Procercopidae. Genus includes new species V. tonyi.
^[231]	Sp. nov	Valid	Gębicki & Walczak in Gębicki, Walczak & Zmarzły	Eocene	Baltic amber	Europe (Baltic Sea region)	A leafhopper belonging to the subfamily Mileewinae.

Hemipteran research[]

Grimaldi & Vea (2021) reinterpret engeli as a scale insect, and reject the interpretation of this insect as an ectoparasite.^[232]
Redescription of the holotype specimen of Paranoika placida is published by Moura-Júnior et al. (2021), who transfer this species to the genus Lethocerus, and consider this species to be the senior synonym of Lethocerus vetus.^[233]
A fulgorid specimen, representing the oldest occurrence of this family in the fossil record in Asia, is described from the Eocene Lunpola Basin (Qinghai-Tibetan Plateau, China) by Xu et al. (2021).^[234]
Revision of the fossil record of Fulgoromorpha and Cicadomorpha from the Jurassic and Cretaceous of eastern Asia is published by Chen (2021).^[235]

Hymenopterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[236]	Gen. et sp. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A bee belonging to the tribe . The type species is A. emunctorii.
^[237]	Sp. nov	In press	Wu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Vespidae.
^[238]	Fam. nov., Gen. et 3 sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family . Genus includes new species A. procerum, A. elongatum and A. grande.
^[236]	Gen. et comb. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber Bitterfeld amber	Germany Russia ( Kaliningrad Oblast)	A bee belonging to the tribe Melikertini. The type species is "" clypeatus Engel (2001).
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Crabroninae. Genus includes new species A. diabolicum.
^[239]	Sp. nov	Valid	Ngô-Muller, Garrouste & Nel in Ngô-Muller et al.	Miocene		Indonesia	An ant, a species of Anochetus.
^[237]	Sp. nov	In press	Wu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Vespidae.
^[240]	Gen. et sp. nov	Valid	Tribull, Pankowski & Colombo	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A Bethylid. The type species is A. eocena.
^[241]	Gen. et sp. nov	Valid	Rasnitsyn	Late Jurassic	Karabastau Formation	Kazakhstan	A member of Proctotrupoidea belonging to the family Peleserphidae. The type species is A. leleji.
^[242]	Sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Cheloninae.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Heterogynaidae. Genus includes new species A. setosus.
^[243]	Sp. nov	Valid	Engel in Engel et al.	Miocene (Langhian)	Zhangpu amber	China	A stingless bee, a species of Austroplebeia.
^[244]	Gen. et 2 sp. nov	Valid	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Braconidae belonging to the subfamily . Genus includes new species B. gracilens and B. grossus.
^[245]	Gen. et sp. nov	Valid	Jouault, Perrichot & Nel	Cretaceous		Myanmar	A member of the family Bethylidae belonging to the subfamily . Genus includes new species B. azevedoi.
^[246]	Gen. et sp. nov	Valid	Zheng et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A sawfly belonging to the family . Genus includes new species B. lii.
^[247]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of Apocrita belonging to the family .
^[248]	Gen. et 2 sp. nov	In press	Herbert & McKellar	Cretaceous	Burmese amber	Myanmar	A member of the family Serphitidae. Genus includes new species B. applanatus and B. myanmarensis.
^[249]	Gen. et sp. nov	Valid	Chen, van Achterberg & Hong	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae belonging to the subfamily . The type species is C. syntoma.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber Rovno amber	Russia Ukraine	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
^[251]	Gen. et 2 sp. nov	In press	Brazidec et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Diapriidae. Genus includes new species C. myanmarensis and C. ohmkuhnlei.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Astatinae. Genus includes new species C. longevus.
^[252]	Gen. et sp. nov	In press	Jouault, Perrichot & Nel	Cretaceous	Burmese amber	Myanmar	A member of Orussoidea. Genus includes new species C. vilhelmseni.
^[253]	Sp. nov	Valid	Jouault, Rasnitsyn & Nel in Jouault et al.	Early Cretaceous	Crato Formation	Brazil	A member of the family Vespidae.
^[254]	Gen. et sp. nov	In press	Santer & Álvarez-Parra in Santer et al.	Early Cretaceous (Albian)	Escucha Formation	Spain	A member of the family Spathiopterygidae. The type species is D. marveni.
^[255]	Sp. nov	Valid	Radchenko	Late Eocene	Rovno amber	Ukraine	An ant belonging to the subfamily Formicinae.
^[256]	Sp. nov	Valid	Belokobylskij in Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily Euphorinae.
^[257]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Aulacidae.
^[258]	Sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Embolemidae.
^[258]	Sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Embolemidae.
^[259]	Sp. nov	Valid	Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Pelecinidae.
^[260]	Gen. et sp. nov	Valid	Archibald, Aase & Nel	Early Eocene	Green River Formation	United States ( Wyoming)	A horntail. Genus includes new species E. lacoi.
^[261]	Sp. nov	Valid	Davidian & Kaliuzhna in Davidian, Kaliuzhna & Perkovsky	Middle Eocene	Sakhalinian amber	Russia	A member of the family Braconidae belonging to the subfamily Aphidiinae.
^[257]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Aulacidae.
Exilaulacus eximius^[257]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Aulacidae.
^[262]	Sp. nov	Valid	Radchenko & Perkovsky in Radchenko, Perkovsky & Vasilenko	Late Eocene	Rovno amber	Ukraine	An ant, a species of Formica.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family . Genus includes new species G. mandibularis.
^[250]	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae; a new genus for "Epyris" bifossatus Brues (1939).
^[263]	Gen. et sp. nov	Valid	Zheng in Zheng et al.	Cretaceous	Burmese amber	Myanmar	A member of the family . The type species is G. rasnitsyni.
^[236]	Gen. et sp. nov	Valid	Engel in Engel & Davis	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A bee belonging to the tribe Melikertini. The type species is H. uraeus.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family . Genus includes new species H. apiformis.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psenini. Genus includes new species H. verrucosus.
^[264]	Gen. et sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae. The type species is H. rasnitsyni.
^[264]	Sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae.
^[250]	Sp. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Rovno amber	Ukraine	A member of the family Bethylidae.
^[265]	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber	Myanmar	A member of Aculeata of uncertain phylogenetic placement.
Holopsenella burmitica^[265]	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber	Myanmar	A member of Aculeata of uncertain phylogenetic placement.
Holopsenella gothica^[265]	Sp. nov	In press	Lepeco & Melo	Cretaceous	Burmese amber	Myanmar	A member of Aculeata of uncertain phylogenetic placement.
^[266]	Sp. nov	Valid	Olmi, Guglielmino & Chen in Olmi et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Dryinidae.
Hybristodryinus viriosus^[267]	Sp. nov	Valid	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Dryinidae.
Hybristodryinus zaifui^[266]	Sp. nov	Valid	Chen, Olmi & Perkovsky in Olmi et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Dryinidae.
^[268]	Sp. nov	Valid	Bremer & Talamas in Bremer, van de Kamp & Talamas	Eocene	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Janzenella.
^[269]	Sp. nov	In press	Won & So	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A wasp belonging to the family .
^[270]	Gen. et 3 sp. nov	Valid	Jouault et al.	Cretaceous	Burmese amber	Myanmar	A member of Evanioidea belonging to the family Othniodellithidae. Genus includes new species K. anubis, K. basilisci and K. kirin.
^[249]	Gen. et sp. nov	Valid	Chen, van Achterberg & Hong	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is K. dolichura.
^[271]	Gen. et sp. nov	Valid	Radchenko & Proshchalykin	Late Eocene	Rovno amber	Ukraine	An ant belonging to the subfamily Myrmicinae. The type species is L. venustus.
^[272]	Sp. nov	Valid	Olmi et al.	Late Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Dryinidae.
^[250]	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Eocene (Priabonian)	Baltic amber	Russia	A member of the family Bethylidae belonging to the subfamily ; a new genus for "Calyoza" longiceps Brues (1923).
^[273]	Gen. et sp. nov	Valid	Belokobylskij & Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae. Genus includes new species M. clavicornis.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Bembicinae. Genus includes new species M. paradoxum.
^[250]	Gen. et comb. nov	Valid	Colombo & Azevedo in Colombo et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Bethylidae belonging to the subfamily ; a new genus for "Epyris" atavellus Cockerell (1920).
^[248]	Gen. et 5 sp. nov	In press	Herbert & McKellar	Cretaceous	Burmese amber	Myanmar	A member of the family Serphitidae. Genus includes new species M. annulus, M. giganteus, M. engeli, M. scutatus and M. viraneacapitis.
^[242]	Sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Meteorus.
^[242]	Sp. nov	Valid	Belokobylskij et al.	Eocene	Rovno amber	Ukraine	A member of the family Braconidae belonging to the subfamily .
^[251]	Gen. et sp. nov	In press	Brazidec et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Diapriidae. Genus includes new species M. parva.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psenini. Genus includes new species O. burmensis.
^[274]	Gen. et 2 sp. nov	Valid	Gao, Engel, Shih, Gao in Gao et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Anaxyelidae. The type species is O. elegans; genus also includes O. thanti.
^[242]	Gen. et sp. nov	Valid	Belokobylskij et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Braconidae belonging to the subfamily . The type species is P. bruesi.
^[275]	Gen. et sp. nov	Valid	Jouault & Nel	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Aulacidae. Genus includes new species P. minutus.
^[276]	Gen. et sp. nov	Valid	Jouault, Aase & Nel	Eocene (Chadronian)	Florissant Formation	United States ( Colorado)	A member of the family Xyelidae. The type species is P. nearctica.
^[277]	Sp. nov	In press	Yu et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Peleserphidae.
^[278]	Sp. nov	Valid	Varela-Hernández & Riquelme	Miocene	Mexican amber	Mexico	An ant, a species of Pheidole.
^[246]	Gen. et comb. nov	Valid	Zheng et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A sawfly belonging to the family Syspastoxyelidae; a new genus for "" pinguis.
^[279]	Sp. nov	Valid	Zheng et al.	Middle Jurassic (Callovian-Bathonian boundary)	Jiulongshan Formation	China	A member of the family Xyelidae.
^[269]	Sp. nov	In press	Won & So	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A wasp belonging to the family Angarosphecidae.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Bembicinae. Genus includes new species P. spectabilis.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psenini. Genus includes new species P. obscurus.
Prosyntexis sennlaubi^[280]	Sp. nov	Valid	Jouault & Nel	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Sepulcidae.
^[245]	Gen. et sp. nov	Valid	Jouault & Nel in Jouault, Perrichot & Nel	Early Cretaceous (Albian)	Hkamti amber	Myanmar	A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species P. myanmarensis.
^[281]	Gen. et sp. nov	Valid	Talamas et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Platygastridae. The type species is P. kleio.
^[276]	Gen. et sp. nov	Valid	Jouault, Aase & Nel	Eocene (Ypresian)	Green River Formation	United States ( Wyoming)	A member of the family Xyelidae. The type species is P. pankowskii.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Heterogynaidae. Genus includes new species P. corniculatus.
^[282]	Gen. et sp. nov	Valid	Jouault & Brazidec	Cretaceous	Burmese amber	Myanmar	A member of the family Bethylidae belonging to the subfamily . Genus includes new species R. platycephala.
^[264]	Gen. et 2 sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae. The type species is R. alexandri; genus also includes R. gracilis.
^[283]	Gen. et sp. nov	Valid	Olmi et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Dryinidae. Genus includes new species R. burmense.
^[264]	Gen. et sp. nov	Valid	Kopylov, Zhang & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Ichneumonidae. The type species is R. braconidicus.
^[284]	Sp. nov	Valid	Manukyan in Manukyan & Zhindarev	Late Eocene	Prussian Formation (Baltic amber)	Russia ( Kaliningrad Oblast)	A member of the family Ichneumonidae.
^[284]	Sp. nov	Valid	Manukyan in Manukyan & Zhindarev	Late Eocene	Prussian Formation (Baltic amber)	Russia ( Kaliningrad Oblast)	A member of the family Ichneumonidae.
^[285]	Gen. et sp. nov	Valid	Simutnik in Simutnik, Perkovsky & Vasilenko	Middle Eocene	Sakhalinian amber	Russia	A member of the family Encyrtidae. The type species is S. leleji.
^[286]	Gen. et sp. nov	Valid	Davidian in Davidian, Manukyan & Belokobylskij	Middle Eocene	Sakhalinian amber	Russia	A member of the family Braconidae belonging to the subfamily Aphidiinae. Genus includes new species S. alexrasnitsyni.
^[279]	Gen. et sp. nov	Valid	Zheng et al.	Middle Jurassic (Callovian-Bathonian boundary)	Jiulongshan Formation	China	A member of the family Xyelidae. The type species is S. daohugouensis.
^[287]	Sp. nov	Valid	Moser in Moser et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Pteromalidae.
^[287]	Sp. nov	Valid	Moser in Moser et al.	Eocene	Baltic amber	Europe (Baltic Sea region)	A member of the family Pteromalidae.
^[238]	Fam. nov., Gen. et sp. nov.	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Apoidea in the new family . Genus includes new species S. aenigmaticus.
^[273]	Gen. et sp. nov	Valid	Belokobylskij & Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Braconidae. Genus includes new species S. longiscapus.
^[243]	Sp. nov	Valid	Engel in Engel et al.	Miocene (Langhian)	Zhangpu amber	China	A stingless bee, a species of Tetragonula.
^[238]	Gen. et sp. nov	Valid	Rosa & Melo	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Heterogynaidae. Genus includes new species T. longicornis.
^[288]	Gen. et sp. nov	In press	Jouault & Nel	Paleocene	Menat Formation	France	An ant belonging to the stem group of the subfamily Myrmeciinae. Genus includes new species T. inopinata.
^[289]	Sp. nov	In press	Abels & Wedmann	Eocene	Messel pit	Germany	A member of the family Vespidae.
^[290]	Sp. nov	Valid	Manukyan in Manukyan & Smirnova	Eocene	Baltic amber	Europe (Baltic Sea region)	A species of Xeris.
Zigrasimecia goldingot^[291]	Sp. nov	In press	Zhuang et al.	Cretaceous	Burmese amber	Myanmar	An ant in the tribe Sphecomyrmini.

Hymenopteran Research[]

Guo et al. (2021) report the discovery of two queen ant specimens from Burmese amber, belonging to the species Haidomyrmex cerberus and including the first known alate queen in Haidomyrmex.^[292]
Cooccurrence of ants belonging to extant Holarctic genus Lasius and a tropical weaver ant in the same piece of amber is reported from the Bitterfeld amber (Germany) by Radchenko & Perkovsky (2021).^[293]
Original claims that both Melittosphex burmensis and Discoscapa apicula were primitive bees have been rejected by Rosa & Melo, who instead consider the former to be an aculeate wasp of uncertain affinity, and the latter to belong within the crabronid wasp subfamily Crabroninae, a lineage not directly related to bees.^[238]

Lepidopterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[294]	Gen. et 7 sp. nov	Valid	Mey, Léger & Lien	Cretaceous	Burmese amber	Myanmar	A moth belonging to the family Lophocoronidae. The type species is A. skalskii; genus also includes A. muelleri, A. bowangi, A. wichardi, A. kuranishii, A. sattleri and A. spinifera.
^[294]	Gen. et sp. nov	Valid	Mey, Léger & Lien	Cretaceous	Burmese amber	Myanmar	A moth belonging to the family Agathiphagidae. The type species is A. perdita.
^[295]	Sp. nov	Valid	Fischer	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Phyllocnistis
^[295]	Sp. nov	Valid	Fischer	Eocene to Oligocene	Baltic amber	Russia ( Kaliningrad Oblast)	A species of Phyllonorycter

Lepidopteran research[]

The first known Cretaceous caterpillar armoured with spines is described from the Burmese amber by Haug & Haug (2021).^[296]
Fossil leaf mine produced by a lyonetiid-like leaf-mining moth, representing the oldest record of such moths reported to date, is described from the Upper Cretaceous (Campanian) Kaiparowits Formation (Utah, United States) by Maccracken et al. (2021), who name a new ichnotaxon spiralae.^[297]

Mecopterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[298]	Sp. nov		Zhang, Shih & Ren in Zhang et al.	Middle Jurassic	Jiulongshan Formation	China	A scorpionfly belonging to the family Holcorpidae.
^[299]	Gen. et sp. nov	Valid	Poschmann & Nel	Permian		Germany	A scorpionfly belonging to the family . Genus includes new species G. virgoferroa.
^[300]	Sp. nov	In press	Lian, Cai & Huang	Late Jurassic	Tiaojishan Formation	China	A scorpionfly belonging to the family Orthophlebiidae.
^[301]	Sp. nov	Valid	Lian, Cai & Huang	Middle Triassic	Tongchuan Formation	China	A member of the family Mesopsychidae.
Mesopsyche liaoi^[301]	Sp. nov	Valid	Lian, Cai & Huang	Late Triassic	Huangshanjie Formation	China	A member of the family Mesopsychidae.

Neuropterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[302]	Gen. et sp. nov	Valid	Badano, Haug & Cerretti in Badano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	An early representative of the clade Myrmeleontiformia. The type species is A. caudatus.
^[303]	Sp. nov	In press	Jouault	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Rhachiberothidae belonging to the subfamily .
^[304]	Gen. et sp. nov	Valid	Cao, Wang & Liu	Cretaceous	Burmese amber	Myanmar	A member of the family Mesochrysopidae. Genus includes new species M. bilobata.
^[305]	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Babinskaiidae. The type species is C. xiai.
^[306]	Gen. et sp. nov	Valid	Xu, Zheng, Shih & Wang in Xu et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Ithonidae. Genus includes new species F. reticulatus.
^[307]	Sp. nov	Valid	Nakamine	Early Cretaceous (Aptian)	Crato Formation	Brazil	A member of the family Mesochrysopidae.
^[306]	Sp. nov	Valid	Xu, Fang, Shih & Wang in Xu et al.	Middle Jurassic	Jiulongshan Formation	China	A member of the family Ithonidae.
^[308]	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Quilchena site	Canada British Columbia	A Osmylid lance lacewing
^[308]	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Klondike Mountain Formation	United States Washington	A Osmylid lance lacewing
^[308]	Comb. nov	Valid	(Hagen)	Priabonian	Baltic amber	Europe	A Osmylid lance lacewing Moved from "" picta (1913)
^[308]	Sp. nov	Valid	Makarkin, Archibald, & Mathewes	Ypresian	Green River Formation Parachute Creek Member	United States Colorado	A Osmylid lance lacewing
^[309]	Gen. et sp. nov	Valid	Khramov	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A beaded lacewing. Genus includes new species O. simpla.
^[310]	Gen. et sp. nov	In press	Nakamine et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Rhachiberothidae. Genus includes new species P. chimaera.
^[311]	Gen. et sp. nov	Valid	Jouault & Nel	Early Cretaceous	Crato Formation	Brazil	A member of the family Babinskaiidae. Genus includes new species P. sennlaubi.
^[312]	Gen. et comb. nov	Valid	Makarkin, Ansorge & Khramov	Early Jurassic (Toarcian)		Germany	A member of the family Ithonidae sensu lato belonging to the subfamily . The type species is "" aequabilis Bode (1953).
^[313]	Gen. et sp. nov	In press	Huang & Nel in Huang et al.	Paleocene		China	A member of the family Chrysopidae. Genus includes new species S. zhangi
^[305]	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Babinskaiidae. The type species is S. punctata.
^[305]	Gen. et sp. nov	Valid	Lu, Wang & Liu	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Babinskaiidae. The type species is X. lepidotricha.

Neuropteran research[]

Review of the known record of spoon-winged lacewing larvae is published Haug, Haug & Haug (2021), who report two specimens from the Cretaceous amber from Myanmar which might represent the first known larvae of members of the subfamily Nemopterinae in the fossil record.^[314]
New lacewing larvae with elongated mouthparts are described from the Cretaceous amber from Myanmar by Haug et al. (2021), who also review the morphological diversity of fossil lacewing larvae.^[315]

Odonatans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[316]	Gen. et sp. nov	Valid	Béthoux, Deregnaucourt & Norrad in Béthoux et al.	Carboniferous (Moscovian)	Sunbury Creek Formation	Canada ( New Brunswick)	A member of of uncertain phylogenetic placement. The type species is B. norradi.
^[317]	Sp. nov	In press	Xia et al.	Late Eocene	Nima Basin	China	A species of Chalcolestes.
^[318]	Gen. et sp. nov	In press	Huang et al.	Early Cretaceous (Aptian)		China	A damselfly belonging to the family Synlestidae. Genus includes new species C. cretacicus.
^[319]	Gen. et sp. nov	Valid	Nel	Late Cretaceous (Maastrichtian)	Fox Hills Formation	United States ( South Dakota)	A dragonfly belonging to the family . Genus includes new species D. spedeni.
^[320]	Sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A member of the family .
^[320]	Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A collective group name for any fossil species referable to the whose orthotaxon generic affinity is unclear. Includes new species D. delinei.
^[321]	Sp. nov	Valid	Felker & Vasilenko	Permian		Russia	A damselfly belonging to the family .
^[322]	Gen. et sp. nov	Valid	Zheng et al.	Miocene		China	A dragonfly belonging to the group Aeshnoidea, possibly a member of the family Telephlebiidae. Genus includes new species J. taoae.
^[323]	Sp. nov	Valid	Felker	Early Triassic		Russia	A member of Protozygoptera belonging to the family .
^[324]	Sp. nov	Valid	Felker	Late Permian		Russia
^[325]	Gen. et sp. nov	Valid	Deregnaucourt et al.	Triassic	Molteno Formation	South Africa	A member of the stem group of Odonata, belonging to the group within the . Genus includes new species M. lindae.
^[320]	Gen. et 8 sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A member of the family Dysagrionidae. The type species is O. threadgillae; genus also includes new species O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum and O. worleyae.
^[320]	Gen. et 3 sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Tranquille Formation	Canada ( British Columbia)	A member of the family Dysagrionidae. The type species is O. macabeensis; genus also includes new species O. fraseri and O. jeppesenorum.
^[326]	Gen. et sp. nov	In press	So & Won	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A damselfly in the family Megapodagrionidae. Genus includes new species P. ryonsangae.
^[323]	Sp. nov	Valid	Felker	Early Triassic		Russia	A member of Protozygoptera belonging to the family Kennedyidae.
^[327]	Fam. et gen. et 2 sp. nov	In press	Zheng, Jarzembowski & Nel in Zheng et al.	Cretaceous	Burmese amber	Myanmar	A stem hemiphlebioid damselfly belonging to the new family . Genus includes new species P. zhangi and P. meiyingae.
^[328]	Gen. et sp. nov	In press	Martynov, Vasilenko & Perkovsky	Late Eocene	Rovno amber	Ukraine	A damselfly belonging to the group Coenagrionoidea. Genus includes new species P. electra.
^[329]	Gen. et sp. nov	Valid	Archibald & Cannings	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A euphaeid damselfly. Type species is R. weatbrooki.
^[320]	Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States ( Washington)	A dysagrionid odonate. Type species is S. westersidei.
Stenophlebia ryonsangensis^[330]	Sp. nov	In press	Won, So & Kim	Early Cretaceous	Sinuiju Formation	North Korea	A stenophlebiid odonate.
^[320]	Fam. Nov., Gen. et sp. nov	Valid	Archibald & Cannings in Archibald et al.	Eocene (Ypresian)	Klondike Mountain Formation	United States	A type genus of , a new family of uncertain phylogenetic placement. The type species is W. millerae.

Odonate research[]

Archibald et al. (2021) establish a new, extinct suborder of Odonata named , close to but distinct from damselflies, composed of the families Dysagrionidae, and possibly Whetwhetaksidae;^[320] their study is subsequently criticized by Nel & Zheng (2021), who consider evidence presented by Archibald et al. to be insufficient to support a new clade, and reject the suborder Cephalozygoptera as unfounded.^[331]^[332]

Orthopterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[333]	Gen. et sp. nov	In press	Du, Xu & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Tridactylidae. Genus includes new species A. cheni.
Archaboilus polyneurus^[334]	Sp. nov	Valid	Gu, Yue & Ren in Gu et al.	Middle Jurassic (Bathonian–Callovian)	Jiulongshan Formation	China	An ensiferan belonging to the group Hagloidea and the family Haglidae.
^[335]	Gen. et sp. nov	In press	Uchida	Cretaceous	Burmese amber	Myanmar	A member of the family Elcanidae. Genus includes new species C. spinocrus.
^[333]	Sp. nov	In press	Du, Xu & Zhang	Cretaceous	Burmese amber	Myanmar	A member of the family Tridactylidae.
^[336]	Sp. nov	In press	Nel & Jouault	Early Cretaceous	Crato Formation	Brazil	A member of the family Elcanidae.
^[335]	Gen. et sp. nov	In press	Uchida	Cretaceous	Burmese amber	Myanmar	A member of the family Elcanidae. Genus includes new species H. gracile.
^[337]	Sp. nov	In press	Kim et al.	Early Cretaceous (Albian)	Jinju Formation	South Korea	An orthopteran belonging to the family Elcanidae.
^[338]	Gen. et sp. nov	Valid	Gorochov	Late Permian		Russia ( Kemerovo Oblast)	A member of Hagloidea. Genus includes new species P. tomica.
^[339]	Gen. et sp. nov	In press	Xu et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Tridactylidae. Genus includes new species P. wangi.
^[336]	Sp. nov	In press	Nel & Jouault	Early Cretaceous	Crato Formation	Brazil	A member of the family Locustopsidae.

Orthopteran research[]

Revision of the fossil record of the family Trigonidiidae is published by Desutter-Grandcolas et al. (2021).^[340]

Plecoptera[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[341]	Gen. et sp. nov	In press	Chen	Eocene	Baltic amber	Europe (Baltic Sea region)	A nemourid stonefly. Genus includes new species B. bulbosus.
^[342]	Gen. et sp. nov	In press	Chen & Xu	Cretaceous	Burmese amber	Myanmar	A perlid stonefly belonging to the family. Genus includes new species B. starki.
^[343]	Sp. nov	Valid	Jouault et al.	Eocene	Baltic amber	Europe (Baltic Sea regon)	A perlodid stonefly.

Psocodeans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[344]	Gen. et sp. nov	Valid	Yoshizawa in Yoshizawa & Yamamoto	Cretaceous (Albian–Cenomanian)	Burmese amber	Myanmar	A member of Homilopsocidea of uncertain phylogenetic placement. The type species is B. lienhardi.
^[345]	Sp. nov	Valid	Hakim, Azar & Huang	Cretaceous	Burmese amber	Myanmar	A member of Trogiomorpha belonging to the family Cormopsocidae.
Cormopsocus neli^[346]	Sp. nov	Valid	Hakim, Azar & Huang in Hakim et al.	Cretaceous	Burmese amber	Myanmar	A member of Trogiomorpha belonging to the family Cormopsocidae.
^[347]	Gen. et sp. nov	Valid	Hakim, Huang & Azar	Late Cretaceous		Russia	A member of Trogiomorpha belonging to the family Empheriidae. Genus includes new species E. rasnitsyni.
^[347]	Sp. nov	Valid	Hakim, Huang & Azar	Late Cretaceous		Russia	A member of Trogiomorpha belonging to the family Trogiidae.
^[348]	Gen. et sp. nov	Valid	Cumming & Le Tirant	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Trogiomorpha belonging to the family Archaeatropidae. The type species is H. morganae.
^[349]	Gen. et 2 sp. nov	In press	Wang, Li & Yao in Wang et al.	Cretaceous	Burmese amber	Myanmar	A member of the family Cormopsocidae. Genus includes new species L. edentatus and L. pedhyalinus.
^[350]	Sp. nov	In press	Hakim, Huang & Azar	Early Cretaceous (Barremian)	Lebanese amber	Lebanon	A member of Psocodea belonging to the family Prionoglarididae.
Palaeosiamoglaris hkamtiensis^[351]	Sp. nov	In press	Jouault et al.	Early Cretaceous	Hkamti amber	Myanmar	A member of Psocodea belonging to the family Prionoglarididae.
^[351]	Sp. nov	In press	Jouault et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.
^[352]	Sp. nov	Valid	Liang & Liu	Cretaceous	Burmese amber	Myanmar	A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.
^[353]	Gen. et sp. nov	In press	Liang & Liu	Cretaceous	Burmese amber	Myanmar	A member of the family Cormopsocidae. Genus includes new species S. jiewenae.

Psocodean research[]

Four bark lice carrying sand granules and organic material atop their back are described from the Cretaceous amber from Myanmar by Kiesmüller et al. (2021), who interpret this finding as the first fossil evidence of masking behaviour in Cretaceous representatives of Psocodea.^[354]

Raphidiopterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Archiinocellia protomaculata^[355]	comb nov	valid	(Engel	Lutetian	Green River Formation	USA Colorado	A raphidiid snakefly. Moved from Archiinocellia protomaculata Engel (2011)^[356]	Archiinocellia protomaculata
^[357]	Sp. nov	In press	Jouault, Engel & Nel	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A snakefly.
^[355]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Tranquille Formation	Canada British Columbia	A raphidiid snakefly.
^[355]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Allenby Formation	Canada British Columbia	A raphidiid snakefly.
^[355]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	Klondike Mountain Formation	United States Washington	A raphidiid snakefly.
^[355]	Sp. nov	Valid	Archibald & Makarkin	Ypresian	"Unnamed unit" Ootsa Group	Canada British Columbia	A raphidiid snakefly.
^[358]	Sp. nov	In press	Ri et al.	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A member of the family Mesoraphidiidae.
Mesoraphidia ryonsangensis^[358]	Sp. nov	In press	Ri et al.	Early Cretaceous (Barremian–Aptian)	Sinuiju Formation	North Korea	A member of the family Mesoraphidiidae.

Trichopterans[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[359]	Sp. nov	Valid	Melnitsky & Ivanov	Late Cretaceous (Santonian)	Kheta Formation (Taymyr amber)	Russia ( Krasnoyarsk Krai)	A member of the family Polycentropodidae.
^[359]	Sp. nov	Valid	Melnitsky & Ivanov	Late Cretaceous (Santonian)	Kheta Formation (Taymyr amber)	Russia ( Krasnoyarsk Krai)	A member of the family Polycentropodidae.
^[360]	Sp. nov	Valid	Wichard & Neumann	Probably early Miocene	Dominican amber	Dominican Republic	A caddisfly, a species of Cernotina.
^[361]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Late Eocene	Rovno amber	Ukraine	A member of the family Hydroptilidae.
^[362]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
^[362]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
^[362]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
^[362]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
^[362]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
^[362]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
^[363]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Eocene (Priabonian)	Rovno amber	Ukraine	A species of Holocentropus.
^[364]	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber	Myanmar	A member of the family Odontoceridae.
Palaeopsilotreta succini^[364]	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber	Myanmar	A member of the family Odontoceridae.
^[365]	Sp. nov	Valid	Melnitsky, Ivanov & Perkovsky	Eocene (Priabonian)	Rovno amber	Ukraine	A species of Plectrocnemia.
^[364]	Sp. nov	Valid	Wichard, Müller & Xu	Cretaceous	Burmese amber	Myanmar	A species of Psilotreta.
^[366]	Sp. nov	Valid	Sukatsheva & Aristov	Early Cretaceous		Russia	A member of the family .
^[362]	Sp. nov	Valid	Sukacheva & Vasilenko	Early Cretaceous		Russia	A larval case of a caddisfly.
^[367]	Sp. nov	In press	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A caddisfly, a species of Wormaldia.
^[367]	Sp. nov	In press	Wang et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A caddisfly, a species of Wormaldia.

Other insects[]

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
^[368]	Gen. et sp. nov	Valid	Aristov	Late Permian		Russia	A member of Polyneoptera belonging to the group and the family . Genus includes new species A. intermittendia.
^[369]	Gen. et sp. nov	In press	Huang & Nel in Huang et al.	Middle Permian		China	A member of Megasecoptera belonging to the family . Genus includes new species A. medianelongata.
^[370]	Fam. et gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)		France	A member of Archaeorthoptera belonging to the new family . The type species is A. schubneli.
^[371]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)		France	A member of Archaeorthoptera belonging to the group Caloneurodea. Genus includes new species A. garroustei.
^[372]	Gen. et sp. nov	In press	Quispe et al.	Carboniferous (Moscovian)		France	A member of belonging to the family . Genus includes new species A. oudardi.
^[373]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)		France	A member of Archaeorthoptera possibly belonging to the family . Genus includes new species A. jouaulti.
^[370]	Gen. et sp. nov	Valid	Nel & Roques	Carboniferous (Moscovian)		France	A member of Archaeorthoptera belonging to the family . The type species is A. gui.
^[368]	Sp. nov	Valid	Aristov	Late Permian		Russia	A member of Polyneoptera belonging to the group Eoblattida and the family .
^[374]	Sp. nov	Valid	Storozhenko & Aristov	Triassic (Ladinian-Carnian)	Madygen Formation	Kyrgyzstan	A member of the family Mesorthopteridae.
^[375]	Sp. nov	Valid	Rasnitsyn & Aristov	Permian (Kungurian)		Russia ( Krasnoyarsk Krai)	A member of the family Blattinopsidae.
^[376]	Gen. et sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of of uncertain phylogenetic placement. Genus includes new species C. magnificus.
^[377]	Gen. et sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)		Germany	A member of Archaeorthoptera of uncertain phylogenetic placement. Genus includes new species C. impar.
^[378]	Sp. nov	Valid	Chen et al.	Carboniferous (Bashkirian to Moscovian)	Yanghugou Formation	China	A member of Archaeorthoptera belonging to the family .
^[379]	Sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin	Germany	A miomopteran belonging to the family .
^[379]	Sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin	Germany	A miomopteran belonging to the family Palaeomanteidae.
^[380]	Sp. nov	In press	Logghe et al.	Carboniferous-Permian (Gzhelian-Asselian)		France	A member of Palaeodictyoptera.
^[381]	Gen. et sp. nov	Valid	Nel & Roques in Nel, Roques & Duquesne	Carboniferous (Moscovian)		France	A member of Archaeorthoptera. Genus includes new species D. gallica.
^[382]	Gen. et comb. nov	Valid	Rasnitsyn & Aristov	Early Permian		United States	A member of Polyneoptera belonging to the group Eoblattida and the family . The type species is "" strictus Carpenter (1976).
^[383]	Gen. et sp. nov	Valid	Falières, Engel & Nel	Earliest Eocene	Oise amber	France	A webspinner belonging to the family Embiidae. The type species is G. raholai.
^[384]	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian		Russia ( Komi Republic)	A member of the family Blattinopsidae.
Glaphyrophlebia iva^[385]	Sp. nov	Valid	Aristov & Rasnitsyn	Middle Permian		Russia	A member of the family Blattinopsidae.
Glaphyrophlebia komia^[384]	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation	Russia ( Komi Republic)	A member of the family Blattinopsidae.
Glaphyrophlebia vorkutensis^[384]	Sp. nov	Valid	Aristov, Rasnitsyn & Naugolnykh	Permian	Inta Formation	Russia ( Komi Republic)	A member of the family Blattinopsidae.
^[386]	Sp. nov	In press	Lai et al.	Cretaceous	Burmese amber	Myanmar	A webspinner belonging to the family Clothodidae.
^[387]	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin	Russia	A member of the (stem-trichopteran or stem-amphiesmenopteran) family .
Kamopanorpa rotunda^[387]	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin	Russia	A member of the family Microptysmatidae.
Kamopanorpa sivchikovi^[387]	Sp. nov	Valid	Bashkuev & Sukatsheva	Permian	Minusinsk Coal Basin	Russia	A member of the family Microptysmatidae.
^[376]	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of Grylloblattida belonging to the family .
^[376]	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of Grylloblattida belonging to the family Liomopteridae.
^[388]	Gen. et sp. nov	Valid	Aristov & Storozhenko	Triassic (Ladinian or Carnian)	Madygen Formation	Kyrgyzstan	A webspinner belonging to the family . The type species is M. rasnitsyni.
^[389]	Fam. et gen. et 3 sp. nov	Valid	Yang et al.	Early Cretaceous	Yixian Formation	China	A member of Polyneoptera of uncertain phylogenetic placement, belonging to the new family . The type species is M. sticta; genus also includes M. elegans and M. antennalis.
^[390]	Gen. et sp. nov	Valid	Béthoux & Anderson	Late Triassic (Carnian)	Molteno Formation	South Africa	A member of Glosselytrodea belonging to the family . Genus includes new species M. parva.
^[386]	Gen. et sp. nov	In press	Lai et al.	Cretaceous	Burmese amber	Myanmar	A webspinner belonging to the family . Genus includes new species M. curvivena.
^[391]	Gen. et sp. nov	Valid	Zessin, Brauckmann & Gröning	Carboniferous (Pennsylvanian)	Osnabrück Formation	Germany	Probably a basal member of Odonatoptera. The type species is O. seppelti.
^[392]	Gen. et sp. nov	Valid	Nel & Poschmann	Early Permian (probably Asselian–?Sakmarian)	Meisenheim Formation	Germany	A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is P. prokopi.
^[377]	Gen. et sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)		Germany	A member of Archaeorthoptera belonging to the group Caloneurodea. Genus includes new species P. brauckmannii.
^[393]	Gen. et sp. nov	Valid	Nel & Poschmann	Early Permian	Saar–Nahe Basin	Germany	A member of Archaeorthoptera. Genus includes new species P. manseri.
^[376]	Gen. et sp. et comb. nov	Valid	Cui et al.	Early Permian		Germany	A member of Grylloblattida. Genus includes new species P. clara, as well as "" germanica Prokop et al. (2021).
^[377]	Gen. et 2 sp. nov	In press	Dvořák et al.	Carboniferous (Moscovian)		France Germany	A member of Archaeorthoptera belonging to the group . Genus includes new species P. punctatum Dvořák et al. and P. avionensis Nel & Roques.^[394]
^[390]	Sp. nov	Valid	Béthoux & Anderson	Late Triassic (Carnian)	Molteno Formation	South Africa	A member of Glosselytrodea belonging to the family Polycytellidae.
^[395]	Fam. et gen. et sp. nov	Valid	Gorochov	Triassic		Kyrgyzstan	A member of Embioptera belonging to the new family . Genus includes new species R. rasnitsyni.
^[396]	Sp. nov	In press	Brauckmann, Herd & Leipner	Carboniferous (Moscovian)		Germany	A member of Palaeodictyoptera belonging to the family .
^[397]	Fam. et gen. et sp. nov	Valid	Schubnel, Roques & Nel in Schubnel et al.	Carboniferous (Moscovian)		France	A member of Titanoptera belonging to the new family . The type species is T. azari.
^[376]	Sp. nov	Valid	Cui et al.	Early Permian	Goldlauter Formation	Germany	A member of Grylloblattida belonging to the family Liomopteridae.

Research[]

A well-preserved full-body impression of an elongate winged insect strongly resembling stick insects is described from the Permian (Wordian) (France) by Logghe et al. (2021), who name a new ichnotaxon radagasti.^[398]
A dragonfly-type odonatopteran larva, preserving ancestral morphological characters maintained in mayflies but lost in modern odonatan larvae, is described from the Cretaceous Burmese amber by Haug, Müller & Haug (2021).^[399]
A study on the anatomy of fossil specimens described as consecutive series of different larval stages of palaeodictyopteran species guttata and fenestrata, evaluating their implications for the knowledge of wing development in palaeodictyopterans, is published by Rosová, Sinitshenkova & Prokop (2021).^[400]

General research[]

Revision of the fossil record of insects from the Triassic of China is published by Zhang et al. (2021).^[401]
A study on the diversity of insect damage types in fossil plants from the Cretaceous (Albian to Cenomanian) Dakota Formation (United States), evaluating their implications for the knowledge of the early evolution of angiosperm florivory and associated pollination, is published by Xiao et al. (2021).^[402]

References[]

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^ Qvarnström, M.; Fikáček, M.; Wernström, J. V.; Huld, S.; Beutel, R. G.; Arriaga-Varela, E.; Ahlberg, P. E.; Niedźwiedzki, G. (June 30, 2021). "Exceptionally preserved beetles in a Triassic coprolite of putative dinosauriform origin". Current Biology. 31 (15): 3374–3381.e5. doi:10.1016/j.cub.2021.05.015. PMID 34197727. S2CID 235691750.
^ Li, Y.-D.; Tihelka, E.; Dahan, L.; Huang, D.-Y.; Cai, C.-Y. (2021). "On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)". Zootaxa. 5082 (3): 223–244. doi:10.11646/zootaxa.5082.3.2. S2CID 245319655.
^ ^a ^b Háva, J. (2021). "Two new genera and species of Dermestidae (Coleoptera) from Cretaceous Burmese amber". Euroasian Entomological Journal. 20 (6): 343–345. doi:10.15298/euroasentj.20.6.08 (inactive 2022-01-18).{{cite journal}}: CS1 maint: DOI inactive as of January 2022 (link)
^ Tihelka, E.; Li, Y.-D.; Huang, D.-Y.; Cai, C.-Y. (2021). "Are wounded-tree beetles living fossils? A new nosodendrid genus from Burmese amber with bilobed tarsi (Coleoptera: Nosodendridae)". Palaeoentomology. 4 (1): 50–56. doi:10.11646/palaeoentomology.4.1.10. S2CID 233894693.
^ ^a ^b Alekseev, V. I.; Bukejs, A. "Two fossil species of Petaliini White (Coleoptera: Ptinidae: Dorcatominae) from Eocene Baltic amber". Baltic Journal of Coleopterology. 21 (1): 11–18.
^ Perkovsky, E.E.; Háva, J.; Zaitsev, A.A. (2021). "The first finding of a skin beetle (Coleoptera, Dermestidae) from Sakhalinian amber". Paleontological Journal. 55 (2): 184–192. doi:10.1134/S0031030121020118. S2CID 233746555.
^ Bukejs, A.; Alekseev, V. I.; Háva, J. (2021). "A new species of Xyletinus Latreille, 1809 (Coleoptera: Ptinidae: Xyletininae) from Eocene Baltic amber, with a key to known fossil species". Caucasian Entomological Bulletin. 17 (1): 179–184. doi:10.23885/181433262021171-179184.
^ Kolibáč, J.; Alekseev, V. I.; Kairišs, K.; Bukejs, A. (2021). "Systematic placement and new data on the checkered beetles Aberrokorynetes Winkler and Visokorynetes Winkler (Coleoptera: Cleridae) from Eocene Baltic amber obtained from X-ray tomography". Historical Biology. in press: 1–9. doi:10.1080/08912963.2021.1994561. S2CID 240056175.
^ Tshernyshev, S. E. (2021). "A new genus and species of soft-winged flower beetles (Coleoptera, Malachiidae) in Baltic amber". Paleontological Journal. 55 (1): 90–95. doi:10.1134/S0031030121010056. ISSN 0031-0301. S2CID 232070647.
^ Legalov, A. A.; Poinar, G. (2021). "A new species of the genus Archaeosciaphilus Legalov, 2012 (Coleoptera: Сurculionidae) with a list of Entiminae weevils described from Baltic amber". Historical Biology. in press: 1–6. doi:10.1080/08912963.2021.2007530.
^ Legalov, A. A. (2021). "First record of the family Chilecarinae (Coleoptera; Ithyceridae) from the Upper Cretaceous Arkagala Formation, Magadan Region, Russian Far East". Cretaceous Research. 128: Article 105001. doi:10.1016/j.cretres.2021.105001.
^ Nabozhenko, M. V.; Bukejs, A. (2021). "The first fossilized comb-clawed beetle of the genus Asiomira Dubrovina, 1973 (Coleoptera: Tenebrionidae: Alleculinae) from Baltic Amber and notes on the distribution of extant species of the genus". Zootaxa. 5082 (2): 177–184. doi:10.11646/zootaxa.5082.2.7. S2CID 245227981.
^ Legalov, A. A. (2021). "First record of orsodacnid leaf beetle (Coleoptera: Orsodacnidae) from the Lower Cretaceous of Brazil". Historical Biology. in press: 1–5. doi:10.1080/08912963.2021.1929204. S2CID 236395989.
^ ^a ^b Bukejs, A.; Legalov, A. A. (2021). "Two new species of the family Rhynchitidae (Coleoptera: Curculionoidea) from Eocene Baltic amber, with key to species and assumed trophic relationships". Fossil Record. 24 (1): 117–127. doi:10.5194/fr-24-117-2021.
^ Szawaryn, K. (2021). "The first fossil Microweiseini (Coleoptera: Coccinellidae) from the Eocene of Europe and its significance for the reconstruction of the evolution of ladybird beetles". Zoological Journal of the Linnean Society. 193 (4): 1294–1309. doi:10.1093/zoolinnean/zlaa180.
^ Poinar, G.; Legalov, A. A. (2021). "First record of the tribe Lymantini (Coleoptera: Curculionidae) from Dominican amber". Historical Biology. in press: 1–5. doi:10.1080/08912963.2021.1893717. S2CID 242522689.
^ Alekseev, V. I.; Bukejs, A. "Cathartosilvanus perkovskyi sp. nov. - first record of Silvanidae (Coleoptera: Cucujoidea) from Eocene Rovno amber". Baltic Journal of Coleopterology. 21 (1): 19–24.
^ Legalov, A. A.; Nazarenko, V. Yu.; Vasilenko, D. V.; Perkovsky, E. E. (2021). "Ceutorhynchus Germar (Coleoptera, Curculionidae) as proxy for Eocene core Brassicaceae: first record of the genus from Rovno amber". Journal of Paleontology: 1–8. doi:10.1017/jpa.2021.82. S2CID 240546489.
^ Li, Y.-D.; Huang, D.-Y.; Cai, C.-Y. (2021). "An aberrant colydiine-like tenebrionoid beetle from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea: Zopheridae)". Palaeoentomology. 4 (6): 614–619. doi:10.11646/palaeoentomology.4.6.10. S2CID 245201548.
^ Zhao, Y.; Liu, Z.; Yu, Y.; Shih, C.; Ślipiński, A.; Ren, D. (2021). "First species of Prionoceridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Cleroidea)". Cretaceous Research. in press: Article 105118. doi:10.1016/j.cretres.2021.105118. S2CID 245058205.
^ ^a ^b Solórzano-Kraemer, M. M.; Hammel, J. U.; Kunz, R.; Xu, C.-P.; Cognato, A. I. (2021). "Miocene pinhole borer ambrosia beetles: new species of Diapus (Coleoptera: Curculionidae: Platypodinae)". Palaeoworld. in press. doi:10.1016/j.palwor.2021.10.001. S2CID 240104595.
^ Legalov, A. A.; Nazarenko, V. Y.; Perkovsky, E. E. (2021). "A new species of the genus Dorytomus Germar, 1817 (Coleoptera, Curculionidae) from Rovno amber". Zootaxa. 5006 (1): 95–100. doi:10.11646/zootaxa.5006.1.12. ISSN 1175-5326. PMID 34810584. S2CID 242796435.
^ ^a ^b ^c ^d ^e ^f Alekseev, V. I.; Bukejs, A. (2021). "Fossil Melandryidae (Coleoptera: Tenebrionoidea) from Eocene Baltic amber of the Sambian peninsula: new genus, six new species, new records and key to described taxa". Zootaxa. 4965 (1): 142–166. doi:10.11646/zootaxa.4965.1.7. PMID 33903499. S2CID 233408753.
^ Vitali, F. "Eurocrossotus alekseevi gen. n., sp. n. (Coleoptera: Cerambycidae) from Baltic amber, the first fossil member of the tribe Crossotini Thomson, 1864". Baltic Journal of Coleopterology. 21 (2): 221–228.
^ Legalov, A. A.; Nazarenko, V. Y.; Perkovsky, E. E. (2021). "A new species of the genus Glaesotropis Gratshev and Zherikhin, 1995 (Coleoptera, Anthribidae) from Rovno amber". Fossil Record. 24 (1): 1–7. doi:10.5194/fr-24-1-2021.
^ Telnov, D.; Perkovsky, E. E.; Vasilenko, D. V.; Yamamoto, S. (2021). "The first fossil Coleoptera record from the Volyn Region, Ukraine, with description of a new Glesoconomorphus (Coleoptera, Mycteridae) in syninclusion with Winterschmidtiidae (Acari) and a key to species". ZooKeys (1068): 189–201. doi:10.3897/zookeys.1068.75391. PMC 8592962. PMID 34819765.
^ Yu, Y.-L.; Li, Y.-D.; Kolibáč, J.; Ślipiński, A.; Ren, D.; Pang, H.; Li, Z.-Q. (2021). "A New Genus and Species of Lophocateridae from Mid-Cretaceous Amber of Myanmar (Coleoptera)". Insects. 12 (12): Article 1052. doi:10.3390/insects12121052. PMC 8709423. PMID 34940140.
^ Kupryjanowicz, J.; Lyubarsky, G. Yu.; Perkovsky, E. E. (2021). "Heterhelus buzina sp.n. (Coleoptera: Kateretidae) from Rovno amber: the first proxy for Sambucus in the Eocene of Eastern Europe". Invertebrate Zoology. 18 (1): 16–24. doi:10.15298/invertzool.18.1.02. S2CID 234840368.
^ Nabozhenko, M. V.; Bukejs, A. (2021). "A new species and a key to Isomira Mulsant, 1856 (Coleoptera: Tenebrionidae: Alleculinae) from Eocene Baltic amber". Caucasian Entomological Bulletin. 17 (1): 51–56. doi:10.23885/181433262021171-5156.
^ Legalov, A. A.; Kupryjanowicz, J.; Perkovsky, E. E. (2021). "A new genus of the tribe Cossonini (Coleoptera: Curculionidae) in Baltic amber (Poland)". Paleontological Journal. 55 (4): 405–409. doi:10.1134/S0031030121040109. S2CID 237378812.
^ Tihelka, E.; Yamamoto, S.; Huang, D.; Cai, C. (2021). "Enigmatic tenebrionoid beetles from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea)". Cretaceous Research. 123: Article 104779. doi:10.1016/j.cretres.2021.104779. S2CID 233908153.
^ Kirejtshuk, A. G.; Reid, C. A. M. (2021). "Lobanoviella andreyi, new genus and species of Lobanoviellini trib. nov. in the subfamily Palophaginae (Coleoptera, Megalopodidae) from the Eocene Baltic amber, with discussion of similarity with Phloeostichidae". Historical Biology. in press: 1–7. doi:10.1080/08912963.2021.1987428. S2CID 243581689.
^ ^a ^b Batelka, J.; Prokop, J. (2021). "The earliest beetle with mouthparts specialized for feeding on nectar is a parasitoid of mid-Cretaceous Hymenoptera". BMC Ecology and Evolution. 21 (1): Article number 207. doi:10.1186/s12862-021-01930-6. PMC 8607574. PMID 34809578.
^ Nabozhenko, M. V.; McKellar, R. C.; Bukejs, A. (2021). "The first described darkling beetle of the tribe Metaclisini (Coleoptera: Tenebrionidae) from Eocene Baltic amber". Zootaxa. 4999 (3): 279–284. doi:10.11646/zootaxa.4999.3.7. PMID 34810485. S2CID 237770026.
^ Li, Y.-D.; Tihelka, E.; Leschen, R. A. B.; Yu, Y.; Ślipiński, A.; Pang, H.; Huang, D.; Kolibáč, J.; Cai, C. (2021). "An exquisitely preserved tiny bark-gnawing beetle (Coleoptera: Trogossitidae) from mid-Cretaceous Burmese amber and the phylogeny of Trogossitidae". Journal of Zoological Systematics and Evolutionary Research. 59 (8): 1939–1950. doi:10.1111/jzs.12515. S2CID 238712220.
^ Bukejs, A.; Bezděk, J.; Kairišs, K.; Alekseev, V. I. (2021). "The first extinct species of Monolepta Chevrolat (Coleoptera: Chrysomelidae: Galerucinae) from Bitterfeld amber, described using X-ray micro-computed tomography". Zootaxa. 4995 (2): 345–356. doi:10.11646/zootaxa.4995.2.7. PMID 34810567. S2CID 237740153.
^ Zhang, Y. (2021). "New replacement name for the genus Latitergum Yu, Ślipiński, Leschen, Ren & Pang, 2014 (Insecta: Coleoptera: Trogossitidae)". Zootaxa. 4980 (2): 395–396. doi:10.11646/zootaxa.4980.2.11. PMID 34186972. S2CID 235687049.
^ Szawaryn, K.; Sontag, E.; Kubisz, D. (2021). "The first described fossil Oedemeridae (Insecta: Coleoptera) from Baltic amber". Journal of Paleontology: 1–4. doi:10.1017/jpa.2021.102. S2CID 242027889.
^ Tihelka, E.; Huang, D.; Cai, C. (2021). "A new genus of giant sooty mould beetles from mid-Cretaceous Burmese amber (Coleoptera: Cyclaxyridae)". Cretaceous Research. 125: Article 104876. doi:10.1016/j.cretres.2021.104876.
^ Legalov, A. A. (2021). "First record of the subfamily Sagrinae (Coleoptera: Chrysomelidae) from the Eocene of North America". Fossil Record. 24 (1): 135–139. doi:10.5194/fr-24-135-2021.
^ Legalov, A.A. (2021). "New tribe Palaeoanoplini trib. nov. (Coleoptera: Curculionidae) in Baltic amber". Paleontological Journal. 55 (2): 179–183. doi:10.1134/S0031030121020076. ISSN 0031-0301. S2CID 233746782.
^ Poinar, G. O.; Brown, A. E.; Legalov, A. A. (2021). "First record of the subfamily Chilecarinae (Coleoptera; Ithyceridae) from mid-Cretaceous Burmese amber". Cretaceous Research. 124: Article 104793. doi:10.1016/j.cretres.2021.104793. S2CID 233806293.
^ Powell, G. S.; Cline, A. R. (2021). "The First Cillaeinae (Coleoptera: Nitidulidae) Described from Amber". Annales Zoologici. 71 (1): 21–26. doi:10.3161/00034541ANZ2021.71.1.003. S2CID 232422254.
^ Tihelka, E.; Li, L.; Fu, Y.; Su, Y.; Huang, D.; Cai, C. (2021). "Angiosperm pollinivory in a Cretaceous beetle". Nature Plants. 7 (4): 445–451. doi:10.1038/s41477-021-00893-2. PMID 33846595. S2CID 233223519.
^ Bao, T.; Wedmann, S.; Grímsson, F.; Beutel, R. G.; Seyfullah, L.; Bao, L.; Jarzembowski, E. A. (2021). "Was the kateretid beetle Pelretes really a Cretaceous angiosperm pollinator?". Nature Plants. in press. doi:10.1038/s41477-021-01044-3. PMID 34949805. S2CID 245471963.
^ Tihelka, E.; Li, L.; Fu, Y.; Su, Y.; Huang, D.; Cai, C. (2021). "Reply to: Pelretes vivificus was a pollinator of Cretaceous angiosperms". Nature Plants. in press. doi:10.1038/s41477-021-01045-2. PMID 34949806. S2CID 245445815.
^ Tihelka, E.; Huang, D.; Perrichot, V.; Cai, C. (2021). "A previously missing link in the evolution of dasytine soft‐winged flower beetles from Cretaceous Charentese amber (Coleoptera, Melyridae)" (PDF). Papers in Palaeontology. 7 (4): 1753–1764. doi:10.1002/spp2.1360. S2CID 234862770.
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^ ^a ^b ^c Oyama, N.; Yukawa, H.; Imai, T. (2021). "New cockroach assemblage from the Lower Cretaceous Kitadani Formation, Fukui, Japan". Palaeontographica Abteilung A. 321 (1–6): 37–52. doi:10.1127/pala/2021/0112. S2CID 239140974.
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^ Bouju, V.; Rosse-Guillevic, S.; Griffon, M.; Bojarski, B.; Szwedo, J.; Perrichot, V. (2021). "The genus Allodia (Diptera: Mycetophilidae) in Miocene Ethiopian amber". Fossil Record. 24 (2): 339–346. doi:10.5194/fr-24-339-2021.
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^ ^a ^b Ševčík, J.; Hippa, H.; Burdíková, N. (2021). "Just a Fragment of Undescribed Diversity: Twenty New Oriental and Palearctic Species of Sciaroidea (Diptera), including DNA Sequence Data and Two New Fossil Genera". Insects. 13 (1): Article 19. doi:10.3390/insects13010019.
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^ Liu, Y.M.; Huang, D.Y. (2021). "A new species of Crenoptychoptera Kalugina, 1985 (Diptera: Ptychopteridae) from the Middle–Late Jurassic of Jiyuan Basin, China". Palaeoentomology. 4 (1): 27–29. doi:10.11646/palaeoentomology.4.1.5. S2CID 233927262.
^ Lukashevich, E. D.; Amorim, D. de S.; Ribeiro, G. C. (2021). "A Gondwanan record of the extinct genus Cretobibio (Diptera: Bibionidae)". Palaeoentomology. 4 (5): 468–474. doi:10.11646/palaeoentomology.4.5.13. S2CID 244210389.
^ ^a ^b ^c ^d Kopeć, K.; Soszyńska-Maj, A.; Kania-Kłosok, I.; Coram, R. A.; Krzemiński, W. (2021). "Morphology of the oldest fossil subfamily of Limoniidae (Diptera, Architipulinae) in the light of exceptionally preserved Mesozoic material". Scientific Reports. 11 (1): Article number 24137. doi:10.1038/s41598-021-03350-4. PMC 8683464. PMID 34921169.
^ Carmo, D. d.D.; Lamas, C. J. E.; Ribeiro, G. C. (2021). "The oldest fossil Stiletto fly: a new genus and species from the Lower Cretaceous Crato Formation of Brazil (Diptera: Therevidae)". Cretaceous Research. 130: Article 105039. doi:10.1016/j.cretres.2021.105039. S2CID 244212150.
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^ ^a ^b ^c ^d Ribeiro, G. C.; Santos, R. R.; Santos, D. (2021). "Four new species of the genus Leptotarsus Guérin-Méneville, 1831 (Insecta: Diptera: Tipulidae) from the Lower Cretaceous Crato Formation of Brazil". Cretaceous Research. 123: Article 104776. doi:10.1016/j.cretres.2021.104776. S2CID 233902054.
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^ Pielowska-Ceranowska, A.; Azar, D.; Szwedo, J. (2021). "New fossil biting midge from the Lower Cretaceous amber of Lebanon (Diptera: Ceratopogonidae)". Cretaceous Research. 129: Article 105004. doi:10.1016/j.cretres.2021.105004. ISSN 0195-6671. S2CID 239639779.
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^ ^a ^b ^c Ševčík, J.; Krzemiński, W.; Skibińska, K. (2021). "Extant Genus in the Mesozoic: Paleoplatyura Meunier (Diptera: Keroplatidae) Found in the Cretaceous Amber of Myanmar". Insects. 13 (1): Article 24. doi:10.3390/insects13010024.
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^ Li, L.; Skibińska, K.; Krzemiński, W.; Wang, B.; Xiao, C.; Zhang, Q. (2021). "A new March fly Protopenthetria skartveiti gen. nov. et sp. nov. (Diptera, Bibionidae, Plecinae) from mid-Cretaceous Burmese amber". Cretaceous Research. 127: Article 104924. doi:10.1016/j.cretres.2021.104924. ISSN 0195-6671.
^ ^a ^b ^c ^d Kania-Kłosok, I.; Jordan-Stasiło, W.; Kopeć, K.; Janiszewska, K.; Krzemiński, W. (2021). "The first stage of the evolution of Rhabdomastix (Diptera, Limoniidae) and the taxonomic implications of genus". The European Zoological Journal. 88 (1): 1152–1169. doi:10.1080/24750263.2021.2003880. S2CID 244765716.
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^ Swanson, D. R.; Heads, S. W.; Taylor, S. J.; Wang, Y. (2021). "A new remarkably preserved fossil assassin bug (Insecta, Heteroptera, Reduviidae) from the Eocene Green River Formation of Colorado". Papers in Palaeontology. 7 (3): 1459–1478. doi:10.1002/spp2.1349.
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^ ^a ^b Zhang, X.; Yao, Y.; Ren, D.; Pang, H.; Chen, H. (2021). "New mimarachnids (Hemiptera, Fulgoromorpha, Fulgoroidea) in mid-Cretaceous Burmese amber". ZooKeys (1057): 37–48. doi:10.3897/zookeys.1057.66434. PMC 8410752. PMID 34539197.
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^ ^a ^b Lian, X.; Cai, C.; Huang, D. (2021). "New species of Mesopsyche Tillyard, 1917 (Mecoptera: Mesopsychidae) from the Triassic of northwestern China". Zootaxa. 4995 (3): 565–572. doi:10.11646/zootaxa.4995.3.10. PMID 34810550. S2CID 237782965.
^ Badano, D.; Fratini, M.; Maugeri, L.; Palermo, F.; Pieroni, N.; Cedola, A.; Haug, J. T.; Weiterschan, T.; Velten, J.; Mei, M.; Di Giulio, A.; Cerretti, P. (2021). "X-ray microtomography and phylogenomics provide insights into the morphology and evolution of an enigmatic Mesozoic insect larva". Systematic Entomology. 46 (3): 672–684. doi:10.1111/syen.12482. S2CID 235363035.
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^ ^a ^b ^c Lu, X.; Wang, B.; Liu, X. (2021). "New Cretaceous antlion-like lacewings promote a phylogenetic reappraisal of the extinct myrmeleontoid family Babinskaiidae". Scientific Reports. 11 (1): Article number 16431. Bibcode:2021NatSR..1116431L. doi:10.1038/s41598-021-95946-z. PMC 8361207. PMID 34385561.
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^ ^a ^b ^c ^d Makarkin, V.N.; Archibald, S.B.; Mathewes, R.W. (2021). "New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks". Zootaxa. 4980 (1): 142–156. doi:10.11646/zootaxa.4980.1.9. PMID 34186986. S2CID 235685548.
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^ Huang, D.; Lian, X.; Jouault, C.; Gao, J.; Nel, A. (2021). "First green lacewing (Insecta: Neuroptera: Chrysopidae) from the Palaeocene Sanshui basin of Guangdong, South China" (PDF). Historical Biology. in press: 1–9. doi:10.1080/08912963.2021.1943381. S2CID 237756011.
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[Afrophaga-115] ^ ^a ^b ^c ^d ^e ^f ^g Vršanský, P.; Vršanská, L.; Vasilenko, D. V.; Puškelová, L.; Biroň, A. (2021). "An isolated Cretaceous analogue of Madagascar on the Adria–Turkey microcontinent indicated by fossils in Brezina, Algeria". Palaeontographica Abteilung A. 321 (1–6): 19–35. doi:10.1127/pala/2021/0107. S2CID 240038689.

[Nadveruzenie-116] Vršanský, P.; Sendi, H.; Hinkelman, J.; Hain, M. (2021). "Alienopterix Mlynský et al., 2018 complex in North Myanmar amber supports Umenocoleoidea/ae status". Biologia. 76 (8): 2207–2224. doi:10.1007/s11756-021-00689-x. S2CID 233784192.

[Arceotermes-117] Jiang, R.X.; Zhang, H.R.; Eldredge, K.T.; Song, X.B.; Li, Y.D.; Tihelka, E.; Huang, D.Y.; Wang, S.; Engel, M.S.; Cai, C.Y. (2021). "Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)". Palaeoentomology. 4 (4): 374–389. doi:10.11646/palaeoentomology.4.4.13. S2CID 239695173.

[Caligoptera-118] Sendi, H. (2021). "Highly specialised basal ectobiid cockroaches (Blattaria: Blattoidea) were rare in Burmese amber". Palaeontographica Abteilung A. 321 (1–6): 109–125. doi:10.1127/pala/2021/0106. S2CID 243471520.

[119] Liang, J.; Wang, Y.; Shih, C.; Ren, D. (2021). "Chuanblatta gen. nov. sexually dimorphic cockroaches of Raphidiomimidae (Blattaria) from the Jiulongshan Formation in China". Palaeontographica Abteilung A. 321 (1–6): 3–17. doi:10.1127/pala/2021/0113. ISSN 0375-0442. S2CID 240180893.

[120] Hinkelman, J. (2021). "Cuniculoblatta brevialata gen. et sp. n., the second case of brachyptery from Cretaceous North Myanmar amber". Palaeontographica Abteilung A. 321 (1–6): 97–107. doi:10.1127/pala/2021/0104. S2CID 240183883.

[121] Chen, X.-Y.; Zhang, H.-C.; Shi, X. (2021). "A new species and genus of Alienopteridae (Blattodea) from mid-Cretaceous amber of northern Myanmar". Zootaxa. 4941 (4): 580–586. doi:10.11646/zootaxa.4941.4.7. PMID 33756925. S2CID 232339877.

[Fractalia-122] Vršanský, P.; Hinkelman, J.; Koubová, I.; Sendi, H.; Kúdelová, T.; Kúdela, M.; Barclay, M. (2021). "A single common ancestor for praying mantids, termites, cave roaches and umenocoleoids" (PDF). AMBA Projekty. 11 (1): 1–16.

[123] Chen, G.; Xiao, L.; Liang, J.; Shih, C.; Ren, D. (2021). "A new cockroach (Blattodea, Corydiidae) with pectinate antennae from mid-Cretaceous Burmese amber". ZooKeys (1060): 155–169. doi:10.3897/zookeys.1060.67216. PMC 8486729. PMID 34690506.

[124] Bouju, V.; Jouault, C.; Perrichot, V. (2021). "The termite genus Glyptotermes (Isoptera: Kalotermitidae) in Miocene amber from Ethiopia". Journal of Paleontology: 1–7. doi:10.1017/jpa.2021.106. S2CID 244426915.

[125] Schneider, J. W.; Scholze, F.; Germann, S.; Lucas, S. G. (2021). "The Late Pennsylvanian nearshore insect fauna of the Kinney Brick Quarry invertebrate and vertebrate fossil Lagerstätte, New Mexico". New Mexico Museum of Natural History and Science Bulletin. 84: 255–285.

[126] Demers‐Potvin, A. V.; Larsson, H. C. E.; Cournoyer, M.; Béthoux, O. (2021). "Wing morphology of a new Cretaceous praying mantis solves the phylogenetic jigsaw of early‐diverging extant lineages". Systematic Entomology. 46 (1): 205–223. doi:10.1111/syen.12457. S2CID 231391397.

[127] Vršanský, P.; Poschmann, M. J.; Vidlička, L. (2021). "Oligocene pseudophyllodromiin cockroach from the Enspel Fossillagerstätte in Germany". Palaeontographica Abteilung A. 321 (1–6): 149–167. doi:10.1127/pala/2021/0110. ISSN 0375-0442. S2CID 240039709.

[128] Luo, C.; Beutel, R. G.; Xu, C.; Jarzembowski, E. A. (2021). "†Laticephalana liuyani gen. et sp. nov., a new bizarre roachoid of †Umenocoleidae (Insecta, Dictyoptera) from mid-Cretaceous Kachin amber". Proceedings of the Geologists' Association. 132 (4): 469–478. doi:10.1016/j.pgeola.2021.04.004. S2CID 236329819.

[129] Perkovsky, E. E.; Nel, A. (2021). "A new Rovno amber termite genus (Isoptera, Rhinotermitidae) from Styr river basin". Palaeontologia Electronica. 24 (1): Article number 24.1.a05. doi:10.26879/1127.

[130] Bezerra, F. I.; DeSouza, O.; Ribeiro, G. C.; Mendes, M. (2021). "A new primitive termite (Isoptera) from the Crato Formation, Araripe Basin, Early Cretaceous of South America". Journal of South American Earth Sciences. 109: Article 103260. Bibcode:2021JSAES.10903260B. doi:10.1016/j.jsames.2021.103260. S2CID 233807443.

[Milesitermes-131] Jouault, C.; Legendre, F.; Grandcolas, P.; Nel, A. (2021). "Revising dating estimates and the antiquity of eusociality in termites using the fossilized birth–death process". Systematic Entomology. 46 (3): 592–610. doi:10.1111/syen.12477. S2CID 235363027.

[Miniblattina-132] Sendi, H. (2021). "Diverse Liberiblattinidae (Insecta: Blattaria) from Lebanese and North Myanmar amber document allometric modifications near lowest size limit". Palaeontographica Abteilung A. 321 (1–6): 127–148. doi:10.1127/pala/2021/0108. S2CID 243475716.

[133] Hinkelman, J. (2021). "Mongolblatta sendii sp. n. (Mesoblattinidae) from North Myanmar amber links record to Laurasian sediments". Palaeontographica Abteilung A. 321 (1–6): 81–96. doi:10.1127/pala/2021/0105. S2CID 240162578.

[PAABlaberidae-134] Šmídová, L.; Vidlička, L.; Wedmann, S. (2021). "Appearance of the family Blaberidae (Insecta: Blattaria) during the Cretaceous and a review of fossils of this family". Palaeontographica Abteilung A. 321 (1–6): 71–79. doi:10.1127/pala/2021/0109. S2CID 243467634.

[Olenablatta-135] Šmídová, L. (2021). "New genus and species of the families Olidae and Corydiidae (Corydioidea, Blattodea) from mid-Cretaceous Kachin amber". Palaeontographica Abteilung A. 321 (1–6): 61–70. doi:10.1127/pala/2021/0117. S2CID 243471507.

[136] Song, Z.; Xu, C.; Li, J.; Jarzembowski, E. A.; Wang, B.; Xiao, C. (2021). "A new species of Pabuonqedidae (Blattaria: Mastotermitoidea) from mid-Cretaceous Kachin amber". Palaeontographica Abteilung A. 321 (1–6): 53–59. doi:10.1127/pala/2021/0111. S2CID 240023055.

[137] So, K. S.; Won, C. G.; Ri, C. J.; Jon, S. H.; Ju, I. Y. (2021). "Paekthoblatta, a New Predatory Cockroach Genus (Insecta: Blattaria: Raphidiomimidae) from the Lower Cretaceous of Paektho-Dong, Sinuiju, Democratic People's Republic of Korea". Paleontological Journal. 55 (8): 906–909. doi:10.1134/S0031030121080074. S2CID 245330342.

[138] Anisyutkin, L. N.; Perkovsky, E. E. (2021). "Periplaneta (?) perialla sp. nov., a cockroach larva of the subfamily Blattinae (Dictyoptera, Blattidae) from Rovno amber". Paleontological Journal. 55 (3): 294–300. doi:10.1134/S0031030121030047. S2CID 235966737.

[OyamaetalPAA-139] Oyama, N.; Yukawa, H.; Imai, T. (2021). "New cockroach assemblage from the Lower Cretaceous Kitadani Formation, Fukui, Japan". Palaeontographica Abteilung A. 321 (1–6): 37–52. doi:10.1127/pala/2021/0112. S2CID 239140974.

[140] So, K. S.; Won, C. G.; Ri, C. J.; Jon, S. H.; Ju, I. Y. (2021). "A New Species of Spinaeblattina Hinkelman, 2019 (Insecta, Blattaria, Mesoblattinidae) from the Lower Cretaceous of Paektho-Dong, Sinuiju, Democratic People's Republic of Korea". Paleontological Journal. 55 (8): 910–912. doi:10.1134/S0031030121080086. S2CID 245330596.

[141] Liuo, C.-H.; Beutel, R. G.; Thomson, U. R; Zheng, D.-R.; Li, J.-H.; Zhao, X.-Y.; Zhang, H.-C.; Wang, B. (2021). "Beetle or roach: systematic position of the enigmatic Umenocoleidae based on new material from Zhonggou Formation in Jiuquan, Northwest China, and a morphocladistic analysis". Palaeoworld. in press. doi:10.1016/j.palwor.2021.01.003. S2CID 234208413.

[142] Luo, C.; Beutel, R. G.; Engel, M. S.; Liang, K.; Li, L.; Li, J.; Xu, C.; Vršanský, P.; Jarzembowski, E. A.; Wang, B. (2021). "Life history and evolution of the enigmatic Cretaceous-Eocene Alienopteridae: A critical review". Earth-Science Reviews. 225: Article 103914. doi:10.1016/j.earscirev.2021.103914. S2CID 245601183.

[143] Taniguchi, R.; Nishino, H.; Watanabe, H.; Yamamoto, S.; Iba, Y. (2021). "Reconstructing the ecology of a Cretaceous cockroach: destructive and high-resolution imaging of its micro sensory organs". The Science of Nature. 108 (5): Article number 45. Bibcode:2021SciNa.108...45T. doi:10.1007/s00114-021-01755-9. ISSN 0028-1042. PMID 34581877. S2CID 238202024.

[Clusiomitidae-144] Roháček, J.; Hoffeins, C. (2021). "Clusiomitidae, a new family of Eocene fossil Acalyptratae, with revision of Acartophthalmites Hennig and Clusiomites Gen. Nov. (Diptera) †". Insects. 12 (12): Article 1123. doi:10.3390/insects12121123. PMC 8705662. PMID 34940210.

[145] Krzemiński, W.; Kopeć, K.; Soszyńska-Maj, A.; Skibińska, K. (2021). "Aleksiana rasnitsyni gen. et sp. nov. (Diptera, Limoniidae) to honour Alexandr Rasnitsyn". Palaeoentomology. 4 (5): 436–440. doi:10.11646/palaeoentomology.4.5.8. S2CID 240522901.

[Palaeoentomology46Nel-146] Nel, A. (2021). "New Cecidomyiidae from the Lowermost Eocene French amber (Diptera)". Palaeoentomology. 4 (6): 620–628. doi:10.11646/palaeoentomology.4.6.11. S2CID 245179601.

[147] Bouju, V.; Rosse-Guillevic, S.; Griffon, M.; Bojarski, B.; Szwedo, J.; Perrichot, V. (2021). "The genus Allodia (Diptera: Mycetophilidae) in Miocene Ethiopian amber". Fossil Record. 24 (2): 339–346. doi:10.5194/fr-24-339-2021.

[PalaeoworldPhoridae-148] Solórzano-Kraemer, M. M.; Bourdeau, C.; Rosse-Guillevic, S.; Hammel, J. U.; Xu, C.; Brown, B. V. (2021). "Parasitic hump-backed flies (Diptera: Phoridae) from Middle Miocene ambers". Palaeoworld. in press. doi:10.1016/j.palwor.2021.11.002. S2CID 244126413.

[149] Lamas, C. J. E.; Sampronha, S.; Ribeiro, G. C. (2021). "A new robber fly from the Lower Cretaceous (Aptian) Crato Formation of NE Brazil (Insecta: Diptera: Asilidae)". Cretaceous Research. 131: Article 105114. doi:10.1016/j.cretres.2021.105114.

[150] Lukashevich, E. D. (2021). "The oldest Diptera (Insecta) from the Upper Buntsandstein (early Middle Triassic) of Europe". Zootaxa. 5067 (1): 135–143. doi:10.11646/zootaxa.5067.1.10. PMID 34810754. S2CID 243901965.

[151] Skartveit, J.; Wedmann, S. (2021). "A revision of fossil Bibionidae (Insecta: Diptera) from the Oligocene of Germany". Zootaxa. 4909 (1): zootaxa.4909.1.1. doi:10.11646/zootaxa.4909.1.1. PMID 33756592. S2CID 232338865.

[152] Skartveit, J. (2021). "A new fossil species of the genus Bibio, with an update on bibionid flies from Baltic and Rovno amber (Diptera, Bibionidae)". Deutsche Entomologische Zeitschrift. 68 (1): 81–99. doi:10.3897/dez.68.60611.

[153] Ševčík, J.; Skartveit, J.; Krzemiński, W.; Skibińska, K. (2021). "A Peculiar New Genus of Bibionomorpha (Diptera) with Brachycera-Like Modification of Antennae from Mid-Cretaceous Amber of Myanmar". Insects. 12 (4): Article 364. doi:10.3390/insects12040364. PMC 8073775. PMID 33923888.

[Burmasymmerus-154] Ševčík, J.; Hippa, H.; Burdíková, N. (2021). "Just a Fragment of Undescribed Diversity: Twenty New Oriental and Palearctic Species of Sciaroidea (Diptera), including DNA Sequence Data and Two New Fossil Genera". Insects. 13 (1): Article 19. doi:10.3390/insects13010019.

[155] Poinar, G.; Vega, F. E. (2021). "A new genus of Apsilocephalidae (Diptera) in Mid-Cretaceous Burmese amber". Biosis: Biological Systems. 2 (1): 183–190. doi:10.37819/biosis.002.01.0091.

[156] Liu, Y.M.; Huang, D.Y. (2021). "A new species of Crenoptychoptera Kalugina, 1985 (Diptera: Ptychopteridae) from the Middle–Late Jurassic of Jiyuan Basin, China". Palaeoentomology. 4 (1): 27–29. doi:10.11646/palaeoentomology.4.1.5. S2CID 233927262.

[157] Lukashevich, E. D.; Amorim, D. de S.; Ribeiro, G. C. (2021). "A Gondwanan record of the extinct genus Cretobibio (Diptera: Bibionidae)". Palaeoentomology. 4 (5): 468–474. doi:10.11646/palaeoentomology.4.5.13. S2CID 244210389.

[SRCretolimonia-158] Kopeć, K.; Soszyńska-Maj, A.; Kania-Kłosok, I.; Coram, R. A.; Krzemiński, W. (2021). "Morphology of the oldest fossil subfamily of Limoniidae (Diptera, Architipulinae) in the light of exceptionally preserved Mesozoic material". Scientific Reports. 11 (1): Article number 24137. doi:10.1038/s41598-021-03350-4. PMC 8683464. PMID 34921169.

[159] Carmo, D. d.D.; Lamas, C. J. E.; Ribeiro, G. C. (2021). "The oldest fossil Stiletto fly: a new genus and species from the Lower Cretaceous Crato Formation of Brazil (Diptera: Therevidae)". Cretaceous Research. 130: Article 105039. doi:10.1016/j.cretres.2021.105039. S2CID 244212150.

[160] Li, L.; Zhang, Q.; Xiao, C. (2021). "A new genus and species of Bibionidae (Diptera: Bibionoidea) from mid-Cretaceous Kachin amber". Acta Palaeontologica Sinica. 60 (2): 243–252. doi:10.19800/j.cnki.aps.2021018.

[161] Giłka, W.; Harbach, R. E.; Perkovsky, E. E. (2021). "Mosquitoes (Diptera: Culicidae) in Eocene amber from the Rovno region, Ukraine". Zootaxa. 5016 (2): 257–270. doi:10.11646/zootaxa.5016.2.6. PMID 34810451. S2CID 238838914.

[HBAnglesea-162] Peñalver, E.; Arillo Aranda, A.; Szadziewski, R.; Stilwell, J. D. (2021). "Biting midges (Diptera: Ceratopogonidae) from the middle Eocene Anglesea amber (Australia) originated in a subpolar greenhouse earth". Historical Biology. in press: 1–11. doi:10.1080/08912963.2021.1924700. S2CID 236275915.

[SRCyttaromyia-163] Kania-Kłosok, I.; Nel, A.; Szwedo, J.; Jordan-Stasiło, W.; Krzemiński, W. (2021). "Phylogeny and biogeography of the enigmatic ghost lineage Cylindrotomidae (Diptera, Nematocera)". Scientific Reports. 11 (1): Article number 13916. Bibcode:2021NatSR..1113916K. doi:10.1038/s41598-021-91719-w. PMC 8260593. PMID 34230501.

[164] Krzemiński, W.; Kania-Kłosok, I.; Krzemińska, E.; Ševčík, J.; Soszyńska-Maj, A. (2021). "Fossils Shed a New Light on the Diversity and Disparity of the Family Limoniidae (Diptera, Nematocera)". Insects. 12 (3): Article 206. doi:10.3390/insects12030206. PMC 8000698. PMID 33804356.

[SRElephantomyia-165] Kania-Kłosok, I.; Krzemiński, W. (2021). "Recent discoveries of new Elephantomyia (Diptera, Limoniidae) fossils in Baltic amber". Scientific Reports. 11 (1): Article number 23647. doi:10.1038/s41598-021-03022-3. PMC 8655017. PMID 34880309.

[166] Szadziewski, R.; Sontag, E.; Pankowski, M. V. (2021). "A new biting midge of the genus Forcipomyia Meigen, 1818 from Miocene Ethiopian amber (Diptera: Ceratopogonidae)". Palaeoentomology. 4 (5): 441–444. doi:10.11646/palaeoentomology.4.5.9. ISSN 2624-2826. S2CID 244172069.

[SRHelius-167] Kania-Kłosok, I.; Krzemiński, W.; Arillo, A. (2021). "Two new long-rostrum cranefly species from the Cretaceous Iberian amber (Diptera, Limoniidae, Helius)". Scientific Reports. 11 (1): Article number 12851. Bibcode:2021NatSR..1112851K. doi:10.1038/s41598-021-91803-1. PMC 8213732. PMID 34145304.

[168] Wedmann, S.; Hörnschemeyer, T.; Engel, M. S.; Zetter, R.; Grímsson, F. (2021). "The last meal of an Eocene pollen-feeding fly". Current Biology. 31 (9): 2020–2026.e4. doi:10.1016/j.cub.2021.02.025. PMID 33705719.

[169] Badano, D.; Zhang, Q.; Fratini, M.; Maugeri, L.; Bukreeva, I.; Longo, E.; Wilde, F.; Yeates, D. K.; Cerretti, P. (2021). "Discovery of Lebambromyia in Myanmar Cretaceous Amber: Phylogenetic and Biogeographic Implications (Insecta, Diptera, Phoroidea)". Insects. 12 (4): Article 354. doi:10.3390/insects12040354. PMC 8074229. PMID 33923404.

[CRLeptotarsus-170] Ribeiro, G. C.; Santos, R. R.; Santos, D. (2021). "Four new species of the genus Leptotarsus Guérin-Méneville, 1831 (Insecta: Diptera: Tipulidae) from the Lower Cretaceous Crato Formation of Brazil". Cretaceous Research. 123: Article 104776. doi:10.1016/j.cretres.2021.104776. S2CID 233902054.

[Mesoacentron-171] Giłka, W.; Zakrzewska, M.; Lukashevich, E. D.; Vorontsov, D. D.; Soszyńska-Maj, A.; Skibińska, K.; Cranston, P. S. (2021). "Wanted, tracked down and identified: Mesozoic non-biting midges of the subfamily Chironominae (Chironomidae, Diptera)". Zoological Journal of the Linnean Society. doi:10.1093/zoolinnean/zlab020.

[172] Pielowska-Ceranowska, A.; Azar, D.; Szwedo, J. (2021). "New fossil biting midge from the Lower Cretaceous amber of Lebanon (Diptera: Ceratopogonidae)". Cretaceous Research. 129: Article 105004. doi:10.1016/j.cretres.2021.105004. ISSN 0195-6671. S2CID 239639779.

[173] Wagner, R. (2021). "Neoarisemus groehni sp. nov., a notable moth fly (Diptera, Psychodidae, Psychodinae) from Ukrainian amber". Palaeoentomology. 4 (1): 19–22. doi:10.11646/palaeoentomology.4.1.3. S2CID 233941588.

[174] Men, Q.; Podenas, S. (2021). "A new genus of Limoniidae (Diptera: Tipuloidea) from the mid-Cretaceous Burmese amber". Cretaceous Research. 126: Article 104915. doi:10.1016/j.cretres.2021.104915.

[Insects12Palaeoglaesum-175] Skibińska, K.; Albrycht, M.; Zhang, Q.; Giłka, W.; Zakrzewska, M.; Krzemiński, W. (2021). "Diversity of the Fossil Genus Palaeoglaesum Wagner (Diptera, Psychodidae) in the Upper Cretaceous Amber of Myanmar". Insects. 12 (3): Article 247. doi:10.3390/insects12030247. PMC 8000030. PMID 33809471.

[Insects131Paleoplatyura-176] Ševčík, J.; Krzemiński, W.; Skibińska, K. (2021). "Extant Genus in the Mesozoic: Paleoplatyura Meunier (Diptera: Keroplatidae) Found in the Cretaceous Amber of Myanmar". Insects. 13 (1): Article 24. doi:10.3390/insects13010024.

[177] Curler, G. R.; Skibińska, K. (2021). "Paleotelmatoscopus, a proposed new genus for some fossil moth flies (Diptera, Psychodidae, Psychodinae) in Eocene Baltic amber, with description of a new species". Zootaxa. 4927 (4): 505–524. doi:10.11646/zootaxa.4927.4.2. ISSN 1175-5326. PMID 33756688. S2CID 232339113.

[178] Li, L.; Skibińska, K.; Krzemiński, W.; Wang, B.; Xiao, C.; Zhang, Q. (2021). "A new March fly Protopenthetria skartveiti gen. nov. et sp. nov. (Diptera, Bibionidae, Plecinae) from mid-Cretaceous Burmese amber". Cretaceous Research. 127: Article 104924. doi:10.1016/j.cretres.2021.104924. ISSN 0195-6671.

[EZJRhabdomastix-179] Kania-Kłosok, I.; Jordan-Stasiło, W.; Kopeć, K.; Janiszewska, K.; Krzemiński, W. (2021). "The first stage of the evolution of Rhabdomastix (Diptera, Limoniidae) and the taxonomic implications of genus". The European Zoological Journal. 88 (1): 1152–1169. doi:10.1080/24750263.2021.2003880. S2CID 244765716.

[180] Zakrzewska, M.; Jankowska, A. (2021). "A new fossil Stempellinella from Eocene Baltic amber, with systematic notes and amendment of the generic diagnosis (Diptera: Chironomidae)". Zootaxa. 4908 (4): 505–514. doi:10.11646/zootaxa.4908.4.4. PMID 33756601. S2CID 232340439.

[181] Kania-Kłosok, I.; Krzemiński, W.; Kopeć, K.; Arillo, A. (2021). "The Oldest Evolutionary Lineage of Trichoneura Loew, 1850 (Diptera, Limoniidae) and the First Evidence of This Genus in Cretaceous Spanish Amber". Insects. 12 (5): Article 411. doi:10.3390/insects12050411. PMC 8147636. PMID 34063589.

[182] Zhang, Q.; Zhang, J. (2021). "New record of Zhangsolva Nagatomi & Yang, 1998 (Diptera, Zhangsolvidae) from mid-Cretaceous Burmese amber and its implications". In S-C. Chang; D. Zheng (eds.). Mesozoic Biological Events and Ecosystems in East Asia. Geological Society, London, Special Publications. The Geological Society of London. doi:10.1144/SP521-2021-150. S2CID 245059011.

[183] Baranov, V. A.; Engel, M. S.; Hammel, J.; Hörnig, M. K.; van de Kamp, T.; Zuber, M.; Haug, J. T. (2021). "Synchrotron-radiation computed tomography uncovers ecosystem functions of fly larvae in an Eocene forest". Palaeontologia Electronica. 24 (1): Article number 24.1.a07. doi:10.26879/1129.

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[185] Godunko, R. J.; Martynov, A. V.; Staniczek, A. H. (2021). "First fossil record of the mayfly family Vietnamellidae (Insecta, Ephemeroptera) from Burmese Amber confirms its Oriental origin and gives new insights into its evolution". ZooKeys (1036): 99–120. doi:10.3897/zookeys.1036.66435. PMC 8128840. PMID 34040482.

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[187] Sinitshenkova, N. D. (2021). "A new mayfly species Triassodotes rasnitsyni sp. nov. of the family Misthodotidae Tillyard, 1932 (Insecta; Ephemerida, Permoplectoptera) from the Triassic deposits of Kuzbass, Russia". Palaeoentomology. 4 (5): 429–432. doi:10.11646/palaeoentomology.4.5.6. S2CID 244195004.

[188] Sroka, P.; Godunko, R. J.; Sinitshenkova, N. D.; Prokop, J. (2021). "Life history, systematics and flight ability of the Early Permian stem-mayflies in the genus Misthodotes Sellards, 1909 (Insecta, Ephemerida, Permoplectoptera)". BMC Ecology and Evolution. 21 (1): Article number 97. doi:10.1186/s12862-021-01820-x. PMC 8142488. PMID 34024284.

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[191] Heiss, E.; Kairišs, K.; Bukejs, A. (2021). "The invisible becomes visible: X-ray micro-CT reconstruction of Aneurus damzeni sp. nov. (Hemiptera: Heteroptera: Aradidae) from Eocene Baltic amber". Zootaxa. 4950 (1): 189–195. doi:10.11646/zootaxa.4950.1.11. PMID 33903327. S2CID 233409823.

[192] Swanson, D. R.; Heads, S. W.; Taylor, S. J.; Wang, Y. (2021). "A new remarkably preserved fossil assassin bug (Insecta, Heteroptera, Reduviidae) from the Eocene Green River Formation of Colorado". Papers in Palaeontology. 7 (3): 1459–1478. doi:10.1002/spp2.1349.

[193] Kim, J.; Chérot, F.; Jung, S. (2021). "A new genus and species of the subfamily Mirinae (Hemiptera: Cimicomorpha: Miridae) from Eocene Baltic amber with a new generic combination for Calocoris antennatus Statz in Statz & Wagner, 1950". Zootaxa. 5057 (4): 571–576. doi:10.11646/zootaxa.5057.4.7. PMID 34811191. S2CID 239959240.

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[195] Souma, J.; Yamamoto, S.; Takahashi, Y. (2021). "A new species of the fossil lace bug genus Burmacader (Hemiptera: Heteroptera: Tingidae) from mid-Cretaceous Kachin amber from northern Myanmar". Cretaceous Research. 126: Article 104911. doi:10.1016/j.cretres.2021.104911. ISSN 0195-6671.

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[197] Liu, G.; Wang, X.; Zhuo, D.; Chen, J. (2021). "A new non-jumping plant-louse (Hemiptera, Sternorrhyncha, Malmopsyllidae) in mid-Cretaceous Kachin amber, northern Myanmar". Cretaceous Research. 124: Article 104816. doi:10.1016/j.cretres.2021.104816. S2CID 233619062.

[198] Qiao, D.; Zhang, W.; Zhang, X.; Ren, D.; Yao, Y. (2021). "New genus and species of Tettigarctidae (Hemiptera: Cicadomorpha: Cicadoidea) from the mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 126: Article 104900. doi:10.1016/j.cretres.2021.104900.

[199] Souma, J.; Yamamoto, S.; Takahashi, Y. (2021). "Discovery of the Smallest Lace Bug from Mid-Cretaceous of Northern Myanmar Supports the Hypothesis of a Miniaturization Phenomenon of Insects in Kachin Amber (Hemiptera: Heteroptera: Tingidae)". Taxonomy. 1 (2): 60–68. doi:10.3390/taxonomy1020007.

[200] Kim, J.; Lim, H.; Jung, S. (2021). "A first member of bamboo-feeding lineage plant bug (Hemiptera: Heteroptera: Miridae: Mecistoscelini) in mid-Cretaceous Burmese amber". Cretaceous Research. 121: Article 104741. doi:10.1016/j.cretres.2020.104741. S2CID 233567365.

[201] Zhang, X.; Yao, Y.; Ren, D.; Pang, H. (2021). "A new species of Cretodorus (Hemiptera: Fulgoromorpha: Fulgoroidea: Mimarachnidae) from Upper Cretaceous amber of northern Myanmar". Cretaceous Research. 128: Article 104988. doi:10.1016/j.cretres.2021.104988. S2CID 238849433.

[202] Fu, Y.; Huang, D. (2021). "A new sinoalid froghopper in mid-Cretaceous amber from northern Myanmar (Hemiptera, Cicadomorpha, Sinoalidae)". Cretaceous Research. 125: Article 104841. doi:10.1016/j.cretres.2021.104841.

[Multistria-203] Zhang, X.; Yao, Y.; Ren, D.; Pang, H.; Chen, H. (2021). "New mimarachnids (Hemiptera, Fulgoromorpha, Fulgoroidea) in mid-Cretaceous Burmese amber". ZooKeys (1057): 37–48. doi:10.3897/zookeys.1057.66434. PMC 8410752. PMID 34539197.

[204] Song, Z.-S.; Zhang, C.-L.; Xi, H.-Y.; Szwedo, J.; Bourgoin, T. (2021). "First record of adult Dorytocidae — Dorytocus jiaxiaoae Song, Szwedo & Bourgoin sp. nov. (Hemiptera: Fulgoromorpha: Fulgoroidea) from mid-Cretaceous Kachin amber". Cretaceous Research. 125: Article 104863. doi:10.1016/j.cretres.2021.104863.

[205] Chen, J.; Jiang, H.; Shang, T.; Zhang, L.; Zheng, Y.; Zhang, Q.; Zhang, H. (2021). "The second leafhopper of the subfamily Ledrinae (Hemiptera, Cicadellidae) in mid-Cretaceous Burmese amber". Cretaceous Research. 122: Article 104763. doi:10.1016/j.cretres.2021.104763. S2CID 233579718.

[206] Kim, J.; Jung, S. (2021). "A new species of extinct genus Electromyiomma Popov & Herczek (Hemiptera: Miridae: Isometopinae) from Eocene Baltic amber, with the key to Electromyiomma species". Zootaxa. 4927 (2): 289–293. doi:10.11646/zootaxa.4927.2.9. PMID 33756714. S2CID 232339534.

[207] Wedmann, S.; Kment, P.; Campos, L. A.; Hörnschemeyer, T. (2021). "Bizarre morphology in extinct Eocene bugs (Heteroptera: Pentatomidae)". Royal Society Open Science. 8 (12): Article ID 211466. Bibcode:2021RSOS....811466W. doi:10.1098/rsos.211466. PMC 8652274. PMID 34909219.

[208] Cumming, R. T.; Le Tirant, S. (2021). "Drawing the Excalibur bug from the stone: adding credibility to the double-edged sword hypothesis of coreid evolution (Hemiptera, Coreidae)". ZooKeys (1043): 117–131. doi:10.3897/zookeys.1043.67730. PMC 8217075. PMID 34163298.

[209] Monferran, M. D.; de Lima, F. J.; Kellner, A. W. A.; Sayão, J. M.; Filho, E. B. S.; Saraiva, A. Á. F.; Sabater, L. M.; Bantim, R. A. M. (2021). "An unusual preservation of a new species of Cicadomorpha (Auchenorrhyncha) from the Crato formation, Lower Cretaceous of Brazil". Historical Biology. in press: 1–9. doi:10.1080/08912963.2021.1966776. S2CID 244239311.

[210] Cifuentes-Ruiz, P.; Brailovsky, H.; Serrano-Sánchez, M. L.; Vega, F. J. (2021). "First fossil Schizopteridae (Hemiptera: Heteroptera) in Miocene amber from Chiapas, Mexico". Boletín de la Sociedad Geológica Mexicana. 73 (3): A170121. doi:10.18268/BSGM2021v73n3a170121. S2CID 245031228.

[211] Shcherbakov, D. E.; Emeljanov, A.F. (2021). "Paradoxical derbid-like planthopper (Homoptera: Fulgoroidea) from Cretaceous Burmese amber". Russian Entomological Journal. 30 (2): 135–139. doi:10.15298/rusentj.30.2.03. S2CID 237815990.

[212] Fu, Y.Z.; Huang, D.Y. (2021). "A new species of Sinoalidae from the topmost Late Jurassic Daohugou Bed (Hemiptera, Cicadomorpha)". Palaeoentomology. 4 (1): 34–38. doi:10.11646/palaeoentomology.4.1.7. S2CID 233954621.

[213] Doweld, A. B. (2021). "Karatauella, a new generic replacement name for fossil Carpenterella Becker-Migdisova, 1968 (Insecta: Hemiptera: Protopsyllidiidae incertae sedis)". Zootaxa. 5047 (1): 95–96. doi:10.11646/zootaxa.5047.1.9. PMID 34811005. S2CID 244258647.

[214] Wang, Y.; Tang, H.; Ren, D.; Yao, Y. (2021). "New Fossil Tingidae (Hemiptera: Heteroptera) from the Mid-Cretaceous of Myanmar, with Remarks on the Phylogenetic Relationships within the Family". Insects. 12 (10): Article 887. doi:10.3390/insects12100887. PMC 8540450. PMID 34680658.

[215] Heiss, E.; Golub, V. B. (2021). "A new genus and species of Tingidae from Cretaceous Burmese amber (Hemiptera, Heteroptera)". Palaeoentomology. 4 (2): 137–140. doi:10.11646/palaeoentomology.4.2.2. S2CID 235539280.

[216] Heiss, E.; Golub, V. B. (2021). "New synonymy in Burmese amber Tingidae (Hemiptera, Heteroptera)". Palaeoentomology. 4 (3): 202. doi:10.11646/palaeoentomology.4.3.2. S2CID 237896242.

[Palaeoentomology44Paleocader-217] Heiss, E.; Golub, V. B. (2021). "Three new species of the fossil Cantacaderinae genus Paleocader Froeschner, 1996 from the Baltic and Rovno amber (Hemiptera, Heteroptera, Tingidae)". Palaeoentomology. 4 (4): 360–366. doi:10.11646/palaeoentomology.4.4.11. S2CID 239623453.

[218] Golub, V. B.; Perkovsky, E. E.; Vasilenko, D. V. (2021). "A new fossil species of the genus Parasinalda Heiss & Golub (Hemiptera: Heteroptera: Tingidae) from Upper Eocene Rovno amber". Zootaxa. 5027 (2): 290–296. doi:10.11646/zootaxa.5027.2.9. ISSN 1175-5326. PMID 34811229. S2CID 239703141.

[219] Kóbor, P.; Roca-Cusachs, M. (2021). "Pseudocaulisoculus longicornis: description of a new yuripopovinid true bug from mid-Cretaceous Burmese amber (Heteroptera: Coreoidea: Yuripopovinidae)". Cretaceous Research. 125: Article 104849. doi:10.1016/j.cretres.2021.104849.

[220] Tang, H.; Wang, Y.; Ren, D.; Yao, Y. (2021). "The first record of vetanthocorids (Hemiptera: Heteroptera: Vetanthocoridae) from mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 131: Article 105090. doi:10.1016/j.cretres.2021.105090. ISSN 0195-6671. S2CID 244081314.

[221] Kment, P. (2021). "A review of the fossil taxa originally assigned to Pyrrhocoridae (Hemiptera: Heteroptera)". Zootaxa. 5081 (4): 579–586. doi:10.11646/zootaxa.5081.4.8. S2CID 245135992.

[222] Dietrich, C. H.; Dmitriev, D. A.; Perkovsky, E. E. (2021). "A new leafhopper (Hemiptera: Cicadellidae) from late Eocene Rovno amber described based on an adult and associated last-instar nymph from Perebrody (Ukraine)". Palaeoentomology. 4 (5): 445–452. doi:10.11646/palaeoentomology.4.5.10. S2CID 240525234.

[223] Luo, C.; Liu, X.; Jiang, T.; Song, Z.; Jarzembowski, E. A. (2021). "The first mimarachnid planthopper (Hemiptera: Fulgoromorpha: Mimarachnidae: Saltissus) from the Wealden (Lower Cretaceous) of southern England". Historical Biology. in press: 1–6. doi:10.1080/08912963.2021.1998035. S2CID 243487825.

[224] Zhang, Q.-Q.; Zheng, D.-R.; Teng, X.; Zhang, H.-C. (2021). "New Scytinopteridae (Hemiptera: Scytinopteroidea) from the middle Triassic Tongchuan Entomofauna of NW China". Historical Biology. in press: 1–6. doi:10.1080/08912963.2021.2010194. S2CID 244916999.

[225] Zhang, Q.-Q.; Chen, J.; Zhang, H.-C. (2021). "New Granulidae (Hemiptera: Scytinopteroidea) from the Middle Triassic Tongchuan Formation of NW China". Historical Biology. in press: 1–6. doi:10.1080/08912963.2021.1903892. S2CID 234836364.

[226] Luo, C.; Wang, B.; Jarzembowski, E. A. (2021). "A Bizarre Planthopper Nymph (Hemiptera: Fulgoroidea) from Mid-Cretaceous Kachin Amber". Insects. 12 (4): Article 318. doi:10.3390/insects12040318. PMC 8066557. PMID 33918264.

[227] Zhang, Q.; Jiang, T.; Szwedo, J.; Zhang, H. (2021). "A new family of Triassic planthoppers (Hemiptera: Fulgoromorpha: Fulgoroidea) from the Shaanxi Province of China". Alcheringa: An Australasian Journal of Palaeontology. 45 (1): 86–90. doi:10.1080/03115518.2021.1919206. S2CID 235748056.

[228] Fu, Y.; Azar, D.; Huang, D. (2021). "The first Dysmorphoptilidae from the Middle Triassic of China (Hemiptera: Cicadomorpha)". Historical Biology. 33 (12): 3506–3512. doi:10.1080/08912963.2021.1874374. S2CID 233710366.

[229] Poinar Jr., G.; Brown, A. E. (2021). "A new species of plant louse from mid-Tertiary Dominican amber (Hemiptera: Psylloidea) with adjacent plant leaves". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 302 (2): 139–146. doi:10.1127/njgpa/2021/1024. S2CID 243913929.

[230] Li, Y.; Chen, J.; Jarzembowski, E. (2021). "The first true procercopid (Hemiptera, Cercopoidea) from England". Cretaceous Research. 127: Article 104933. doi:10.1016/j.cretres.2021.104933.

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[232] Grimaldi, D. A.; Vea, I. M. (2021). "Insects with 100 million-year-old dinosaur feathers are not ectoparasites". Nature Communications. 12 (1): Article number 1469. Bibcode:2021NatCo..12.1469G. doi:10.1038/s41467-021-21751-x. PMC 7935990. PMID 33674573.

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[234] Xu, X.-T.; Deng, W.-Y.-D.; Zhou, Z.-K.; Wappler, T.; Su, T. (2021). "The first Fulgoridae (Hemiptera: Fulgoromorpha) from the Eocene of the central Qinghai–Tibetan Plateau". Fossil Record. 24 (2): 263–274. doi:10.5194/fr-24-263-2021.

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[Aethemelikertes-236] Engel, M. S.; Davis, S. R. (2021). "New genera of melikertine bees with facial modifications in Baltic amber (Hymenoptera: Apidae)". Journal of Melittology. 103 (103): 1–52. doi:10.17161/jom.i103.15655. S2CID 237834516.

[CRVespidae-237] Wu, Q.; Yang, Y.; Shih, C.; Zhao, Y.; Ren, D. (2021). "Two new species of Vespidae (Hymenoptera) in mid-Cretaceous amber from northern Myanmar". Cretaceous Research. 132: Article 105117. doi:10.1016/j.cretres.2021.105117.

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[239] Ngô-Muller, V.; Garrouste, R.; Schubnel, T.; Pouillon, J.-M.; Christophersen, V.; Christophersen, N.; Nel, A. (2021). "The first representative of the trap-jaw ant genus Anochetus Mayr, 1861 in Neogene amber from Sumatra (Hymenoptera: Formicidae)". Comptes Rendus Palevol. 20 (2): 21–27. doi:10.5852/cr-palevol2021v20a2.

[240] Tribull, C. M.; Pankowski, M. V.; Colombo, W. D. (2021). "A new genus and species of Pristocerinae (Hymenoptera, Bethylidae) from upper Eocene Baltic amber with a review of conspecific association from insect fossils". Journal of Hymenoptera Research. 85: 119–133. doi:10.3897/jhr.85.68658.

[241] Rasnitsyn, A. P. (2021). "First Jurassic representative of the extinct family Peleserphidae (Hymenoptera, Proctotrupoidea)". Journal of Hymenoptera Research. 84: 295–300. doi:10.3897/jhr.84.65493.

[Palaeocolastes-242] Belokobylskij, S. A.; Pankowski, M. G.; Pankowski, M. V.; Zaldívar-Riverón, A. (2021). "A new genus and three new species of fossil braconid wasps (Hymenoptera, Ichneumonoidea) from Eocene Baltic and Rovno ambers". Journal of Paleontology. 95 (6): 1259–1272. doi:10.1017/jpa.2021.47. S2CID 237921761.

[JM105-243] Engel, M. S.; Herhold, H.; Davis, S.; Wang, B.; Thomas, J. (2021). "Stingless bees in Miocene amber of southeastern China (Hymenoptera: Apidae)". Journal of Melittology. 105 (105): 1–83. doi:10.17161/jom.i105.15734. S2CID 239647786.

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[Burmapyris-245] Jouault, C.; Perrichot, V.; Nel, A. (2021). "New flat wasps from mid-Cretaceous Burmese amber deposits highlight the bethylid antiquity and paleobiogeography (Hymenoptera: Chrysidoidea)" (PDF). Cretaceous Research. 123: Article 104772. doi:10.1016/j.cretres.2021.104772. S2CID 234041758.

[Burmoxyela-246] Zheng, Y.; Hu, H.; Zhang, H.; Chen, J.; Rasnitsyn, A. P.; Zhuo, D. (2021). "New genus and species of syspastoxyelid sawflies (Insecta, Hymenoptera) from the mid-Cretaceous Kachin amber with a review of the family Syspastoxyelidae". Cretaceous Research. 127: Article 104940. doi:10.1016/j.cretres.2021.104940.

[247] Li, L.; Shih, C.; Yang, J.; Gao, Y.; Ren, D. (2021). "Two new wasps from mid-Cretaceous Myanmar amber (Hymenoptera: Apocrita)". Cretaceous Research. 131: Article 105115. doi:10.1016/j.cretres.2021.105115. S2CID 244885445.

[Buserphites-248] Herbert, M. C. M.; McKellar, R. C. (2021). "New genera Buserphites and Mesoserphites (Hymenoptera: Serphitidae) from mid-Cretaceous amber of Myanmar" (PDF). Cretaceous Research. 130: Article 105025. doi:10.1016/j.cretres.2021.105025. S2CID 240331883.

[Chainochora-249] Chen, H.; van Achterberg, C.; Hong, P. (2021). "Two new genera of Protobraconinae (Hymenoptera, Braconidae) from mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 126: Article 104914. doi:10.1016/j.cretres.2021.104914.

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[Christophus-251] Brazidec, M.; Perrichot, V.; Nel, A.; Jouault, C. (2021). "Two new genera of belytine wasps (Hymenoptera: Diapriidae) from Cretaceous Burmese amber". Proceedings of the Geologists' Association. 132 (5): 597–604. doi:10.1016/j.pgeola.2021.09.006. S2CID 244613026.

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[253] Jouault, C.; Rasnitsyn, A. P.; Pouillon, J.-M.; Nel, A. (2021). "The first Curiosivespa Rasnitsyn, 1975 (Hymenoptera: Vespidae) from the Lower Cretaceous Crato formation (Brazil)". Annales de la Société Entomologique de France. Nouvelle Série. 57 (6): 509–513. doi:10.1080/00379271.2021.1976672. S2CID 244229524.

[254] Santer, M.; Álvarez-Parra, S.; Nel, A.; Peñalver, E.; Delclòs, X. (2021). "New insights into the enigmatic Cretaceous family Spathiopterygidae (Hymenoptera: Diaprioidea)". Cretaceous Research. in press: Article 105128. doi:10.1016/j.cretres.2021.105128. S2CID 245556639.

[255] Radchenko, A. G. (2021). "New species of the fossil ant genus Drymomyrmex (Hymenoptera, Formicidae, Formicinae) from the late Eocene Rovno amber (Ukraine)". Palaeoentomology. 4 (6): 544–549. doi:10.11646/palaeoentomology.4.6.4. S2CID 245149637.

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[CRLietalAulacidae-257] Li, L.; Shih, P. J. M.; Zhang, X.; Peng, C.; Li, D.; Ren, D. (2021). "Geometric Morphometric Analysis of two genera confirm three new wasps from the mid-Cretaceous of Myanmar (Hymenoptera: Aulacidae)". Cretaceous Research. 130: Article 105057. doi:10.1016/j.cretres.2021.105057. S2CID 242552161.

[Palaeoentomology41Embolemus-258] Olmi, M.; Guglielmino, A.; Müller, P.; Capradossi, L.; Perkovsky, E. E. (2021). "Descriptions of the first females of Embolemus (Hymenoptera: Chrysidoidea) from Kachin (Myanmar) amber". Palaeoentomology. 4 (1): 44–49. doi:10.11646/palaeoentomology.4.1.9. S2CID 233944242.

[259] Jouault, C. (2021). "Mid-Cretaceous Burmese amber pelecinid wasps (Hymenoptera, Pelecinidae) support the hypothesis of an Asian origin of the family" (PDF). Annales de Paléontologie. 107 (1): Article 102464. doi:10.1016/j.annpal.2020.102464. S2CID 234321579.

[260] Archibald, S. B.; Aase, A.; Nel, A. (2021). "The second North American fossil horntail wood-wasp (Hymenoptera: Siricidae), from the early Eocene Green River Formation". Zootaxa. 4999 (4): 325–334. doi:10.11646/zootaxa.4999.4.2. PMID 34810481. S2CID 237762599.

[261] Davidian, E. M.; Kaliuzhna, M. O.; Perkovsky, E. E. (2021). "First aphidiine wasp from the Sakhalinian amber". Acta Palaeontologica Polonica. 66 (Supplement to 3): S59–S65. doi:10.4202/app.00843.2020. S2CID 236723591.

[262] Radchenko, A. G.; Perkovsky, E. E.; Vasilenko, D. V. (2021). "Formica species (Hymenoptera, Formicidae, Formicinae) in late Eocene Rovno amber". Journal of Hymenoptera Research. 82: 237–251. doi:10.3897/jhr.82.64599.

[263] Zheng, Y.; Kopylov, D. S.; Zhang, Q.; Zhang, H.-G.; Qin, M.; Chen, J.; Zhang, H.-C. (2021). "Discovery of a new symphytan wasp in the mid-Cretaceous Kachin amber (Hymenoptera, Syspastoxyelidae)". Palaeoentomology. 4 (6): 629–636. doi:10.11646/palaeoentomology.4.6.12. S2CID 245173388.

[Heteroichneumon-264] Kopylov, D. S.; Zhang, Q.; Zhang, H.-C. (2021). "The Darwin wasps (Hymenoptera: Ichneumonidae) of Burmese amber". Palaeoentomology. 4 (6): 592–603. doi:10.11646/palaeoentomology.4.6.8. S2CID 245157688.

[CRHolopsenella-265] Lepeco, A.; Melo, G. A. R. (2021). "The wasp genus †Holopsenella in mid-Cretaceous Burmese amber (Hymenoptera: †Holopsenellidae stat. nov.)". Cretaceous Research. 131: Article 105089. doi:10.1016/j.cretres.2021.105089. ISSN 0195-6671. S2CID 244055567.

[JHR81-266] Olmi, M.; Chen, H.-Y.; Shih, C.; Müller, P.; Capradossi, L.; Ren, D.; Perkovsky, E. E.; Guglielmino, A. (2021). "New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts". Journal of Hymenoptera Research. 81: 43–55. doi:10.3897/jhr.81.57792.

[267] Wang, Z.; Olmi, M.; Shih, C.; Ren, D.; Gao, T. (2021). "New species of Hybristodryinus from mid-Cretaceous Myanmar amber (Hymenoptera, Dryinidae)". Cretaceous Research. 125: Article 104861. doi:10.1016/j.cretres.2021.104861. ISSN 0195-6671.

[268] Bremer, J.; van de Kamp, T.; Talamas, E. J. (2021). "Janzenella theia Bremer & Talamas (Platygastroidea, Janzenellidae): a new species from Baltic amber". Journal of Hymenoptera Research. 87: 223–233. doi:10.3897/jhr.87.67256.

[WonSoCRwasps-269] Won, C. G.; So, K. S. (2021). "Two new wasps (Insecta: Hymenoptera) from the Lower Cretaceous Sinuiju Formation of Ryonsang-dong, North Phyongan Province, Democratic People's Republic of Korea". Cretaceous Research. 131: Article 105086. doi:10.1016/j.cretres.2021.105086. S2CID 243805918.

[270] Jouault, C.; Maréchal, A.; Condamine, F. L.; Wang, B.; Nel, A.; Legendre, F.; Perrichot, V. (2021). "Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth–death process". Zoological Journal of the Linnean Society. doi:10.1093/zoolinnean/zlab034.

[271] Radchenko, A. G.; Proshchalykin, M. Yu. (2021). "New extinct ant genus (Hymenoptera, Formicidae, Myrmicinae) from late Eocene Rovno amber". Journal of Hymenoptera Research. 84: 271–282. doi:10.3897/jhr.84.65238.

[272] Olmi, M.; Vasilenko, D. V.; Capradossi, L.; Perkovsky, E. E.; Guglielmino, A. (2021). "A new species of Lonchodryinus (Hymenoptera, Dryinidae) from Upper Eocene Baltic amber". Zootaxa. 5020 (2): 328–336. doi:10.11646/zootaxa.5020.2.5. PMID 34810999. S2CID 238658382.

[Megalyrhyssalus-273] Belokobylskij, S. A.; Jouault, C. (2021). "Two new striking braconid genera (Hymenoptera: Braconidae) from the mid-Cretaceous Burmese amber". Proceedings of the Geologists' Association. 132 (4): 426–437. doi:10.1016/j.pgeola.2021.04.003. S2CID 236271642.

[274] Gao, J.; Engel, M. S.; Shih, C.; Ren, D.; Gao, T. (2021). "A new genus of anaxyelid wood wasps from the mid-Cretaceous and the phylogeny of Anaxyelidae (Hymenoptera)". Journal of Hymenoptera Research. 86: 151–169. doi:10.3897/jhr.86.73161.

[275] Jouault, C.; Nel, A. (2021). "A new genus and species of Aulacidae (Hymenoptera: Evanioidea) from mid-Cretaceous Burmese amber" (PDF). Cretaceous Research. 124: Article 104799. doi:10.1016/j.cretres.2021.104799. S2CID 233846750.

[Paleoxyela-276] Jouault, C.; Aase, A.; Nel, A. (2021). "Past ecosystems drive the evolution of the early diverged Symphyta (Hymenoptera: Xyelidae) since the earliest Eocene". Fossil Record. 24 (2): 379–393. doi:10.5194/fr-24-379-2021.

[277] Yu, Y.; Hu, G.; Zhu, Y.; Li, Y.; Jiang, W.; Rasnitsyn, A. P.; Zhang, H.; Zhang, Q. (2021). "New material of Peleserphidae (Proctotrupoidea, Hymenoptera) in the mid-Cretaceous Burmese (Kachin) Amber". In S-C. Chang; D. Zheng (eds.). Mesozoic Biological Events and Ecosystems in East Asia. Geological Society, London, Special Publications. The Geological Society of London. doi:10.1144/SP521-2021-128. S2CID 244645180.

[278] Varela-Hernández, F.; Riquelme, F. (2021). "A New Ant Species of the Genus Pheidole Westwood, 1839 from Miocene Mexican Amber". Southwestern Entomologist. 46 (1): 75–82. doi:10.3958/059.046.0107. S2CID 232357684.

[Scleroxyela-279] Zheng, Y.; Hu, H.; Chen, D.; Chen, J.; Zhang, H.; Rasnitsyn, A. P. (2021). "New fossil records of Xyelidae (Hymenoptera) from the Middle Jurassic of Inner Mongolia, China". European Journal of Taxonomy (733): 146–159. doi:10.5852/ejt.2021.733.1229.

[280] Jouault, C.; Nel, A. (2021). "A new species of Prosyntexis Sharkey, 1990 (Hymenoptera: Sepulcidae) from the Lower Cretaceous Crato Formation of Brazil confirmed by geometric morphometric analysis". Palaeoentomology. 4 (2): 171–177. doi:10.11646/palaeoentomology.4.2.6. S2CID 235511059.

[281] Talamas, E. J.; Popovici, O.; Shih, C.; Ren, D. (2021). "Prototeleia Talamas, Popovici, Shih & Ren: A new genus of Platygastridae from Burmese amber". Journal of Hymenoptera Research. 87: 67–80. doi:10.3897/jhr.87.65472.

[282] Jouault, C.; Brazidec, M. (2021). "A new protopristocerine wasp (Hymenoptera: Bethylidae) from the mid-Cretaceous Burmese amber". Annales de Paléontologie. 107 (4): Article 102522. doi:10.1016/j.annpal.2021.102522. S2CID 244791642.

[283] Olmi, M.; Guglielmino, A.; Parise, G.; Capradossi, L.; Perkovsky, E. E. (2021). "Discovery of a new genus and species of Dryinidae (Hymenoptera: Chrysidoidea) from Kachin (Myanmar) amber: Rasnitsynum burmense gen. et sp. nov". Palaeoentomology. 4 (5): 416–420. doi:10.11646/palaeoentomology.4.5.3. S2CID 240513429.

[Palaeoentomology46Rovenosa-284] Manukyan, A. R.; Zhindarev, L. A. (2021). "Fossil Darwin wasps (Hymenoptera: Ichneumonidae) from Baltic amber". Palaeoentomology. 4 (6): 637–647. doi:10.11646/palaeoentomology.4.6.13. S2CID 245191818.

[285] Simutnik, S. A.; Perkovsky, E. E.; Vasilenko, D. V. (2021). "Sakhalinencyrtus leleji Simutnik gen. et sp. nov. of earliest Encyrtidae (Hymenoptera, Chalcidoidea) from Sakhalinian amber". Journal of Hymenoptera Research. 84: 361–372. doi:10.3897/jhr.84.66367.

[286] Davidian, E. M.; Manukyan, A. R.; Belokobylskij, S. A. (2021). "A new genus and species of the aphid parasitoids (Hymenoptera: Braconidae: Aphidiinae) from the Eocene Sakhalinian amber". Palaeoentomology. 4 (6): 537–543. doi:10.11646/palaeoentomology.4.6.3. S2CID 245166207.

[PeerJSpalangiopelta-287] Moser, M.; Burks, R. A.; Ulmer, J. M.; Heraty, J. M.; van de Kamp, T.; Krogmann, L. (2021). "Taxonomic description and phylogenetic placement of two new species of Spalangiopelta (Hymenoptera: Pteromalidae: Ceinae) from Eocene Baltic amber". PeerJ. 9: e10939. doi:10.7717/peerj.10939. PMC 8162234. PMID 34113480.

[288] Jouault, C.; Nel, A. (2021). "The oldest Cenozoic ant fossil: †Tyrannomecia gen. nov. (Formicidae: Myrmeciinae) from the Palaeocene Menat Formation (France)". Historical Biology. in press: 1–8. doi:10.1080/08912963.2021.2010192. S2CID 245194843.

[289] Abels, J.; Wedmann, S. (2021). "A fossil wasp (Hymenoptera: Vespidae: Vespula? hassiaca) from the Eocene of Messel". Palaeontographica Abteilung A. in press. doi:10.1127/pala/2021/0122. S2CID 245248645.

[290] Manukyan, A.R.; Smirnova, A.V. (2021). "New data on the family Siricidae (Hymenoptera, Symphyta) in Baltic amber". Paleontological Journal. 55 (2): 157–165. doi:10.1134/S003103012102009X. ISSN 0031-0301. S2CID 233746801.

[291] Zhuang, Y.; Ran, H.; Li, X.; Feng, C.; Liu, Y. (2021). "A new species of the iron maiden ant based on an alate female from mid-Cretaceous Burmese amber (Hymenoptera: Formicidae: †Zigrasimecia)". Cretaceous Research. 130: Article 105056. doi:10.1016/j.cretres.2021.105056. S2CID 240812443.

[292] Guo, Y.; Shih, C.; Zhuo, D.; Ren, D.; Zhao, Y.; Gao, T. (2021). "The first queen-worker association for Cretaceous Formicidae: the winged caste of Haidomyrmex cerberus". ZooKeys (1048): 69–78. doi:10.3897/zookeys.1048.66920. PMC 8280096. PMID 34305419.

[293] Radchenko, A. G.; Perkovsky, E. E. (2021). "Wheeler's dilemma revisited: first Oecophylla–Lasius syninclusion and other ants syninclusions in the Bitterfeld amber (late Eocene)". Invertebrate Zoology. 18 (1): 47–65. doi:10.15298/invertzool.18.1.05. S2CID 234800986.

[Acanthocorona-294] Mey, W.; Léger, T.; Lien, V. V. (2021). "New taxa of extant and fossil primitive moths in South-East Asia and their biogeographic significance (Lepidoptera, Micropterigidae, Agathiphagidae, Lophocoronidae)". Nota Lepidopterologica. 44: 29–56. doi:10.3897/nl.44.52350.

[ZittelianaFischer-295] Fischer, T. C. (2021). "In search for the unlikely: Leaf-mining caterpillars (Gracillariidae, Lepidoptera) from Upper Cretaceous and Eocene ambers". Zitteliana. 95: 135–145. doi:10.3897/zitteliana.95.63317. S2CID 245086588.

[296] Haug, J. T.; Haug, C. (2021). "A 100 million-year-old armoured caterpillar supports the early diversification of moths and butterflies". Gondwana Research. 93: 101–105. Bibcode:2021GondR..93..101H. doi:10.1016/j.gr.2021.01.009. S2CID 233548334.

[297] Maccracken, S. A.; Sohn, J.-C.; Miller, I. M.; Labandeira, C. C. (2021). "A new Late Cretaceous leaf mine Leucopteropsa spiralae gen. et sp. nov. (Lepidoptera: Lyonetiidae) represents the first confirmed fossil evidence of the Cemiostominae". Journal of Systematic Palaeontology. 19 (2): 131–144. doi:10.1080/14772019.2021.1881177. S2CID 233194385.

[298] Zhang, Y.; Shih, P. J. M.; Wang, J.; McNamara, M. E.; Shih, C.; Ren, D.; Gao, T. (2021). "Jurassic scorpionflies (Mecoptera) with swollen first metatarsal segments suggesting sexual dimorphism". BMC Ecology and Evolution. 21 (1): Article number 47. doi:10.1186/s12862-021-01771-3. PMC 7981964. PMID 33743581.

[299] Poschmann, M. J.; Nel, A. (2021). "The first Permian scorpionfly from Germany (Insecta, Panorpida: Protomeropidae)". Palaeoentomology. 4 (3): 231–236. doi:10.11646/palaeoentomology.4.3.10. S2CID 237881817.

[300] Lian, X.; Cai, C.; Huang, D. (2021). "A new orthophlebiid scorpionfly (Insecta, Orthophlebiidae) from the Late Jurassic Linglongta biota of northern China". Historical Biology. 33 (12): 3585–3589. doi:10.1080/08912963.2021.1878512. S2CID 233934478.

[Zootaxa49953-301] Lian, X.; Cai, C.; Huang, D. (2021). "New species of Mesopsyche Tillyard, 1917 (Mecoptera: Mesopsychidae) from the Triassic of northwestern China". Zootaxa. 4995 (3): 565–572. doi:10.11646/zootaxa.4995.3.10. PMID 34810550. S2CID 237782965.

[302] Badano, D.; Fratini, M.; Maugeri, L.; Palermo, F.; Pieroni, N.; Cedola, A.; Haug, J. T.; Weiterschan, T.; Velten, J.; Mei, M.; Di Giulio, A.; Cerretti, P. (2021). "X-ray microtomography and phylogenomics provide insights into the morphology and evolution of an enigmatic Mesozoic insect larva". Systematic Entomology. 46 (3): 672–684. doi:10.1111/syen.12482. S2CID 235363035.

[303] Jouault, C. (2021). "A new species of thorny lacewing (Neuroptera: Rhachiberothidae: Paraberothinae) from mid-Cretaceous Burmese amber with novel raptorial foreleg structure". Proceedings of the Geologists' Association. in press. doi:10.1016/j.pgeola.2021.11.001. S2CID 244457471.

[304] Cao, L.-R.; Wang, B.; Liu, X.-Y. (2021). "A new genus and species of Mesochrysopidae (Neuroptera) from the mid-Cretaceous Burmese amber". Palaeoentomology. 4 (1): 77–84. doi:10.11646/palaeoentomology.4.1.12. S2CID 233963511.

[Calobabinskaia-305] Lu, X.; Wang, B.; Liu, X. (2021). "New Cretaceous antlion-like lacewings promote a phylogenetic reappraisal of the extinct myrmeleontoid family Babinskaiidae". Scientific Reports. 11 (1): Article number 16431. Bibcode:2021NatSR..1116431L. doi:10.1038/s41598-021-95946-z. PMC 8361207. PMID 34385561.

[Fuscopolystoechotes-306] Xu, Y.; Zheng, B.; Fang, H.; Shih, C.; Ren, D.; Wang, Y. (2021). "New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)". Zootaxa. 5067 (2): 259–266. doi:10.11646/zootaxa.5067.2.7. ISSN 1175-5326. PMID 34810745. S2CID 243981058.

[307] Nakamine, H. (2021). "A new species of Mesochrysopidae (Neuroptera) from the Lower Cretaceous Crato Formation of Brazil". Palaeodiversity. 14 (1): 115–119. doi:10.18476/pale.v14.a4.

[Protosmylinae2021-308] Makarkin, V.N.; Archibald, S.B.; Mathewes, R.W. (2021). "New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks". Zootaxa. 4980 (1): 142–156. doi:10.11646/zootaxa.4980.1.9. PMID 34186986. S2CID 235685548.

[309] Khramov, A. V. (2021). "Osmyloberotha, an unusual new genus of beaded lacewings (Neuroptera: Berothidae) from Burmese amber". Zootaxa. 5060 (2): 95–99. doi:10.11646/zootaxa.5060.2.5. PMID 34811173. S2CID 240194221.

[310] Nakamine, H.; Yamamoto, S.; Takahashi, Y.; Li, H.; Liu, X. (2021). "A remarkable new thorny lacewing from mid-Cretaceous amber from northern Myanmar (Neuroptera: Rhachiberothidae)". PalZ. in press. doi:10.1007/s12542-021-00589-0. S2CID 244780786.

[311] Jouault, C.; Nel, A. (2021). "A new myrmeleontoid genus and species from the Lower Cretaceous Crato Formation (Neuroptera, Babinskaiidae)" (PDF). Palaeoentomology. 4 (3): 204–208. doi:10.11646/palaeoentomology.4.3.4. S2CID 237819174.

[312] Makarkin, V. N.; Ansorge, J.; Khramov, A. V. (2021). "Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s.l. (Neuroptera)". Palaeoentomology. 4 (6): 516–531. doi:10.11646/palaeoentomology.4.6.1. S2CID 245181387.

[313] Huang, D.; Lian, X.; Jouault, C.; Gao, J.; Nel, A. (2021). "First green lacewing (Insecta: Neuroptera: Chrysopidae) from the Palaeocene Sanshui basin of Guangdong, South China" (PDF). Historical Biology. in press: 1–9. doi:10.1080/08912963.2021.1943381. S2CID 237756011.

[314] Haug, G. T.; Haug, C.; Haug, J. T. (2021). "The morphological diversity of spoon-winged lacewing larvae and the first possible fossils from 99 million-year-old Kachin amber, Myanmar". Palaeodiversity. 14 (1): 133–152. doi:10.18476/pale.v14.a6.

[315] Haug, J. T.; Baranov, V.; Müller, P.; Haug, C. (2021). "New extreme morphologies as exemplified by 100 million-year-old lacewing larvae". Scientific Reports. 11 (1): Article number 20432. doi:10.1038/s41598-021-99480-w. PMC 8517013. PMID 34650142.

[316] Béthoux, O.; Norrad, R. E.; Stimson, M. R.; King, O. A.; Allen, L. F.; Deregnaucourt, I.; Hinds, S. J.; Lewis, J. H.; Schneider, J. W. (2021). "A unique, large-sized stem Odonata (Insecta) found in the early Pennsylvanian of New Brunswick (Canada)". Fossil Record. 24 (2): 207–221. doi:10.5194/fr-24-207-2021.

[317] Xia, G.; Zheng, D.; Krieg-Jacquier, R.; Fan, Q.; Chen, Y.; Nel, A. (2021). "The oldest-known Lestidae (Odonata) from the late Eocene of Tibet: palaeoclimatic implications". Geological Magazine. in press: 1–8. doi:10.1017/S0016756821001102. S2CID 242028737.

[318] Huang, D.; Fu, Y.; Lian, X.; Gao, J.; Nel, A. (2021). "The oldest malachite damselfly (Odonata: Synlestidae) from the Lower Cretaceous of China". Cretaceous Research. 129: Article 105023. doi:10.1016/j.cretres.2021.105023. S2CID 239681781.

[319] Nel, A. (2021). "Maastrichtian representatives of the dragonfly family Aeschnidiidae question the entomofaunal turnover of the early Late Cretaceous". Palaeoentomology. 4 (3): 209–212. doi:10.11646/palaeoentomology.4.3.5. S2CID 237876331.

[ZT49341-320] ^ ^a ^b ^c ^d ^e ^f ^g Archibald, S. B.; Cannings, R. A.; Erickson, R. J.; Bybee, S. M.; Mathewes, R. W. (2021). "The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America". Zootaxa. 4934 (1): zootaxa.4934.1.1. doi:10.11646/zootaxa.4934.1.1. PMID 33756770. S2CID 232337536.

[321] Felker, A.; Vasilenko, D. (2021). "A new species of the ʻprotozygopteranʼ damselfly (Odonata: Permagrionidae) from the Lower-Middle Permian of Russia". Palaeoentomology. 4 (5): 462–467. doi:10.11646/palaeoentomology.4.5.12. S2CID 240547006.

[322] Zheng, D.; Li, S.; Zhao, Y.; Zhang, H. (2021). "A potential telephlebiid dragonfly (Odonata: Anisoptera: Aeshnoidea) from Miocene of Yunnan, southwestern China". Palaeoentomology. 4 (3): 227–242. doi:10.11646/palaeoentomology.4.3.11. S2CID 237825025.

[PJ552Kennedyidae-323] Felker, A. S. (2021). "The first damselflies of the family Kennedyidae (Odonata: Protozygoptera) from the Permian-Triassic boundary deposits of the Kuznetsk Basin, Russia". Paleontological Journal. 55 (2): 166–173. doi:10.1134/S0031030121020052. S2CID 233746783.

[324] Felker, A. S. (2021). "New damselflies of the family Kennedyidae (Odonata: Protozygoptera) from the Upper Permian of the Vologda Region". Paleontological Journal. 55 (4): 396–404. doi:10.1134/S0031030121040067. S2CID 237375821.

[325] Deregnaucourt, I.; Bardin, J.; Anderson, J. M.; Béthoux, O. (2021). "The wing venation of a new fossil species, reconstructed using geometric morphometrics, adds to the rare fossil record of Triassic Gondwanian Odonata". Arthropod Structure & Development. 63: Article 101056. doi:10.1016/j.asd.2021.101056. PMID 33984598. S2CID 234494778.

[326] So, K. S.; Won, C. G. (2021). "The first 'Megapodagrionidae' (Odonata, Zygoptera) from the Lower Cretaceous of Democratic People's Republic of Korea". Cretaceous Research. 130: Article 105054. doi:10.1016/j.cretres.2021.105054. S2CID 244226045.

[327] Zheng, D.; Jarzembowski, E. A.; Zhuo, D.; Nel, A. (2021). "Protohemiphlebiidae fam. nov., a stem hemiphlebioid damselfly from Cretaceous amber in Kachin". In S-C. Chang; D. Zheng (eds.). Mesozoic Biological Events and Ecosystems in East Asia. Geological Society, London, Special Publications. The Geological Society of London. doi:10.1144/SP521-2020-249. S2CID 244087958.

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[329] Archibald, S. B.; Cannings, R. A. (2021). "A new genus and species of Euphaeidae (Odonata, Zygoptera) from the early Eocene Okanagan Highlands locality at Republic, Washington, U.S.A.". Zootaxa. 4966 (3): 392–400. doi:10.11646/zootaxa.4966.3.11. PMID 34186607. S2CID 235557114.

[330] Won, C.G.; So, K.S.; Kim, H.C. (2021). "A new odonatan (Stenophlebiidae) from the Lower Cretaceous Sinuiju Formation of North Phyongan Province, the Democratic People's Republic of Korea". Palaeoworld. in press. doi:10.1016/j.palwor.2021.02.002. ISSN 1871-174X. S2CID 233959783.

[331] Nel, A.; Zheng, D. (2021). "The recently proposed odonatan 'suborder' Cephalozygoptera: fact or fiction". Palaeoentomology. 4 (2): 165–170. doi:10.11646/palaeoentomology.4.2.5. S2CID 235536486.

[332] Archibald, S. B.; Cannings, R. A. (2021). "The head of Cephalozygoptera (Odonata)". Zootaxa. 5047 (1): 97–100. doi:10.11646/zootaxa.5047.1.10. PMID 34811004. S2CID 244256746.

[Amberotridactylus-333] Du, X.; Xu, C.; Zhang, L. (2021). "Two new pygmy mole crickets from mid-Cretaceous Kachin amber of northern Myanmar (Orthoptera: Tridactyloidea: Tridactylidae)". Cretaceous Research. 129: Article 105002. doi:10.1016/j.cretres.2021.105002. ISSN 0195-6671. S2CID 238648688.

[334] Gu, J.-J.; Yang, X.; Huang, R.; Yue, Y.; Ren, D. (2021). "New species and material of Hagloidea (Insecta, Ensifera) from the Yanliao biota of China". ZooKeys (1033): 183–190. doi:10.3897/zookeys.1033.63571. PMC 8084857. PMID 33958925.

[Caelielca-335] Uchida, K. (2021). "Two new genera of the Elcanidae (order: Orthoptera) from middle Cretaceous Burmese amber of northern Myanmar". Cretaceous Research. 131: Article 105092. doi:10.1016/j.cretres.2021.105092. S2CID 244377365.

[HBgrasshoppers-336] Nel, A.; Jouault, C. (2021). "New grasshoppers (Orthoptera: Elcanidae, Locustopsidae) from the Lower Cretaceous Crato Formation suggest a biome homogeneity in Central Gondwana". Historical Biology. in press: 1–9. doi:10.1080/08912963.2021.2000602. S2CID 244167833.

[337] Kim, D.-Y.; Lee, M.; Nam, G.-S.; Park, T.-Y. S. (2021). "The first orthopteran fossils from the Lower Cretaceous (Albian) Jinju Formation of Korea: ethological implications for elcanids". Cretaceous Research. 125: Article 104843. doi:10.1016/j.cretres.2021.104843.

[338] Gorochov, A. V. (2021). "The oldest representative of the superfamily Hagloidea (Orthoptera) from the Upper Permian of Russia, with remarks on katydid-like forms from the Paleozoic". Paleontological Journal. 55 (5): 552–558. doi:10.1134/S0031030121050063. S2CID 238640705.

[339] Xu, X.; Wang, B.; Fan, L.; Jarzembowski, E. A.; Fang, Y.; Wang, H.; Li, T.; Zhuo, D.; Ding, M.; Engel, M. S. (2021). "Widespread mimicry and camouflage among mid-Cretaceous insects". Gondwana Research. 101: 94–102. doi:10.1016/j.gr.2021.07.025. ISSN 1342-937X.

[340] Desutter-Grandcolas, L.; Hugel, S.; Nel, A.; Warren, B. H.; Souza-Dias, P.; Chintauan-Marquier, I. C. (2021). "Updated diagnoses for the cricket family Trigonidiidae (Insecta: Orthoptera: Grylloidea) and its subfamilies (Trigonidiinae, Nemobiinae), with a review of the fossil record". Zoologischer Anzeiger. 294: 80–91. doi:10.1016/j.jcz.2021.06.004.

[341] Chen, Z.-T. (2021). "Balticonemoura bulbosus gen. et sp. nov., a new stonefly of Nemouridae (Insecta: Plecoptera) from Eocene Baltic amber". Historical Biology. in press: 1–4. doi:10.1080/08912963.2021.1922399. S2CID 235528609.

[342] Chen, Z.-T.; Xu, C.-P. (2021). "New perlid stonefly (Insecta: Plecoptera) from mid-Cretaceous Burmese amber". Cretaceous Research. 127: Article 104954. doi:10.1016/j.cretres.2021.104954.

[343] Jouault, C.; Legendre, F.; Condamine, F. L.; Nel, A. (2021). "A new stonefly species (Plecoptera: Perlodidae) from Eocene Baltic amber and questions on the wing venation potential for species diagnostic of fossil Plecoptera" (PDF). Palaeoentomology. 4 (3): 243–256. doi:10.11646/palaeoentomology.4.3.12. S2CID 237847715.

[344] Yoshizawa, K.; Yamamoto, S. (2021). "The earliest fossil record of the suborder Psocomorpha (Insecta: Psocodea) from mid-Cretaceous Burmese amber, with description of a new genus and species". Insecta Matsumurana. New Series. 77: 1–15. hdl:2115/82942.

[345] Hakim, M.; Azar, D.; Huang, D.-Y. (2021). "A new species of Cormopsocidae from Burmese amber (Psocodea; Trogiomorpha)". Palaeoentomology. 4 (3): 213–217. doi:10.11646/palaeoentomology.4.3.6. S2CID 237857111.

[346] Hakim, M.; Azar, D.; Fu, Y.-Z.; Cai, C.-Y.; Huang, D.-Y. (2021). "A new cormopsocid from mid-Cretaceous Burmese amber (Psocodea: Trogiomorpha: Cormopsocidae)". Palaeoentomology. 4 (2): 178–185. doi:10.11646/palaeoentomology.4.2.7. ISSN 2624-2826. S2CID 236620426.

[Empherium-347] Hakim, M.; Huang, D.-Y.; Azar, D. (2021). "New fossil psocids from Cretaceous Siberian ambers (Psocodea: Trogiomorpha: Atropetae)". Palaeoentomology. 4 (2): 196–198. doi:10.11646/palaeoentomology.4.2.8. S2CID 235564657.

[348] Cumming, R. T.; Le Tirant, S. (2021). "Review of the Cretaceous †Archaeatropidae and †Empheriidae and description of a new genus and species from Burmese amber (Psocoptera)". Faunitaxys. Revue de Faunistique, Taxonomie et Systématique morphologique et moléculaire. 9 (16): 1–11.

[349] Wang, Q.; Li, S.; Ren, D.; Yao, Y. (2021). "New genus and species of †Cormopsocidae (Psocodae: Trogiomorpha) from mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 128: Article 104992. doi:10.1016/j.cretres.2021.104992.

[350] Hakim, M.; Huang, D.-Y.; Azar, D. (2021). "Earliest record of Prionoglarididae from the Lower Cretaceous Lebanese amber (Psocodea; Trogiomorpha)". Cretaceous Research. 132: Article 105121. doi:10.1016/j.cretres.2021.105121. S2CID 245304011.

[JouaultetalCRPsocodea-351] Jouault, C.; Yoshizawa, K.; Hakim, M.; Huang, D.; Nel, A. (2021). "New psocids (Psocodea: Prionoglarididae, Psyllipsocidae) from Cretaceous Burmese amber deposits" (PDF). Cretaceous Research. 126: Article 104890. doi:10.1016/j.cretres.2021.104890.

[352] Liang, F.; Liu, X. (2021). "A new species of Psyllipsocus (Psocodea: Trogiomorpha: Psyllipsocidae) from the mid-Cretaceous amber of Myanmar". Zootaxa. 5072 (1): 81–87. doi:10.11646/zootaxa.5072.1.9. S2CID 244682740.

[353] Liang, F.; Liu, X. (2021). "A new genus and species of the family Cormopsocidae (Psocodea: Trogiomorpha) from mid-Cretaceous amber of Myanmar". Cretaceous Research. 130: Article 105049. doi:10.1016/j.cretres.2021.105049. S2CID 244225553.

[354] Kiesmüller, C.; Haug, J. T.; Müller, P.; Hörnig, M. K. (2021). "Debris-carrying behaviour of bark lice immatures preserved in 100 million years old amber". PalZ. in press. doi:10.1007/s12542-021-00567-6.

[Archibald2021-355] Archibald, S. B.; Makarkin, V. N. (2021). "Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation". Zootaxa. 4951 (1): 41–79. doi:10.11646/zootaxa.4951.1.2. PMID 33903413. S2CID 233411745.

[Engel2011a-356] Engel, Michael S. (2011). "A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae)". Transactions of the Kansas Academy of Science. 114 (1–2): 77–87. doi:10.1660/062.114.0107. S2CID 85011364.

[357] Jouault, C.; Engel, M. S.; Nel, A. (2021). "A new baissopterid snakefly (Raphidioptera: Baissopteridae) from mid-Cretaceous amber of northern Myanmar". Cretaceous Research. 129: Article 105028. doi:10.1016/j.cretres.2021.105028. S2CID 239253683.

[CRMesoraphidia-358] Ri, C.-J.; Won, C.-G.; So, K.-S.; Ho, M.-S. (2021). "New snakeflies (Insecta: Raphidioptera: Mesoraphidiidae) from the Lower Cretaceous Sinuiju Formation of the Democratic People's Republic of Korea". Cretaceous Research. 132: Article 105105. doi:10.1016/j.cretres.2021.105105. S2CID 244700502.

[FEE444-359] Melnitsky, S. I.; Ivanov, V. D. (2021). "Two new species of the genus Archaeopolycentra (Trichoptera: Polycentropodidae) from Cretaceous Taimyr amber". Far Eastern Entomologist. 444: 1–7. doi:10.25221/fee.444.1. ISSN 1026-051X. S2CID 244880131.

[360] Wichard, W.; Neumann, C. (2021). "The polycentropodid genus Cernotina (Insecta, Trichoptera) in Miocene Dominican amber". Fossil Record. 24 (1): 129–133. doi:10.5194/fr-24-129-2021.

[361] Melnitsky, S. I.; Ivanov, V. D.; Perkovsky, E. E. (2021). "A new species of the fossil genus Electrotrichia (Insecta: Trichoptera: Hydroptilidae) from Rovno amber (Zhytomyr region, Olevsk locality)". Palaeoentomology. 4 (5): 421–424. doi:10.11646/palaeoentomology.4.5.4. S2CID 240542805.

[PJ555-362] ^ ^a ^b ^c ^d ^e ^f ^g Sukacheva, I. D.; Vasilenko, D. V. (2021). "Caddisflies (Insecta, Trichoptera) from the upper Mesozoic Chernovskie Kopi locality (Transbaikalia): part 2. Larval cases". Paleontological Journal. 55 (5): 545–551. doi:10.1134/S0031030121050154. S2CID 238640223.

[363] Melnitsky, S. I.; Ivanov, V. D.; Perkovsky, E. E. (2021). "A new species of the genus Holocentropus (Trichoptera: Polycentropodidae) from Rovno amber". Zoosystematica Rossica. 30 (2): 298–302. doi:10.31610/zsr/2021.30.2.298. S2CID 245185514.

[Palaeoentomology44Odontoceridae-364] Wichard, W.; Müller, P.; Xu, C.-P. (2021). "The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber". Palaeoentomology. 4 (4): 331–338. doi:10.11646/palaeoentomology.4.4.8. S2CID 239650542.

[365] Melnitsky, S. I.; Ivanov, V. D.; Perkovsky, E. E. (2021). "A new species of Plectrocnemia (Trichoptera: Polycentropodidae) from Rovno amber". Zootaxa. 5006 (1): 106–109. doi:10.11646/zootaxa.5006.1.14. PMID 34810582. S2CID 239644442.

[366] Sukatsheva, I. D.; Aristov, D. S. (2021). "New data on Cretaceous caddisflies family Vitimotauliidae (Insecta, Trichoptera) from Buryatia (Khasurty locality)". Paleontological Journal. 55 (4): 391–395. doi:10.1134/S0031030121330017. S2CID 237376766.

[WangetalCRWormaldia-367] Wang, J.; Zhang, W.; Shih, C.; Ren, D. (2021). "Two new mid-Cretaceous caddisflies (Trichoptera, Philopotamidae) from amber of Northern Myanmar". Cretaceous Research. 126: Article 104891. doi:10.1016/j.cretres.2021.104891.

[Anakitala-368] Aristov, D. S. (2021). "New eoblattid insects (Insecta: Eoblattida) from the Upper Permian of Russia". Paleontological Journal. 55 (1): 72–75. doi:10.1134/S0031030121010020. S2CID 232070926.

[369] Huang, D.; Fu, Y.; Lian, X.; Gao, J.; Nel, A. (2021). "The first Chinese Protohymenidae (Palaeoptera: Megasecoptera)". Historical Biology. in press: 1–4. doi:10.1080/08912963.2021.1929950. S2CID 236405039.

[Archaeogerarus-370] Nel, A.; Roques, P. (2021). "A new family and two new genera from Avion, Northern France, confirm the high Moscovian (late Carboniferous) diversity of the insect superorder Archaeorthoptera". Acta Palaeontologica Polonica. 66 (4): 879–884. doi:10.4202/app.00940.2021. S2CID 244879144.

[371] Nel, A.; Roques, P. (2021). "The second Caloneurodea from the Moscovian of Avion, France (Insecta, Archaeorthoptera)". Palaeoentomology. 4 (4): 320–322. doi:10.11646/palaeoentomology.4.4.5. ISSN 2624-2826. S2CID 239666503.

[372] Quispe, L.; Roques, P.; Garrouste, R.; Nel, A. (2021). "Carboniferous Blattinopsidae: revision of Klebsiella and new genus and species from Avion (Insecta, Paoliida)" (PDF). Historical Biology. in press: 1–7. doi:10.1080/08912963.2021.1916817. S2CID 234869567.

[373] Nel, A.; Roques, P. (2021). "A new strange Archaeorthoptera from the Moscovian of Avion (France) (Insecta, Polyneoptera)" (PDF). Historical Biology: 1–3. doi:10.1080/08912963.2021.1978082. S2CID 239222016.

[374] Storozhenko, S. Yu.; Aristov, D. S. (2021). "The latest record of the genus Belmophenopterum (Eoblattida: Mesorthopteridae) from the Middle Triassic of Kyrgyzstan". Far Eastern Entomologist. 437: 31–36. doi:10.25221/fee.437.6. S2CID 239678300.

[375] Rasnitsyn, A. P.; Aristov, D. S. (2021). "New species of the genus Blattinopsis Giebel, 1867 (Insecta: Blattinopsida: Blattinopsidae) from the Permian of Taimyr Peninsula, Russia". Far Eastern Entomologist. 437: 6–9. doi:10.25221/fee.437.2. S2CID 239636286.

[Cabarzopterum-376] Cui, Y.; Brauner, S.; Schneider, J. W.; Béthoux, O. (2021). "Grylloblattidan insects from Sperbersbach and Cabarz (Germany), two new early Permian and insect-rich localities". Journal of Paleontology: 1–20. doi:10.1017/jpa.2021.101. S2CID 243486826.

[Contracladus-377] Dvořák, T.; Pecharová, M.; Leipner, A.; Nel, A.; Prokop, J. (2021). "New archaeorthopteran insects from the Pennsylvanian of Piesberg reveal unexpected mosaic of morphological traits and colouration pattern of the tegmina". Historical Biology. 33 (12): 3375–3383. doi:10.1080/08912963.2020.1867127. S2CID 234189184.

[378] Chen, L.; Gu, J.-J.; Yang, Q.; Ren, D.; Blanke, A.; Béthoux, O. (2021). "Ovipositor and mouthparts in a fossil insect support a novel ecological role for early orthopterans in 300 million years old forests". eLife. 10: e71006. doi:10.7554/eLife.71006. PMC 8631945. PMID 34844668.

[NJGPA3003-379] Nel, A.; Poschmann, M. J. (2021). "Early Permian acercarian insects (Miomoptera, Palaeomanteidae) from the Saar-Nahe Basin (Rhineland-Palatinate, SW Germany)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 300 (3): 267–277. doi:10.1127/njgpa/2021/0989. S2CID 236233140.

[380] Logghe, A.; Garrouste, R.; Steyer, J.-S.; Pouillon, J.-M.; Nel, A. (2021). "The first Palaeodictyoptera (Insecta) from the Carboniferous-Permian basin of Graissessac (France)". Historical Biology. in press: 1–7. doi:10.1080/08912963.2021.1929953. S2CID 236354115.

[381] Nel, A.; Roques, P.; Duquesne, H. (2021). "A further strange Archaeorthoptera from the Moscovian of Nord-Pas-de-Calais (France) (Insecta, Polyneoptera)". Zootaxa. 5047 (2): 165–170. doi:10.11646/zootaxa.5047.2.5. PMID 34810773. S2CID 244251921.

[382] Rasnitsyn, A. P.; Aristov, D. S. (2021). "Review of the family Tococladidae Carpenter (Insecta: Eoblattida) from the Permian of North America and Europe". Paleontological Journal. 55 (2): 151–156. doi:10.1134/S003103012102012X. S2CID 233746323.

[383] Falières, E.; Engel, M. S.; Nel, A. (2021). "Earliest occurrence of Embiidae: A new genus from earliest Eocene Oise amber (Insecta: Embiodea)". Comptes Rendus Palevol. 20 (38): 799–805. doi:10.5852/cr-palevol2021v20a38.

[PJ556Glaphyrophlebia-384] Aristov, D. S.; Rasnitsyn, A. P.; Naugolnykh, S. V. (2021). "New Blattinopsidae (Insecta: Blattinopsida) in the Permian of the Pechora Basin (Komi Republic, Russia) in the context of landscape and vegetation evolution". Paleontological Journal. 55 (6): 641–649. doi:10.1134/S0031030121060022. S2CID 245009855.

[385] Aristov, D. S.; Rasnitsyn, A. P. (2021). "New and little known Blattinopsidae (Insecta: Blattinopsida) from the Middle Permian of Russia". Paleontological Journal. 55 (3): 288–293. doi:10.1134/S0031030121030059. S2CID 235966366.

[Multivena-386] Lai, Z.; Yang, Q.; Shi, C.; Zhang, W.; Ren, D. (2021). "New webspinners (Insecta: Embioptera) from Upper Cretaceous amber of northern Myanmar". Cretaceous Research. 130: Article 105040. doi:10.1016/j.cretres.2021.105040. S2CID 239202878.

[Kamopanorpa-387] Bashkuev, A. S.; Sukatsheva, I. D. (2021). "New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)". Palaeoentomology. 4 (5): 453–461. doi:10.11646/palaeoentomology.4.5.11. ISSN 2624-2826. S2CID 240525412.

[388] Aristov, D. S.; Storozhenko, S. Yu. (2021). "A new genus of the family Alexarasniidae (Insecta: Embiida) from the Triassic of Kyrgyzstan". Palaeoentomology. 4 (5): 433–435. doi:10.11646/palaeoentomology.4.5.7. S2CID 240549301.

[389] Yang, H.; Shih, C.; Rasnitsyn, A. P.; Ren, D.; Gao, T. (2021). "Early Cretaceous enigmatic insect group showing unique wing venations and antennal sensilla". Papers in Palaeontology. doi:10.1002/spp2.1402. S2CID 244172848.

[Moltenojurina-390] Béthoux, O.; Anderson, J. M. (2021). "The Polycytellidae viewed as Gondwanan Glosselytrodea". Palaeoentomology. 4 (6): 550–558. doi:10.11646/palaeoentomology.4.6.5. S2CID 245173292.

[391] Zessin, W.; Brauckmann, C.; Gröning, E. (2021). "A new insect (probably basal Odonatoptera) from the Pennsylvanian (Late Carboniferous) of the Piesberg Fossil-Lagerstätte, Osnabrück, Germany". Palaeoentomology. 4 (6): 532–536. doi:10.11646/palaeoentomology.4.6.2. S2CID 245170114.

[392] Nel, A.; Poschmann, M. J. (2021). "A new representative of the "orthopteroid" family Cnemidolestidae from the lower Permian of Germany". Acta Palaeontologica Polonica. 66 (3): 641–646. doi:10.4202/app.00879.2021.

[393] Nel, A.; Poschmann, M. J. (2021). "A new archaeorthopteran insect from the Early Permian of the Saar-Nahe Basin, SW Germany". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 301 (1): 79–84. doi:10.1127/njgpa/2021/0998. S2CID 237729949.

[394] Nel, A.; Roques, P. (2021). "The second species of the cnemidolestid genus Piesbergopterum from the Upper Carboniferous of Avion, Northern France (Archaeorthoptera: Cnemidolestidae)". Palaeoentomology. 4 (4): 323–325. doi:10.11646/palaeoentomology.4.4.6. S2CID 239687328.

[395] Gorochov, A. V. (2021). "New Family, Genus and Species of the Primitive Embioptera from the Triassic of Kyrgyzstan and Remarks on the Classification of this Order". Entomological Review. 101 (6): 837–845. doi:10.1134/S0013873821060087. S2CID 245108349.

[396] Brauckmann, C.; Herd, K. J.; Leipner, A. (2021). "First Eugereonidae (Insecta: Palaeodictyoptera) from the Pennsylvanian (Late Carboniferous) of the Piesberg site near Osnabrück, Germany". PalZ. in press. doi:10.1007/s12542-021-00578-3. S2CID 244745023.

[397] Schubnel, T.; Legendre, F.; Roques, P.; Garrouste, R.; Cornette, R.; Perreau, M.; Perreau, N.; Desutter-Grandcolas, L.; Nel, A. (2021). "Sound vs. light: wing-based communication in Carboniferous insects". Communications Biology. 4 (1): Article number 794. doi:10.1038/s42003-021-02281-0. PMC 8266802. PMID 34239029.

[398] Logghe, A.; Nel, A.; Steyer, J.-S.; Ngô-Muller, V.; Pouillon, J.-M.; Garrouste, R. (2021). "A twig-like insect stuck in the Permian mud indicates early origin of an ecological strategy in Hexapoda evolution". Scientific Reports. 11 (1): Article number 20774. Bibcode:2021NatSR..1120774L. doi:10.1038/s41598-021-00110-2. PMC 8531326. PMID 34675246.

[399] Haug, J. T.; Müller, P.; Haug, C. (2021). "Fossil dragonfly-type larva with lateral abdominal protrusions and implications on the early evolution of Pterygota". iScience. 24 (10): Article 103162. Bibcode:2021iSci...24j3162H. doi:10.1016/j.isci.2021.103162. PMC 8501664. PMID 34646993.

[400] Rosová, K.; Sinitshenkova, N. D.; Prokop, J. (2021). "Evidence for wing development in the Late Palaeozoic Palaeodictyoptera revisited". Arthropod Structure & Development. 63: Article 101061. doi:10.1016/j.asd.2021.101061. PMID 34098321. S2CID 235369956.

[401] Zhang, Q.; Zheng, D.; Wang, B.; Zhang, H. (2021). "Review of Triassic insects in China". In S-C. Chang; D. Zheng (eds.). Mesozoic Biological Events and Ecosystems in East Asia. Geological Society, London, Special Publications. The Geological Society of London. doi:10.1144/SP521-2021-121. S2CID 244645537.

[402] Xiao, L.; Labandeira, C.; Dilcher, D.; Ren, D. (2021). "Florivory of Early Cretaceous flowers by functionally diverse insects: implications for early angiosperm pollination". Proceedings of the Royal Society B: Biological Sciences. 288 (1953): Article ID 20210320. doi:10.1098/rspb.2021.0320. PMC 8207559. PMID 34132112.

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