2020 in paleobotany

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List of years in paleobotany
In paleontology
2017
2018
2019
2020
2021
2022

This article records new taxa of fossil plants that are scheduled to be described during the year 2020, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2020.

Flowering plants[]

Alismatales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[1]

Sp. nov

Valid

Low, Su & Xing in Low et al.

Late Oligocene

 China

A member of the family Araceae.

Apiales[]

Name Novelty Status Authors Age Type locality Location Notes Images

Paleopanax puryearensis[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

 United States
( Tennessee)

A member of the family Araliaceae.

Arecales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Arecaceae.

Palmoxylon ceroxyloides[4]

Sp. nov

In press

Khan, Hazra & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

 India

A petrified palm stem of a member of the subfamily Ceroxyloideae.

Palmoxylon dindoriensis[5]

Sp. nov

Valid

Khan, Roy & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

 India

A petrified palm stem.

Sabalites dawsonii[6]

Sp. nov

Valid

Greenwood & Conran

Eocene

 Canada
( British Columbia)

Sabalites karondiensis[7]

Sp. nov

In press

Roy, Hazra & Khan in Roy et al.

Late Cretaceous-Paleocene (latest Maastrichtian-earliest Danian)

Deccan Intertrappean Beds

 India

A palm frond .

[8]

Sp. nov

Valid

Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal

Late Cretaceous (Maastrichtian)

Lago Colhué Huapí Formation

 Argentina

A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

Buxales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Pachysandra. Announced in 2019; the final version of the article naming it was published in 2020.

Caryophyllales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant, possibly produced by members of the family Amaranthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Chloranthales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[11]

Gen. et sp. nov

Valid

Pipo, Iglesias & Bodnar

Late Cretaceous (early–middle Campanian)

Santa Marta Formation

Antarctica

A member of the family Chloranthaceae. Genus includes new species S. sanjosense.

Cornales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family .

[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A tupelo.

[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A tupelo.

[14]

Sp. nov

In press

Denk et al.

Late Oligocene

 Russia

[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

 United States
( Colorado)

A member of the family Mastixiaceae.

Crossosomatales[]

Name Novelty Status Authors Age Type locality Location Notes Images
[15] Sp. nov In press Zhu & Manchester Oligocene Renova Formation  United States
( Montana)
A species of Staphylea.

Cucurbitales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[16]

Sp. nov

Valid

Barreda, Palazzesi & Tellería in Renner et al.

Late Cretaceous (Campanian–early Maastrichtian)

Lopez de Bertodano Formation
Santa Marta Formation
Snow Hill Island Formation

Antarctica

Pollen grains similar to those of extant members of the genus Coriaria.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Cayaponia. Announced in 2020; the final version of the article naming it was published in 2021.

Ericales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

 United States
( Tennessee)

A member of the family Theaceae.

[17]

Gen. et sp. nov

Valid

Smith & Manchester

Middle Eocene

Clarno Formation

 United States
( Oregon)

A member of the family Theaceae. Genus includes new species A. andersonae.

Fabales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[18]

Sp. nov

Valid

Li et al.

Early Miocene

 China

A species of Cercis. Announced in 2020; the final version of the article naming it was published in 2021.

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Gleditsia. Announced in 2019; the final version of the article naming it was published in 2020.

[19] Sp. nov In press Baez & Crisafulli Miocene  Argentina Fossil wood of a member of the family Fabaceae.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Parkia. Announced in 2020; the final version of the article naming it was published in 2021.

[20]

Sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

 Panama

A species of Prioria.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Polygalaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of an eudicot, probably a member of the family Fabaceae.

Fagales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

An alder.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

 China

A member of the family Fagaceae.

[23]

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

 China

A species of Carpinus.

[23]

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

 China

A species of Carpinus

[24]

Sp. nov

In press

Swinehart & Farlow

Late Neogene

Pipe Creek Sinkhole

 United States
( Indiana)

A hickory.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A member of the family Fagaceae.

[25] Sp. nov Valid Mantzouka, Ivanov & Bozukov Late Miocene–early Pliocene (late Messinian–early Zanclean)  Bulgaria A species of Castanopsis. Announced in 2020; the final version of the article naming it was published in 2021.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

A species of Lithocarpus.

[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

An oak.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

 China

An oak.

Garryales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Eucommia. Announced in 2019; the final version of the article naming it was published in 2020.

Gentianales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[26]

Gen. et 2 sp. nov

Valid

Del Rio et al.

Early Eocene

 China

An asclepiadoid Apocynaceae genus.
Included species A. marginatum and A. ellipticum.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Rauvolfia. Announced in 2020; the final version of the article naming it was published in 2021.

Icacinales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[27]

Sp. nov

In press

Del Rio et al.

Early Oligocene

Wenshan Basin

 China

A member of the family Icacinaceae.

[28]

Sp. nov

Valid

Del Rio & De Franceschi

Early Eocene

 France

A member of the family Icacinaceae.

[29]

Gen. et sp. nov

In press

Stull & Rozefelds in Rozefelds et al.

Cenozoic (mid-Miocene or, more likely, middle Eocene)

 Australia

A member of the family Icacinaceae. Genus includes new species M. australis.

[30]

Sp. nov

Valid

Stull et al.

Early Oligocene

 Egypt

A species of Pyrenacantha.

Lamiales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Aegiphila. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Acanthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolpites pardoi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant, possibly produced by members of the genus Amphilophium. Announced in 2020; the final version of the article naming it was published in 2021.

Laurales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[31]

Gen. et sp. nov

In press

Akkemik et al.

Eocene (Lutetian)

 Turkey

A member of the family Lauraceae. Genus includes new species A. zileensis.

[32]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

 Argentina

A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

[32]

Gen. et sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

 Argentina

A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

[33]

Gen. et sp. nov

Valid

Chambers & Poinar

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra.

[34]

Gen. et sp. nov

Valid

Poinar et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Possibly a member of Laurales related to the families Monimiaceae and Atherospermataceae. Genus includes new species V. pleristaminis.

Liliales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[35]

Sp. nov

Valid

Dong et al.

Middle Miocene

 China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

[35]

Sp. nov

Valid

Dong et al.

Middle Miocene

 China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Magnoliales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[36]

Sp. nov

Valid

Huang et al.

Late Oligocene

 China

A species of Magnolia.

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Magnolia. Announced in 2019; the final version of the article naming it was published in 2020.

[37]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

Potomac Group

 United States
( Virginia)

A relative of extant early-diverging members of the Magnoliales. Genus includes new species M. virginiensis.

Malpighiales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[38]

Sp. nov

Valid

Narita et al.

Miocene

 Japan

A willow.

Malvales[]

Name Novelty Status Authors Age Type locality Location Notes Images
[19] Gen. et sp. nov In press Baez & Crisafulli Miocene  Argentina Fossil wood of a member of the family Malvaceae. Genus includes new species B. antiqua.
[39] Sp. nov Valid Khan, Spicer & Bera in Khan et al. Late Cretaceous (Maastrichtian) Deccan Intertrappean Beds  India A species of Dipterocarpus.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites jaramilloi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Hibiscus. Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites titanicus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Malachra. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the genus Ceiba. Announced in 2020; the final version of the article naming it was published in 2021.

[20]

Gen. et sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

 Panama

A member of the family Malvaceae. Genus includes new species V. panamense.

Myrtales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[40]

Sp. nov

Valid

Gandolfo & Zamaloa in Zamaloa, Gandolfo & Nixon

Eocene (Ypresian)

Huitrera Formation

 Argentina

A species of Eucalyptus.

[41]

Gen. et comb. nov

In press

Moya & Brea

Late Miocene?

Ituzaingó Formation

 Argentina

A member of the family Combretaceae; a new genus for "" areniensis Lutz (1979).

[32]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

 Argentina

A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

[42]

Gen. et sp. nov

Valid

Li & Li in Li et al.

Late Eocene to early Miocene

 China
 Czech Republic
 France
 Germany

A member of Trapoideae. Genus includes new species P. weichangensis, as well as "Carpolithus" pomelii Saporta (1878) and "Hemitrapa" alpina Su & Zhou in Su et al. (2018).

Nymphaeales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[43]

Gen. et sp. nov

Valid

Barrón, Peris & Labandeira in Peris et al.

Cenomanian

Burmese amber

 Myanmar

Pollen of a member of Nymphaeaceae.
Genus includes new species P. cenomaniensis.

Oxalidales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[44]

Gen. et sp. nov

Valid

Jud & Gandolfo

Paleocene (early Danian)

 Argentina

A member of the family Cunoniaceae. Genus includes new species C. bicarpellata. Announced in 2020; the final version of the article naming it was published in 2021.

[45]

Sp. nov

In press

Liu et al.

Late Oligocene

 China

A species of Elaeocarpus.

[45]

Sp. nov

In press

Liu et al.

Late Miocene

 China

A species of Elaeocarpus.

[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

[45]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

 China

A species of Elaeocarpus.

[45]

Sp. nov

In press

Liu et al.

Miocene

 China

A species of Elaeocarpus.

Poales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen produced by members of the family Poaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[46]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

 Argentina

A member of Cyperaceae. Genus includes new species R. cyperoides.

Proteales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[47]

Sp. nov

Valid

Carpenter in Carpenter & Milne

Late Eocene

 Australia

A species of Banksia.

Banksieaeidites zanthus[47]

Sp. nov

Valid

Milne in Carpenter & Milne

Late Eocene

 Australia

A Banksia-like pollen.

[48]

Sp. nov

Valid

Huegele, Spielbauer & Manchester

Miocene

 United States

A species of Platanus.

Ranunculales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Cissampelos. Announced in 2020; the final version of the article naming it was published in 2021.

[13]

Sp. nov

Valid

Hably

Miocene

 Hungary

A species of Clematis.

[50]

Sp. nov

Valid

Jia et al.

Late Miocene

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[49]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[51]

Gen. et sp. nov

Valid

Han et al.

Paleocene

 China

A member of the family Menispermaceae. Genus includes new species P. parvum.

[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

[51]

Sp. nov

Valid

Han et al.

Paleocene

 China

A species of Stephania

[51]

Sp. nov

Valid

Han et al.

Paleocene

 China

A species of Stephania

[49]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

 China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Rosales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[52]

Sp. nov

Valid

Zhou, Wang & Huang in Zhou et al.

Late Eocene

 China

A member of the family Rhamnaceae

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A species of Crataegus. Announced in 2019; the final version of the article naming it was published in 2020.

[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

 Kazakhstan

A member of the family Ulmaceae.

[3]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis.

Sapindales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

 Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Burseraceae.

[53]

Sp. nov

In press

Wang et al.

Miocene

 China

A species of Choerospondias.

[54] Gen. et sp. nov Rodríguez-Reyes, Estrada-Ruiz & Gasson Oligocene–Miocene  Panama A member of the family Anacardiaceae. Genus includes new species L. sandovalii.

[55]

Gen. et sp. nov

Valid

Atkinson

Late Cretaceous (Campanian)

 Canada
( British Columbia)

A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis.

[56]

Gen. et sp. nov

Valid

Franco et al.

Miocene

Ituzaingó Formation

 Argentina

A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum.

[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

 Mexico

Pollen of a flowering plant, probably a member of the family Meliaceae

[57]

Gen. et sp. nov

Valid

Manchester & Disney in Manchester, Disney & Pham

Eocene

Clarno Formation
Tepee Trail Formation

 United States
( Oregon
 Wyoming)

A fossil fruit with affinities with the Rutaceae. Genus includes new species Q. obovata.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Saxifragales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[58]

Sp. nov

Valid

Quirk & Hermsen

Early Pliocene

Gray Fossil Site

 United States
( Tennessee)

A species of Corylopsis. Announced in 2020; the final version of the article naming it was published in 2021.

[59]

Sp. nov

In press

Huang & Tian in Tian et al.

Oligocene

 China

A species of Itea.

[60]

Gen. et sp. nov

Valid

Scharfstein, Stockey & Rothwell

Late Cretaceous (Coniacian)

 Canada
( British Columbia)

A member of the family Altingiaceae. Genus includes new species P. comoxense.

Solanales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[61]

Sp. nov

Valid

Deanna, Wilf & Gandolfo

Early Eocene

Laguna del Hunco Formation

 Argentina

A species of Physalis.

Trochodendrales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[62]

Gen. et sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

 United States
( Wyoming)

A member of the family Trochodendraceae. Genus includes new species E. polystylum. Announced in 2020; the final version of the article naming it was published in 2021.

Paraconcavistylon[62]

Gen. et comb. nov

Valid

Manchester, Kvaček & Judd

Ypresian

Klondike Mountain Formation

 United States
( Washington)

A Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018). Announced in 2020; the final version of the article naming it was published in 2021.

[62]

Sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

 United States
( Wyoming)

A species of Trochodendron. Announced in 2020; the final version of the article naming it was published in 2021.

Vitales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[63]

Sp. nov

Valid

He & Wang

Miocene

 China

A species of Yua. Announced in 2020; the final version of the article naming it was published in 2021.

Other angiosperms[]

Name Novelty Status Authors Age Type locality Location Notes Images

[64]

Sp. nov

Valid

Vera et al.

Late Cretaceous

 Argentina

A fossil dicot wood

[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 Portugal
 United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species A. virginiensis.

[13]

Sp. nov

Valid

Hably & Erdei in Hably

Miocene

 Hungary

A fossil fruit of a flowering plant of uncertain phylogenetic placement.

[66]

Gen. et sp. nov

In press

Friis, Crane & Pedersen

Early Cretaceous

 Portugal

An early flowering plant. Genus includes new species C. dolichostemon.

[67]

Gen. et sp. nov

Valid

Siegert & Hermsen

Early Pliocene

Gray Fossil Site

 United States
( Tennessee)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil endocarps. Genus includes new species C. pratchettii.

[68]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Genus includes new species C. zeugostylus.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[69]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Genus includes new species C. papillosa.

[70]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus.

[71]

Gen. et sp. nov

Valid

Liu et al.

Miocene

Dominican amber

 Dominican Republic

A eudicot of uncertain phylogenetic placement. Genus includes new species D. pentamera.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[72]

Gen. et sp. nov

Valid

Han & Wang

Early Cretaceous

Yixian Formation

 China

An infructescence including a central axis and five fruits resembling . Genus includes new species E. liutiaogouensis.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species L. elegans.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides echibaculatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides romeroae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[73]

Gen. et sp. nov

Valid

Uhl, Paudayal & El Atfy in Uhl et al.

Paleocene (Thanetian)

 France

A eudicot of uncertain phylogenetic placement. Genus includes new species M mosbruggeri. Announced in 2020; the final version of the article naming it was published in 2021.

[65]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Aptian-early Albian)

 Portugal

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species M. portugallica.

[74]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla.

[75]

Sp. nov

Valid

Halamski & Kvaček in Halamski et al.

Late Cretaceous (Santonian)

 Poland

Trifoliolate platanoid leaves.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Psilaperiporites lunaris[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[76]

Gen. et sp. nov

Valid

Xing & Gu

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A possible epizoochorous fruit. Genus includes new species R. auricularus.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retibrevitricolporites? toigoae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

[77]

Gen. et sp. nov

Valid

Ramteke, Manchester & Nagrale in Ramteke et al.

Late Cretaceous (Maastrichtian)

Deccan Intertrappean Beds

 India

A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species S. kapgatei.

[78]

Gen. et sp. nov

Valid

Liu & Wang in Liu, Chen & Wang

Early Cretaceous (BarremianAptian)

Yixian Formation

 China

An early monocot. Genus includes new species S. ningchengensis. Announced in 2020; the final version of the article naming it was published in 2021.

[79]

Gen. et sp. nov

In press

Liu et al.

Early Cretaceous

Yixian Formation

 China

An early flowering plant. Genus includes new species V. lingyuanensis.

[80]

Gen. et sp. et comb. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian to Santonian)

 United States
( Alabama
 Massachusetts
 New Jersey
 New York)

A winged fruit of a eudicot of uncertain phylogenetic placement. Genus includes new species W. viccallii, as well as W. major (Hollick).

Pinales[]

Name Novelty Status Authors Age Type locality Location Notes Images

[81]

Sp. nov

Valid

Oskolski et al.

Late Oligocene–early Miocene

 China

A species of Agathis

Agathoxylon cozzoi[82]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

 Argentina

[83]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

 Argentina

A species of Araucaria.

[84]

Sp. nov

Valid

Stockey & Rothwell

Late Cretaceous (Campanian)

 Canada
( British Columbia)

A species of Araucaria.

[85]

Sp. nov

Valid

Shi et al.

Eocene

Antarctica
(King George Island)

A species of Araucaria

[86]

Sp. nov

Valid

Rossetto‐Harris in Rossetto‐Harris et al.

Early Eocene

Laguna del Hunco Formation

 Argentina

A species of Araucaria.

[83]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

 Argentina

A species of Araucaria.

Araucarites pachacuteci[87] Sp. nov

In press

Martínez in Martínez et al.

Early Cretaceous (BerriasianValanginian)

 Peru

A member of the family Araucariaceae

[88]

Sp. nov

Valid

Jiang et al.

Early Cretaceous

 China

A conifer wood. Announced in 2020; the final version of the article naming it was published in 2021.

Brachyphyllum sattlerae[89] Sp. nov Valid Batista et al. Early Cretaceous (Aptian) Crato Formation  Brazil A member of the family Pinidae.

[90]

Gen. et sp. nov

In press

Kvaček & Mendes

Early Cretaceous (late Aptian–early Albian)

Lusitanian Basin

 Portugal

A pollen cone of a member of Araucariaceae. Genus includes new species C. sousai.

[91]

Sp. nov

Valid

Akkemik

Early Miocene

 Turkey

A species of Cedrus. Announced in 2020; the final version of the article naming it was published in 2021.

[92]

Sp. nov

In press

Xia et al.

Middle Jurassic (Callovian)

 China

Possibly a member of the family Podocarpaceae.

[93]

Gen. et sp. nov

Valid

Stockey, Rothwell & Atkinson

Late Cretaceous (early Coniacian)

 Canada
( British Columbia)

A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species C. rossii.

[94]

Nom. nov

Valid

Blokhina

Miocene

 Russia

A member of the family Cupressaceae; a replacement name for Cupressinoxylon biotoides Blokhina (1989).

[82]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

 Argentina

Cupressinoxylon manuelii[95] Sp. nov In press Ríos-Santos, Cevallos-Ferriz & Pujana Late Cretaceous (Campanian-Maastrichtian) Cabullona Group  Mexico

[96]

Sp. nov

In press

Gou & Feng in Gou et al.

Middle Jurassic

Xishanyao Formation

 China

A conifer stem.

[97]

Gen. et sp. nov

In press

Mendes & Kvaček

Early Cretaceous (late Aptian – early Albian)

Lusitanian Basin

 Portugal

An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica.

[98]

Sp. nov

In press

Akkemik

Early Miocene

Hançili Formation

 Turkey

A member of the family Cupressaceae.

[99]

Sp. nov

Valid

Akkemik & Mantzouka in Akkemik, Mantzouka & Kıran Yıldırım

Miocene

Divriği Formation

 Turkey

A member of the family Pinaceae.

[100]

Sp. nov

Valid

Frolov & Mashchuk

Middle Jurassic

Prisayan Formation

 Russia

A member of the family Taxaceae.

[101]

Gen. et sp. nov

Valid

Stockey, Nishida & Rothwell

Late Cretaceous (late Campanian—early Maastrichtian)

 Japan

A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species M. satoi.

[102]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

 China

Fossil wood of a member of the family Pinaceae. Announced in 2020; the final version of the article naming it was published in 2021.

[103]

Sp. nov

Valid

Dolezych & Reinhardt

Paleogene

Eureka Sound Group

 Canada
( Nunavut)

A member of the family Pinaceae described on the basis of fossil wood

[102]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

 China

Announced in 2020; the final version of the article naming it was published in 2021.

[104]

Sp. nov

Valid

Wu & Zhou in Wu et al.

Late Paleogene

 China

A species of Tsuga. Announced in 2019; the final version of the article naming it was published in 2020.

Other seed plants[]

Name Novelty Status Authors Age Type locality Location Notes Images

[105]

Sp. nov

In press

Shi, Yu & Sun

Permian (Lopingian)

 China

A root of a member of Cordaitales

[106]

Sp. nov

In press

Bodnar et al.

Late Triassic

 Argentina

A member of Cycadales.

[107]

Gen. et sp. nov

Valid

Seyfullah, Roberts, Schmidt & Kunzmann in Seyfullah et al.

Early Cretaceous (Aptian-Albian)

Crato

 Brazil

A seed plant belonging to the group Erdtmanithecales. Genus includes new species A. resinosus.

[108]

Gen. et sp. nov

Valid

Naugolnykh

Carboniferous

 Russia

A member of Pinophyta belonging to the group . Genus includes new species A. progressus.

[109]

Gen. et sp. nov

In press

Mendes, Pedersen & Friis

Early Cretaceous

 Portugal

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species B. hirsutum.

[110]

Sp. nov

Valid

Gómez et al.

Early Cretaceous (Aptian)

La Cantera Formation

 Argentina

A fossil seed, possibly produced by a member of Gnetales.

[111]

Gen. et sp. nov

Valid

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae.

[112]

Gen. et 2 sp. nov

In press

Zhou et al.

Late Permian

 China

A gigantopterid. Genus includes new species F. asymmetrica and F. dahaia.

[113]

Sp. nov

In press

Deng, Yang & Zhou

Early Cretaceous

 China

A species of Ginkgo.

[114]

Gen. et sp. nov

Valid

Yang, Wang & Ferguson

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A member of Ephedraceae. Genus includes new species J. verticillata.

[115]

Gen. et sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales. Genus includes new species J. multinerve.

[116]

Gen. et comb. nov

Valid

Anderson in Anderson et al.

Permian (possibly Lopingian)

Um Irna Formation

 Jordan

A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017).

Kaokoxylon brasiliensis[111]

Sp. nov

Valid

Conceição, Neregato & Iannuzzi in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

 Brazil

A conifer described on the basis of fossil wood.

Nilssonia mirovanae[117]

Sp. nov

Valid

Čepičková & Kvaček

Late Cretaceous (Cenomanian)

Peruc-Korycany Formation

 Czech Republic

A cycad.

[118]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

 Brazil

A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi.

[119]

Sp. nov

Valid

Naugolnykh & Linkevich

Permian (Artinskian)

 Russia
( Sverdlovsk Oblast)

A member of Ginkgoales belonging to the family .

[120]

Gen. et sp. nov

In press

Wan, Yang & Wang

Late Permian

 China

A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale.

[121]

Sp. nov

Valid

Nosova in Zolina et al.

Late Cretaceous–Paleocene (MaastrichtianDanian)

 Russia
( Chukotka Autonomous Okrug)

A member of Czekanowskiales

[115]

Sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales

[122]

Sp. nov

Valid

Conran et al.

Middle Miocene

 New Zealand

A cycad

Pteruchus frenguellii[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A pollen organ of a seed fern

Pteruchus lepidus[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A pollen organ of a seed fern

Samaropsis jinchangensis[124]

Sp. nov

Valid

Hua & Sun in Hua et al.

Early Permian

 China

A seed fossil. Announced in 2019; the final version of the article naming was published in 2020.

[125]

Sp. nov

In press

Nosova

Middle Jurassic

 Uzbekistan

Umaltolepis sogdianica[125]

Sp. nov

In press

Nosova

Middle Jurassic

Angren Formation

 Uzbekistan

Umkomasia aequatorialis[123]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

 Jordan

A cupulate structure of a seed fern

[115]

Gen. et sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

 United States
( Texas)

A member of Voltziales. Genus includes new species W. acaulis.

[126]

Gen. et sp. nov

Valid

Pšenička et al.

Early Permian

 China

A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii.

[127]

Gen. et comb. nov

In press

Zhou et al.

Permian (Asselian)

 China

A seed fern; a new genus for "Alethopteris" ascendens Halle.

[118]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

 Brazil

A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi.

Other plants[]

Name Novelty Status Authors Age Type locality Location Notes Images

Annularia paisii[128]

Sp. nov

Valid

Correia et al.

Carboniferous (Gzhelian)

 Portugal

[129]

Sp. nov

In press

Li et al.

Early Cretaceous

 China

A liverwort belonging to the family Blasiaceae.

[130]

Sp. nov

In press

He et al.

Permian (Lopingian)

 China

A fern

Calamites cambrensis[131]

Sp. nov

Valid

Thomas

Carboniferous (Pennsylvanian)

 United Kingdom

[132]

Gen. et sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

 China

A green alga belonging to the group Dasycladales. Genus includes new species C. gigantia.

[133]

emend. nov.

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

 Spain

Thallus of . Both taxa has been firstly found annatomically attached

[134]

Sp. nov

Vršanská & Hinkelman

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Chlamydomonas

[46]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

 Argentina

A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides.

[135]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

var. [136]

comb. nov

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

 Spain

Clavatoracean species.

[137] Sp. nov Valid Bucur & Rigaud in Bucur et al. Late Triassic (Norian)  United States
( Idaho)
A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae.

[135]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Coniopteris sandaolingensis[138]

Sp. nov

Valid

Yuan & Sun in Yuan et al.

Middle Jurassic

 China

[139]

Gen. et comb. nov

Valid

Thomas & Cleal

Carboniferous

 United Kingdom

A lycophyte; a new genus for "Lepidodendron" hickii.

[140]

Sp. nov

Valid

Yu & Xie in Yu et al.

Late Miocene

 China

A fern belonging to the family Polypodiaceae.

[141]

comb. nov

Valid

Pérez-Cano, Bover-Arnal et MArtín-Closas in Pérez-Cano et al.

Barremian

 Spain

A member of .

[142]

Sp. nov

Valid

Zhang & Yan in Zhang et al.

Late Triassic

 China

A member of Equisetales. Announced in 2020; the final version of the article naming was published in 2021.

[143]

Sp. nov

Valid

Aung et al.

Late Miocene

 Vietnam

A species of Equisetum

[143]

Sp. nov

Valid

Aung et al.

Late Pliocene

 Vietnam

A species of Equisetum

[144] Gen. et sp. nov In press Sokač & Grgasović Early Paleocene  Croatia A green alga belonging to the group Dasycladales. Genus includes new species F. barattoloi.

[145]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A liverwort, a species of Frullania

[146]

Sp. nov

Valid

Mamontov et al.

Eocene

Rovno amber

 Ukraine

A liverwort, a species of Frullania

[147]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

 Australia

A member of Zosterophyllaceae. Genus includes new species G. minutus.

[148] Gen. et sp. nov Valid Pfeiler & Tomescu Devonian (Emsian) Battery Point  Canada
( Quebec)
An early euphyllophyte. Genus includes new species G. tetraxylopteroides. Announced in 2020; the final version of the article naming it was published in 2021.
[137] Sp. nov Valid Bucur & Peybernes in Bucur et al. Late Triassic  Japan A green alga belonging to the group Dasycladales and the family .
[149] Gen. et sp. nov In press Zhou et al. Permian (Asselian)  China An anachoropterid fern. Genus includes new species H. uncinatus.
[137] Sp. nov Valid Bucur & Fucelli in Bucur et al. Late Triassic (Norian)  United States
( Nevada)
A green alga belonging to the group Dasycladales and the family Triploporellaceae.
Holosporella? rossanae[137] Sp. nov Valid Bucur & Del Piero in Bucur et al. Late Triassic (Norian)  Canada
( Yukon)
A green alga belonging to the group Dasycladales and the family Triploporellaceae.

[150]

Gen. et comb. nov

Valid

LoDuca et al.

Silurian

 United States

An alga related to the group Bryopsidales. Genus includes "" divaricata White (1901), "B." newlini White (1901), "B." lesquereuxi Grote & Pitt (1876) and "Chondrites" verus Ruedemann (1925). Announced in 2020; the final version of the article naming it was published in 2021.

[132]

Sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

 China

A green alga belonging to the group Dasycladales.

[151]

Gen. et sp. nov

Valid

Chen et al.

MiddleLate Jurassic

Jiulongshan Formation

 China

A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis.

[152]

Gen. et sp. nov

Valid

Champreux, Meyer-Berthaud & Decombeix

Devonian (Famennian)

Mandowa Mudstone Formation

 Australia

A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae.

[153]

Gen. et sp. nov

Valid

Mamontov in Kopylov et al.

Early Cretaceous

 Russia

A liverwort belonging to the group Marchantiidae. Genus includes new species K. ginkgoides.

Lepidodendron demkinae[154]

Sp. nov

Valid

Mosseichik

Carboniferous (Viséan)

 Russia

[155]

Sp. nov

Valid

Xu et al.

Middle Triassic

 China

A member of Equisetales.

[156]

Sp. nov

Valid

Naugolnykh et al.

Paleocene

 China

A species of Lygodium. Announced in 2019; the final version of the article naming it was published in 2020.

Metzgeriites kujiensis[157]

Sp. nov

Valid

Katagiri in Katagiri & Shinden

Late Cretaceous (Santonian)

 Japan

A liverwort.

[158]

emend. nov

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Lebanon

 Lebanon

Clavatoracean thallus.

[159] Gen. et sp. nov In press Gnaedinger et al. Middle Triassic  Argentina A member of Equisetales. Genus includes new species N. gondwanaensis.

[160]

Sp. nov

Valid

Cheng et al.

Cretaceous

 China

A member of the family Osmundaceae

[160]

Sp. nov

Valid

Cheng et al.

Cretaceous

 China

A member of the family Osmundaceae

[161]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

 Argentina

A green alga belonging to the group Zygnematales.

[161]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

 Argentina

A green alga belonging to the group Zygnematales.

[162]

Sp. nov

In press

Liu et al.

Permian (Asselian)

 China

A member of the family Calamitaceae.

[147]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

 Australia

A member of Zosterophyllaceae. Genus includes new species P. timsiae.

[137] Sp. nov Valid Bucur & Rigaud in Bucur et al. Late Triassic (Norian)  United States
( Oregon)
A green alga belonging to the group Dasycladales and the family Polyphysaceae.
Patruliuspora pacifica[137] Sp. nov Valid Bucur, Del Piero & Peyrotty in Bucur et al. Late Triassic (Norian)  Canada
( Yukon)
A green alga belonging to the group Dasycladales and the family Polyphysaceae.
[163] Sp. nov Valid Li et al. Middle Jurassic  China A liverwort belonging to the family Pelliaceae.

[164]

Sp. nov

In press

Hiller et al.

Eocene

 Vietnam

A member of Osmundaceae

Pleuromeia shaolinii[165]

Sp. nov

Valid

Zhang & Wang in Zhang et al.

Middle Triassic

 China

[135]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla

 Argentina

Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it was published in 2020.

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

[166]

Sp. nov

In press

Álvarez-Vázquez, Bek & Drábková

Carboniferous (Pennsylvanian)

 Spain

A member of Isoetales

[167]

Sp. nov

Valid

Kvaček in Kvaček & Teodoridis

Oligocene

 Czech Republic

A fern, a species of Polystichum

[168]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis.

[169]

Sp. nov

Valid

Tang et al.

Mesoproterozoic
circa 1 Ga

 China

An early siphonocladalean chlorophyte

[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D’Apolito

Miocene

Solimões Basin

 Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

[170]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Middle Miocene

 Spain

A charophyte.

[171]

Gen. et sp. nov

Valid

Huang et al.

Late Devonian

Wutong Formation

 China

A plant of uncertain phylogenetic placement, possibly a lycopsid or a sphenopsid. Genus includes new species Q. mira.

[172]

Sp. nov

In press

Wan et al.

Early Permian

 China

A fern belonging to the group Marattiales.

Sigillaria pfefferkornii[173]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

 China

Sigillaria wudensis[173]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

 China

[170]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Miocene

 Lebanon
 Spain

A charophyte.

[174]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A species of Thyrsopteris

Tomiostrobus sinensis[175]

Sp. nov

Valid

Feng in Feng et al.

Early Triassic (Induan)

 China

A member of the family Isoetaceae.

[176]

Sp. nov

Valid

Naugolnykh

Permian (Roadian)

Pechora coal basin

 Russia

A fern belonging to the family Gleicheniaceae.

[177]

Sp. nov

Valid

Tang et al.

Late Permian

 China

A lycopsid belonging to the family

[178]

Gen. et sp. nov

Valid

Mosseichik & Filimonov

Carboniferous (Tournaisian)

 Russia
( Krasnoyarsk Krai)

A member of Lycopodiopsida. Genus includes new species U. asiaticum.

[179]

Sp. nov

Valid

Thomas et al.

Carboniferous (Bashkirian)

 United Kingdom

A fern

General research[]

  • A study on the evolutionary history of green plants is published by Nie et al. (2020).[180]
  • Description of new fossil material of uniformis from the Cambrian Yanjiahe Formation (China) and a study on the phylogenetic relationships and possible life cycle of this organism is published by Shang et al. (2020), who consider Y. uniformis to be a likely green microalga.[181]
  • A study on the phylogenetic relationships of extant and fossil complex thalloid liverworts (Marchantiidae) is published by Flores et al. (2020).[182]
  • Evidence of development of dichotomous roots in euphyllophytes that were extant during the Devonian and Carboniferous periods is presented by Hetherington, Berry & Dolan (2020), who interpret their findings as indicating that dichotomous root branching evolved in both lycophytes and euphyllophytes.[183]
  • An early land plant producing multiple spore size classes is described from the Lower Devonian Campbellton Formation (Canada) by Bonacorsi et al. (2020).[184]
  • A study on the impact of the appearance and evolution of herbivorous tetrapods on the evolution of land plants from the Carboniferous to the Early Triassic is published by Brocklehurst, Kammerer & Benson (2020).[185]
  • A study on the production of periderm in Late Paleozoic arborescent lycopsids is published by D'Antonio & Boyce (2020), who argue that these lycopsids did not grow from sporelings into large trees through the production of a periderm cylinder, because physiological limitations would have prohibited the production of thick periderm.[186]
  • A study on the architecture and development of the Carboniferous arborescent lycopsid is published by DiMichele & Bateman (2020).[187]
  • New information on the anatomy of Weichselia reticulata is presented by Blanco‐Moreno, Decombeix & Prestianni (2020).[188]
  • A study on the phylogenetic placement of the extinct fern genus Coniopteris is published by Li et al. (2020).[189]
  • New information on the morphology of patagonica is presented by Benedetti et al. (2020), who evaluate the implications of this taxon for the knowledge of the evolution of water ferns.[190]
  • A study aiming to determine which ferns were most likely to be the producers of spores from earliest Paleocene plant localities across western North America, and were most likely to be among the first plants in western North America to thrive in the immediate aftermath of the Cretaceous–Paleogene extinction event, is published by Berry (2020).[191]
  • A study on the morphology and development of , and on its implications for the knowledge of the evolutionary origins of seed development, is published by Meade, Plackett & Hilton (2020).[192]
  • A pollen organ resembling seed fern pollen organs and is described from the Lopingian Umm Irna Formation (Jordan) by Zavialova et al. (2020), who interpret this finding as evidence of persistence of lyginopterid seed ferns until the late Permian.[193]
  • Evidence of increasing atmospheric CO2 concentration at the onset of the end-Triassic extinction event, based on data from fossil leaves of the seed fern Lepidopteris ottonis from southern Sweden, is presented by Slodownik, Vajda & Steinthorsdottir (2020), who confirm L. ottonis as a valid proxy for pCO2 reconstructions.[194]
  • A study on the anatomy of the seed cone scales of mongolica is published by Herrera et al. (2020), who argue that K. mongolica and Podozamites harrisii are the seed cones and leaves of the same extinct plant, and name a new family Krassiloviaceae within the order Voltziales.[195]
  • A study on the microscopic wood anatomy of a fossil tree trunk of Agathoxylon arizonicum with the characteristic external features of a fire scar from the Upper Triassic Chinle Formation (Petrified Forest National Park, Arizona, United States) is published by Byers et al. (2020), who evaluate the implications of this specimen for the knowledge of the evolution of fire-adapted plant traits.[196]
  • A putative bamboo "Chusquea" oxyphylla from the early Eocene Laguna del Hunco biota (Argentina) is reinterpreted as a conifer by Wilf (2020), who transfers this species to the genus Retrophyllum.[197]
  • A study on evolutionary history of conifers as indicated by fossil and molecular data, aiming to determine whether the rise of angiosperms drove the decline of conifers and other gymnosperms, is published by Condamine et al. (2020).[198]
  • Presence of secretory tissues is reported in extinct flowers from the Cretaceous amber from Myanmar and Cenozoic Dominican amber (including specimens preserved while in the process of emitting compounds) by Poinar & Poinar (2020).[199]
  • Fossil pollen of flowering plants is reported from the Aptian and Albian of Australia by Korasidis & Wagstaff (2020), who evaluate the implications of their findings for the knowledge of the timing of the appearance and diversification of the flowering plants in the high-latitude southern basins of Australia.[200]
  • A study on the morphology of palm and palm-like pollen from the Eocene (Myanmar), and on the implications of these fossils for the knowledge of distribution and diversity of Eocene palms across the globe, is published by Huang et al. (2020).[201]
  • Fossils fruits of , representing the second fossil occurrence of Illigera worldwide and the first in Asia, are described from the Eocene (central Tibetan Plateau) by Wang et al. (2020), who evaluate the implications of this finding for the knowledge of the climate in the central Tibetan Plateau during the early middle Eocene, and for the knowledge of the floristic links between Asia and North America during the Paleogene.[202]
  • A study on the morphology and phylogenetic relationships of Montsechia vidalii is published by Gomez et al. (2020).[203]
  • Eocene leaves of members of the family Urticaceae with stinging trichomes are described from the Okanogan Highlands (British Columbia, Canada) by DeVore et al. (2020).[204]
  • A revision of the fossil record of the family Nothofagaceae from South America is published by Pujana et al. (2020).[205]
  • A study on the extinction of plants from south polar terrestrial ecosystems during the Permian–Triassic extinction event and on their recovery after this extinction event, based on data from the Sydney Basin (Australia), is published by Mays et al. (2020).[206]
  • A study on the impact of ecological disturbances around the Permian–Triassic boundary (from the Wuchiapingian to Ladinian) on land plant communities is published by Nowak, Vérard & Kustatscher (2020).[207]
  • A study on the age of the Paleogene Kanaka Creek fossil flora (Huntingdon Formation; British Columbia, Canada) and on its implications for reconstructions of the contemporaneous paleoclimate and paleoenvironment is published by Mathewes, Greenwood & Love (2020).[208]
  • Evidence from Eocene plant fossils from the Bangong-Nujiang suture indicating that the Tibetan Plateau area hosted a diverse subtropical ecosystem approximately 47 million years ago and that this area was both low and humid at the time is presented by Su et al. (2020), who also report that the composition of this flora is similar to Early-Middle Eocene floras in both North America and Europe, but shows little affinity to Eocene floras from the Indian Plate.[209]
  • A study aiming to estimate leaf dry mass per area in fossil plants from 22 western North American sites spanning the Eocene–Oligocene transition is published online by Butrim & Royer (2020), who evaluate the implications of their findings for the knowledge of the impact of the environmental changes occurring during the Eocene–Oligocene transition on leaf‐economic strategies of plants.[210]
  • A study on the Neogene paleobotanical record and climate in the northernmost part of the Central Andean Plateau, based on data from the (Peru), is published by Martínez et al. (2020), who report the earliest evidence of a puna-like ecosystem in the Pliocene and a montane ecosystem without modern analogs in the Miocene.[211]
  • Fossil fruits (mericarps) of the neoendemic Apiaceae Melanoselinum (Daucus) decipiens were reported from the lacustrine and fluvial sediments of Porto da Cruz, Madeira, dated 1.3 Ma, by Góis-Marques et al. 2020.[212] This paper not only reports the oldest Daucus s.l. fossil known to date but also the first fossil evidence of a plant with insular woodiness (see Island gigantism).
  • The leaf fossil Mesodescolea plicata from the Early Cretaceous of Patagonia, first interpreted as a cycad with affinities with extant Stangeria, is reinterpreted as an angiosperm leaf with affinities with Austrobaileyales or Chloranthales by Coiro et al. 2020,[213] with implications for the evolution of leaf shape in the early radiation of the angiosperms.
  • A study on the phylogenetic relationships of 10 Cretaceous flower taxa ( endressii, Dakotanthus cordiformis, lusitanicus, mirabilis, svitkoana, chevalieri, Paradinandra suecica, mauldiniensis, crossmanensis and calycanthoides) is published by Schönenberger et al. (2020).[214]

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