Paleobiota of the Posidonia Shale
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The Sachrang Formation or "Posidonienschiefer" Formation (Whose Vulgar name is "Posidonia Shale") is a geological formation of southwestern Germany, northern Switzerland, northwestern Austria, southeast Luxembourg and the Netherlands, that spans about 3 million years during the Early Jurassic period (early Toarcian stage). It is known for its detailed fossils, especially sea fauna, listed below.[1] Composed mostly by black shale, the formation is a Lagerstätte, where fossils show exceptional preservation (Including exquisite soft tissues), with a thickness that varies from about 1 m to about 40 m on the Rhine level, being on the main quarry at Holzmaden between 5 and 14 m.[1] Some of the preserved material has been transformed into fossil hydrocarbon Jet, specially wood remains, used for jewelry.[2] The exceptional preservation seen on the Posidonia Shale has been studied since the late 1800s, finding that a cocktail of chemical and environmental factors let to such an impressive conservation of the marine fauna.[2] The most common theory is the changes on the oxygen level, where the different anoxic events of the Toarcian left oxygen-depleted bottom waters, with the biota dying and falling to the bottom without any predator able to eat the dead bodies.[3]
Microbial activity[]
Color key
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Notes Uncertain or tentative taxa are in small text; |
Genus | Species | Location | Material | Notes | Images |
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Stromatolites[4] |
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Traces of Microbial Activity |
Non-fenestrate stromatolite crusts formed in Aphotic deep-water environments during intervals of very low sedimentation.[4] Abundant on the Precambrian, but after it, and concretely on the Jurassic-Cretaceous, the appearance of the Corallinaceae algae and related biota forced the stromatolite-forming microbes to withdraw to extreme habitats such as hypersaline lagoons and possibly to deep-water settings. On the Sachrang Formation are related with plankton, mainly coccoliths and the problematic (A Haptophytan Alga), but also typical deep-sea forms including various groups of cephalopods, and articulated skeletons of fishes and reptiles.[4] The Stromatolites of this region have evidence of live on a deeper shelf environment with a quietwater deposit which suffered repeated phases of stagnant bottom waters, where a depth water habitat developed, probably at more than 100 meters depth.[4] There is a thin, southern widespread Stromatolite crust on the Top of the Sachrang Formation, called "Wittelshofener Bank", that has made rethink the depth of the major southern basin of the formation, where with the absence of phototrophic calcareous benthic organisms (probably due to the lack of light), shows the deph character of the Basin.[4] On the "Wittelshofener Bank" there is also the only occurrence of Ooids, presumably formed in the same deep-water environment.[4] |
![]() Example of Stromatolites fossil trace, done probably by Cyanobacteria |
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Possible traces of Microbial Activity |
Probably related with Archaea activity.[4] Although Frutexites is a cryptic microfossil and an important element of many deep water stromatolites, with an Inorganic origin proposed, where are interpreted as dendritic shrubs to purely inorganic growth of Aragonitic crystals, but also resemble shrubs of the cyanobacteria .[4] On the Posidonia a cryptoendopelitic mode of life is assumed, being only possible for Heterotrophic bacteria or Fungi.[4] As seen on the Stromatolites of the Posidonia, Frutexites acted mainly as a dweller or secondary binder of the deep-water stromatolites, not as their major constructor.[4] |
Cyanobacteria[]
Genus | Species | Location | Material | Notes | |
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Girvanella[5] |
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Crypt laminites |
A Cyanobacteria, member of the family Oscillatoriales. The only difference compared to the Paleozoic forms is the much thinner celular wall of the specimens described, which otherwise seems to be always thicker, and the more irregular shape of the individual tubes. Girvanella is almost rock-forming in the Lower and Upper levels, and is very common, but can only rarely be detected in the bituminous clay marl slate due to conservation reasons.[5] The Girvanella specimens recovered on the Posidonienschiefer can come from Diatomeas rather than Cyanobacteria, but its assignation is rather controversial.[5] |
Rhizaria[]
Foraminifera[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Shells |
A benthonic Foraminiferan, member of inside the family (). An extant genus. Its shell resemble a mixture between an ammonite conch, due to having a lower spiral, and a mussel. |
![]() Drawing of an Astacolus shell | |
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Shells |
A benthonic Foraminiferan, member of inside the family (). Its conch has a Myriapod-like segmented built. |
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Shells |
A benthonic Foraminiferan, member of inside the family (). |
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Shells |
A benthonic Foraminiferan, type member of inside the family (). |
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Shells |
A benthonic Foraminiferan, member of inside the family (). |
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Shells |
A benthonic Foraminiferan, member of inside the family (). |
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Shells |
A benthonic Foraminiferan, member of inside the family (). |
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Shells |
A benthonic Foraminiferan, type member of inside the family (). Round-spiral shell morphology |
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Shells |
A benthonic Foraminiferan, member of inside the family Nodosariacea (). Dentalina is an extant genus, with an elongated shell, that resemble a small worm. |
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Shells |
A benthonic Foraminiferan, member of inside the family Nodosariacea (). |
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Shells |
A benthonic Foraminiferan, type member of inside the family . Another genus with a Myriapod-like segmented built. |
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Shells |
A benthonic Foraminiferan, type member of inside the family (). Dentalina is an extant genus, with an elongated shell, that resemble a small worm. |
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Shells |
A benthonic Foraminiferan, member of inside the family Robertinida. It resembles a small ammonite. |
Dinoflagellata[]
Dinoflagellate cysts[]
The evolutionary burst of the Toarcian Dinoflajellates led the first appearance and rapid radiation of the (, , , and related forms).[8] This occurred at the time of a widespread Lower Toarcian bituminous anoxia-derived shale of the Posidonienschiefer Formation. Is recovered on the Posidonienschiefer, Pozzale, Italy, Asturias, Spain, Bornholm, Denmark, the Lusitanian Basin of Portugal, the in Yorkshire and to the "Schistes Carton" in northern France. Whether there is a causal connection in this co-occurrence of and bituminous facies is a problem still to be resolved. This family has its acme in diversity and quantity in the latest Toarcian and became less important in the Aalenian.[8]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Cysts |
A Dinoflagellate cyst, member of Dinophyceae of the family . On the Lias Epsylon Interval (Lowermost Toarcian), most of the assemblages are dominated by Nannoceratopsis gracilis. Nannoceratopsis senex becomes highly abundant until the uppermost Tenuicostatum.[9] |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . An Ectoparasitic dinoflagellate, whose hosts are normally Tunicates |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . |
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Cysts |
A Dinoflagellate cyst from the family . The specific epithet, fenestrata, refers to the openings, or fenestrae, in the periphragm. |
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Cysts |
A Dinoflagellate cyst, type member of . Dominant genera on some layers of the Lias Delta Stage.[9] |
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Cysts |
A Dinoflagellate cyst, member of . Commonly found along the genus Beaumontella.[10] |
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Cysts |
A Dinoflagellate cyst, type member of . Luehndea spinosa is common on the medium layers of the lower Sachrang Formation, while restricted to some areas on the Lias delta.[9] |
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Cysts |
A Dinoflagellate cyst, member of . Common on Pliensbachian levels, become present but rare on lower Toarcian.[10] |
Algae[]
Includes abundant variety of algae, such as the genus of colonial Green algae Botryococcus,[16] or the unicellular algal bodies , and other small examples. Algae are a good reference for changes on the oxygen conditions along the Toarcian.[17]
Algae Acritarchs[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Millions of specimens |
An Acritarch probably from Algal origin. Veryhachium fossils represent open marine and transgressive conditions. It has high presence on most of the samples studied from the Sachrang Formation, being nearly the 50% of the Acritarch fraction on some locations. |
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Millions of specimens |
An Acritarch probably from Algal origin. Its fossils indicate nearshore or estuarine to shallow lagoon and/or slightly brackish-water environments. It is the dominant on the nearshore sections. |
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Millions of specimens |
An Acritarch probably from Algal origin. Related to estuarine deposits. |
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Millions of specimens |
An Acritarch probably from Algal origin. Related to open shelf deposits |
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Millions of specimens |
An Acritarch probably from Algal origin. Related to open shelf deposits |
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Millions of specimens |
An Acritarch probably from Algal origin. Related to open shelf deposits |
Haptophyta[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Millions of specimens |
A member of the family inside . Shore deposits genus. The abundance drop of M. jansae further characterise the T-OAE perturbation, where becomes the dominant Genus on most of the Saxony Basin. |
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Millions of specimens |
Type member of the family inside . |
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Millions of specimens |
Type member of the family inside . Towards the Pliensbachian-Toarcian extincion this genus gets a decrease in abundance and size that shows the change and biotic crisis. |
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Millions of specimens |
Type member of the family inside . |
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Millions of specimens |
Type member of the family inside . |
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Millions of specimens |
Type member of the family inside . |
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Millions of specimens |
Type member of the family inside . |
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Millions of specimens |
A member of the family inside . |
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Millions of specimens |
A member of the family inside . |
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Millions of specimens |
A member of the family inside . |
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Millions of specimens |
A member of the family inside . |
Chlorophyta[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Millions of specimens |
A member of Dinophyceae. |
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Millions of specimens |
A member of Peridiniaceae inside Dinophyceae. |
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Millions of specimens |
A member of inside Dinophyceae. |
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Millions of specimens |
A member of Prasinophyceae. It the main genus present on silt and sand horizons, trending to be absent on black argillaceous layers. |
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Millions of specimens |
A member of Prasinophyceae. A genus common on green clays and other upper strata on the formation. |
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Millions of specimens |
A member of Prasinophyceae. A genus common on green clays and other upper strata on the formation. |
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Millions of specimens |
A member of Prasinophyceae. A genus common on green clays and other upper strata on the formation. |
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Millions of specimens |
A member of the Prasinophyceae. Basinal deposits genus |
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Millions of specimens |
A member of the Prasinophyceae. Basinal deposits genus |
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Millions of specimens |
A member of the Prasinophyceae. Basinal deposits genus |
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Millions of specimens |
A member of the Prasinophyceae. Basinal deposits genus |
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Millions of specimens |
A member of the Prasinophyceae. Basinal deposits genus |
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Millions of specimens |
A member of the family Pyramimonadales inside Prasinophyceae. Basinal deposits genus |
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Millions of specimens |
A member of the family inside Chlorodendrales. Basinal deposits genus |
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Botryococcus[18] |
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Millions of specimens |
Type member of the family Botryococcaceae inside Trebouxiales. Freshwater or Deltaic Genus |
![]() Modern Botryococcus |
Fungi[]
Fungal Spores, hypae and undeterminated remains are a rare element of the otherwise openmarine deposits of the Posidonienschiefer formation, but where recovered at Dormettingen.[25] This fungal remains are composed mostly by indeterminate spores and indicate oxygenated environments and suitable transportation by rivers.[25]
Plantae[]
![](http://upload.wikimedia.org/wikipedia/commons/thumb/8/80/Phragmites_australis_kz01.jpg/205px-Phragmites_australis_kz01.jpg)
The macroflora of the Posidonia slate can be described as extremely poor in species.[26] Apart from the remains of Horsetails, it is without exception the remains of coarse branches and fronds from gymnosperms, in which one has a certain can assume transport resistance. Remains of Ferns are completely missing, except for tall arboreal ferns (Peltaspermales).[27] Mostly of the flora was reported from the area of Braunschweig.[26] The major explanation for the flora could be that the plants in question are mono-or oligotypic stands on the edge of the waters that flow into the Posidonienschiefer sea, probably tear away in the course of flood events, easily fragmented during transport and wave waves, possibly especially in the occasional storm events postulated.[28] In terms of taphonomy, this would result in a comparison with today's reed Phragmites, which can form extensive stocks on the edge of shallower and slowly flowing waters ("Reed belts").[26] The Wood remnants clearly indicate one higher diversity of Coniferous flora in the delivery area than the remains of leafy branches.[26] This fact is likely to be proportionate, similar to that frequent occurrence of charcoalized or gagged trunks, mostly of them are believed to be "driftwoods" that only take a long time drifting also suggests a frequent settlement with mussels and full-grown Sea Lilies.[26][28] The deposition settings are at large distance from the nearest coastline (for southern Germany about 100 kilometers), making only plants strong to transportation able to resist enough to get deposited.[29][30] At Irlbach and Kheleim, NE of Regensburg, where the Posidonienschiefer has its near mainland deposit with abundant sand, a rich deposit filled with plant remains of different kind (Seds, Reproductive organs, Leafs, Stems, Cuticles and wood) with traces of coal was recovered, however, it was never studied in depth.[31] Of all the plant material expected only a few Bennetites leafs and two conifer branches with leaves where cited and none studied.[31] At the Austrian realm The sachrang Member was developed in the basinal area, while the Unken Member, sandwiched between red, often condensed limestones, represents the marginal facies.[20] Due to be more marginal and connected with the southern Vindelician land, the most diverse palynological assemblages of the formation are found, transported from zonas with moldanuvian granites as proven by the feldspar accumulations.[20]
Phytoclasts[]
Phytoclasts have been recovered from several sections on the formation, but only studied in depth from the Dotternhausen and specially Dormettingen.[25] Here two kinds of Phytoclasts where recovered, opaque phytoclasts (charcoal, indicator of wildfire activity on nearby landmasses, indicator of seasonal alterations of the water column) and translucent phytoclasts (indicator of proximal landmasses with high aviability of wood and other plant material, as well transport conditions).[25] On the lowermost part of the section opaque phytoclasts are low (15% of the total organic matter) while tranlucent are incredibily abundant (40%), lowering its abundance to a 20-10% on the next section.[25] The Exaratum Subzone is the only one with an inverse trend and more abundance of opaque phytoclasts. On the Bifrons level, both types reach between a 15% and a 30%, showing a rapid increase, to decrease on the end of the section to values of less than a 10%.[25] Opaque Phytoclasts, for a supposed marine deposit are relatively abundant on some sections, whose decreasing on others suggest (along with increasing levels of Kaolinite) an increased delivery of land plant material by rivers, from areas with wetter climate and less frequent fires, while its rise suggest the opposite, nearby continental setting with dry climate and continuous wildfire activity.[25]
Palynology[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Spores |
Affinities with Pteridopsida. Spores from several types of ferns, relatively rare, present only on 2 samples. |
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Spores |
Affinities with Selaginellaceae and probably Lycopsida. A rare element on the palynological records of the German Basin, although more abundant than any other Spore recovered locally. |
![]() Extant Selaginella, typical example of Selaginellaceae. Genera like probably come from a similar or a related Plant | |
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Spores |
Affinities with the family Osmundaceae inside Polypodiopsida. Near Fluvial currents ferns, reted to the modern Osmunda regalis |
![]() Example of extant Osmunda specimens, come probably from similar genera or maybe a species from the genus | |
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Affinities with the family Cyatheaceae inside Cyatheales. Arboreal Fern Spores |
Example of extant Cyathea, come probably from similar genera | |
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Spores |
Affinities with Dicksoniaceae inside Pteridopsida. Tree Fern Spores |
![]() Modern Dicksonia, come probably from similar genera | |
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Spores |
Affinities with the Ophioglossaceae inside Filicopsida. Spores related with modern floor Ferns, that appear on abundant water locations. The Unken Member is considered a more basinal deposit, where Wood and Sporomorph remains are more common. |
![]() Ophioglossum, come probably from similar genera | |
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Spores |
Affinities with Gleicheniaceae inside Gleicheniales. Suggest relative increase of humidity on the rivers flowing towards the Austrian realm. Most abundant Fern spore in this region. |
![]() Dicranopteris, come probably from similar genera | |
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Pollen |
Affinities with the families Peltaspermaceae, Corystospermaceae or Umkomasiaceae inside Peltaspermales. Pollen of Uncertain provenance, that can be derived from any of the members of the Peltaspermales. The lack o distinctive characters and bad conservation are among the main factors to make this Palynological residues difficult to classify. Arboreal to arbustive seed ferns. |
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Pollen |
Affinities with Gnetopsida and probably Gnetophyta. Has Been considered Pollen of Chloranthaceae. However, it is to old for belonging to advanced Angiosperms. It probably comes from cones related to the Genera kuesperti from the Lowermost Jurassic of Germany, resembling pollen of extant Ephedra and Welwitschia. |
![]() Closer Look of Ephedra cones, a common Gnetal. maybe come from a related plant | |
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Pollen |
Affinities with the Cycadopsida inside Cycadales. Pollen related with modern Cycas, arbustive to lower floor plants, relatively abundant, present on various of the measured samples. The mos common found on the Austrian realm, indicator of dry settings. |
![]() Encephalartos, come probably from similar genera | |
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Pollen |
Affinities with Cycadaceae and probably Cycadales. Alternatively can be Pollen from Bennettitales. It is the most abundant non conifer Pollen recovered on the formation, recovered on all the major sampled areas. Probably derived from arbustive cycads, this genus is related with dry settings, even from desertic regions. |
![]() Closer Look of Macrozamia cones, a common Cycad. maybe come from a related plant | |
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Pollen |
Affinities with Pinaceae inside Coniferae. |
![]() Extant Pinus Cone, example of the Pinidae. is similar to the pollen found on this genus | |
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Pollen |
Affinities with the Pinidae inside Coniferae. Abundant on the Lower Jurassic of NW Europe. Its identification on the Posidonienschiefer is rather complex due to the bad preservation of the Pollen Grains. |
![]() Extant Pinus cembra Cone, example of the Pinidae. is similar to the pollen found on this genus | |
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Pollen |
Affinities with Abietoideae inside Coniferae. Pollen From arbustive to arboreal plants, the second most abundant conifer pollen recovered, resembling the pollen of the modern genus Tsuga (Section Micropeuce).[36] The differences observed between Cerebropollenites and Tsuga are no greater than the differences observed between the pollen of the two Sections of Tsuga, Hesperopeuce and Micropeuce.[36] The greater difference with the modern Tsuga is the relation of this genus with an arid/ecuatorial range, as this genus is indicative of dry, arid and even desertic conditions, compared with the cold locations where this genus is found today. Probably has a similar role to the modern species Pinus pinea.[36] |
![]() Extant Tsuga Cone, example of the Abietoideae. is similar to the pollen found on this genus | |
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Pollen |
Affinities with Cheirolepidiaceae and Araucariaceae inside Pinaceae. Non concreted affinities |
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Pollen |
Affinities with Cheirolepidiaceae inside Pinaceae. Abundant on the Lower Jurassic of NW Europe. co-occurs on the coeval Sorthat Formation with cuticles of ramonensis, and after a test of relationships it was found a highly significant correlation that may suggest that the species S. psilatus was produced by the conifer genus Dactyletrophyllum.[37] |
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Pollen |
Affinities with the Cheirolepidiaceae inside Pinales. Pollen from Arboreal to Arbustive Plants. It is rare on the Samples measured. |
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Pollen |
Affinities with Cheirolepidiaceae inside Coniferae. Abundant on the Lower Jurassic of North and Southern Europe, represents pollen of medium to large arboreal plants, specially coniferales. The abundance of pollen of Classopollis and other thermophile plants was observed in this region in the lower Toarcian from the end of the antiquum (= tenuicostatum) zone to the middle of commune zone.[38] Classopollis is correlated with evaporites and are therefore associated with desert basins, but the shrubs may have also lived in xeric upland areas with seasonal fires. Evidence of fires is absent on the marine Posidonienschiefer, but has been recovered on the coeval nearshore calcareous sandstones.[38] It increases with the appearance of charcoal phytoclasts, as derived from dry settings with increased wilfires.[25] |
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Pollen |
Affinities with Cheirolepidiaceae inside Coniferae. Pollen of medium to large arboreal plants, specially coniferales. |
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Pollen |
Affinities with the family Cupressaceae inside Pinopsida. Pollen that resembles extant genera such as the Genus Actinostrobus and Austrocedrus, probably derived from Dry environments. |
![]() Extant Austrocedrus. maybe come from a related plant | |
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Pollen |
Affinities with Podocarpaceae inside Pinopsida. Pollen From arbustive to arboreal plants |
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Equisetaceae[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Stems |
Affinities with Equisetaceae inside Equisetopsida. Number of mostly very fragmented and not particularly well preserved, but clear horsetail remains described. So far recognizable, leaf sheaths where developed in most cases, but the state of preservation does not allow a more precise determination.[40] |
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Stems and incomplete axes |
Affinities with Equisetaceae inside Equisetopsida. Neocalamites is the most common more distributed of all the Posidonia Shale, being even found on Luxembourg Posidonia Strata.[41] Mostly of the Stems reported come from Aeolian-Dunar related deposits, or from nearshore-basinal deposition. Probably was related to the seashore.[41] Some stems are big, resembling the rates of growth seen on modern Bamboo specimens, suggesting +6–7 m tall Equisetopsids.[41] |
Neocalamites merianii specimen | |
?[42] |
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Pollen Cones |
Affinities with Calamitaceae inside Equisetopsida. It was an approximately 2–3 m high shrub-like plant related to swamp environments. Calamitaceans are common in late Palaeozoic wetland plant communities, so the find of a jurassic specimen is rare.[42] Maybe is a mistaken new genera of Pollen cone, but definitely come from an Equisetalean. This genus is found associated with Annularia, being both part of the Calamites plant. It can be related with the Equisetites stems found on the formation. |
![]() drawing of a specimen |
Pteridospermatophyta[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Affinities with Umkomasiaceae inside Corystospermaceae. Is based on bipinnate leaves, rachis longitudinally striated, with a long petiole and secondary rachises. It belongs to large tree ferns. Sachrang Formation Specimen is characterized for its large size and probably where attached to trunks similar in built to the Cretaceous genus Tempskya.[43] |
![]() The only Pachypteris specimen know from the Sachrang Formation |
Bennettitales[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Affinities with Cycadeoidaceae inside Bennettitales. It is the most abundant medium-sized plant on the environment. Found specially on seashore depositional settings, but also on deltaic and lagoonar environments. It was a low arbustive-arboreal Bennetite related to arid environments, with a leave similar of that of the modern genus Encephalartos, specially Encephalartos munchii, but also Dioon mejiae. Otozamites has been considered synonym with , but since the 1990s everybody used the name Otozamites, and was forgotten.[46] |
![]() Otozamites gracilis specimen from the Sachrang Formation | |
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Affinities with Cycadeoidaceae inside Bennettitales. This Bennettitalean is related to Shrub built. Some specimens were assigned to Dioonites acutifolium (Junior synonym). Consists on leaves with pinnate, leaflets perpendicular or oblique to the rhachis, on the top of the leaf axis. It comes from fragments of fairly large fronds. In its external form it closely follows the modern genus Dioon, only the leaflets are of our kind wider and shorter, they are further apart, but the tendency to rhachis is the same. |
Pterophyllum fossil | |
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Affinities with Williamsoniaceae inside Bennettitales. Arboreal Cycadaceans, some with the presence of flower-like structures. It resembles the leaf of the modern Microcycas calocoma, and probably had a similar arboreal built, being the leave of tall Bennetite trees such as , found on the middle jurassic of England. |
![]() Ptilophyllum sp. specimen from the Sachrang Formation | |
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A member of Williamsoniaceae inside Bennettitales. Identified originally as Zamites oblongifolius |
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A member of Williamsoniaceae inside Bennettitales. It has been interpreted as a cycad in the family Cycadaceae or a Bennettitalean plant, and also a late surviving member of Noeggerathiales. Leaflets somewhat removed, oval-oblong, a little narrower near the base, rounded at the tip, nerves partially diverging from the base towards the edge. It was assigned to Pterophyllum oblongifolium and on the genus Glossozamites. This genus was the leaf of arboreal Bennetites, similar in appearance to the modern Encephalartos woodii. |
![]() Zamites mandelslohi specimen |
Ginkgoales[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Affinities with Ginkgoaceae inside Ginkgoales. Various leaves are known from Ohmden, coming from marine deposits and where identified as Ginko digitata, identified originally as the only Ginko specimen reported on the Sachrang Formation. In the southern and northern Germany, there are regular remains of coal, which are initially reminiscent of small Ginkgo leaves. The leaves are hard to identify, more or less regularly concentric structures, as they sometimes appear like the coarse fruiting bodies of wood-dwelling fungi, such as the genus Trametes. |
![]() Ginkgoites digitata specimens | |
Sphenobaiera[30] |
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Affinities with Karkeniaceae inside Ginkgoales. |
Pinophyta[]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. Represent various kinds of cones from diverse conifer origin. |
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Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. Pagiophyllum araucarinum predominates among the two types of leafy coniferous branches that have become known from the Posidonia. However, there is no indication whether this fact reflects their respective share in the vegetation of the delivery area. Cheirolepidiaceae Pollen is the most abundant and diverse found on the formation, what is correlated with the abundance of this genus. Other factor that puts local Pagiophyllum on Cheirolepidiaceae is the dominance of an arid climate, the preferred for this type of conifers. |
![]() Pagiophyllum kurri specimen from Banz | |
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Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales. Specimens whose spiral foliage of the branch, in which the individual leaves open about 2/3 of their length (without the tip) are fused with the branch they hold in their free part fits tightly, suggests belonging to the genus Brachyphyllum. |
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Hirmeriella[26] |
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Ovuliferous dwarf-shoots |
Affinities with Cheirolepidiaceae. Is the type genus of this family, and is related with Arid Settings. Coming from The land-plant-taphocoenose from posidonia slate 3, is ready for processing. |
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A possible ancestral member of the Callitroideae inside Cupressaceae, or a member of Cheirolepidiaceae. Named also "" liasinus, represents probably Arbustive to arboreal-derived axis. |
liasinus specimen from the Sachrang Formation |
Fossil wood[]
Fossil Wood increases on the marginal Unken Member, with great amounts of logs and fragments of more than 1 m. Surface studies suggest relationships with the wood genera identified on the coeval Úrkút Manganese Ore Formation.[52]
Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
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Fossil wood |
Affinities with the Cupressaceae inside Pinales. Probably related to the coastal settings. It maybe covered a similar role that modern Swamp Cypress, Taxodium distichum. |
![]() Taxodium distichum specimen, whose role was probably covered by the makers of the Taxodioxylon woods. | |
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Fossil wood |
Affinities with Cupressaceae inside Pinales. Wood with a morphology similar to the modern genus Metasequoia. Probably related to Widdringtonites liasinus. |
![]() Metasequoia specimen, whose role was probably covered by the makers of the Cupressinoxylon woods. | |
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Fossil wood |
Affinities with Podocarpaceae inside Pinales. Acmopyle pancheri is the only living conifer that has real resemblance with the wood assigned to . Xenoxylon phyllocladoides and C. grandiporosum are commonly found associated, for example in Braunschweig area. |
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Fossil Wood |
Affinities with Podocarpaceae inside Pinales. resembling modern genera such as Dacrycarpus, with other specimens resembling Juniperus. Includes wood more related to nearshore arbustive Conifers (columnar or low-spreading Shrubs with long, trailing branches), being the most abundant, but also medium to large arboreal conifers from nearshore forests. |
specimen | |
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Fossil wood |
Affinities with Podocarpaceae inside Pinales. Phyllocladus is the extant genus of conifer whose wood is more similar. |
![]() Phyllocladus specimen, whose role was probably covered by the makers of the Phyllocladoxylon woods. | |
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Fossil wood |
Affinities with Podocarpaceae and Cupressaceae inside Pinales. The assignation of this genus is rather complex. |
![]() Bivalves added to a driftwood | |
Agathoxylon[56] |
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Fossil wood. |
Affinities with Araucariaceae inside Pinales. Has araucarioid structures, but lacks proper characters of extant genera such as Araucaria & Agathis The largest known rafting wood on the fossil record is assigned to this genus, with a length of 18 m. The rafts were populated with Crinoid colonies, and a wide variety of organisms.[56] |
![]() Araucarioxylon (=Agathoxylon) reconstruction |
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Fossil wood |
Affinities with Abietinae inside Pinales. This genus was found to be related with found on the coeval Whitby Mudstone, crucial to the interpretation of the fossil record of Early Abietinae. |
![]() Abies specimen, whose role was probably covered by the makers of the Araucariopytis woods. | |
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Fossil wood |
Affinities with Cheirolepidiaceae inside Pinales. Includes large sized trunks up to 1.7 m tall and 115 cm wide. Wood of other species of this genus, such as Protocupressinoxylon purbeckensis grown on flood-influenced settings of vegetated soil with brackish-water, like modern Swamp Cypress. |
catenatum specimen | |
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Fossil wood |
Affinities with Cheirolepidiaceae inside Pinales. |
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Fossil wood |
Wood of the Protopinaceae, a possible "morpho-group" of the family Cheirolepidiaceae. Belongs to an invalid group of mostly Paleozoic Woods, and all the woods assigned to the family should be threated[clarification needed] as Incertade Sedis.[64] |
![]() Woodworthia, example of Protopinaceae | |
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Fossil wood |
Affinities with Coniferales, concretely is closer to the Podocarpaceae, Cupressaceae and in a lesser extend to the Cheirolepidiaceae. Finally can be a member of the extinct family . It is the more abundant genus of wood present on the Bohemian Realm of the Sachrang Formation. |
Ichnofossils[]
The major ichnological analyses of the Posidonian Shale come from Dotternhausen/Dormettingen, where the ichnogenus formed the so-called Tafelfleins and Seegrasschiefer.[68] The Tafelflein bed was deposited under anoxic bottom and pore water, where a recover of oxygen allow the Phymatoderma-producers return.[68] The two organic-rich layers (Tafelfleins and Seegrasschiefer) are characterized by the dense occurrence of trace fossils such as Chondrites and , done episodically due to the fall of the oxygen levels.[68] The Coeval more nearshore Swiss deposits referred Posidonian Shale () hosted similar trace fossils asthose recovered on SW Germany.[68] Tougth this setting apparently evolved faster to more oxic-to-dysoxic bottom waters.[68] At Unken, laminated deposits of red limestone suggest well oxygenated active waters (as lack shale), where high amounts of Chondrites are found.[20]
Genus | Species | Location | Material | Notes | Images |
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Zoophycos[69] |
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Burrowing and track ichnofossils. |
Burrow-like ichnofossils. It has been related to Echiuran annelids, but also from moving and feeding polychaete worms.[70] |
![]() Example of Zoophycos fossil |
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Burrowing and track ichnofossils |
U-shaped tubes with sinuous, bifurcating or planispiral spreite, that can be related to Crustaceans, Annelids and Fishes, being both Domichnia and/or fodinichnia. |
![]() Specimens | |
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Burrowing and track ichnofossils |
Burrow-like ichnofossils, that can be related to Crustaceans, Annelids and Fishes.[71] The presence of this burrows changed along the different depositional layers, interpreted as result of relative magnitudes and durations of a series oxygenation events.[71] Increased Oxygen conditions eventually led to a level that permitted both the survival of larger Chondrites and Thalassinoides producing organisms, as well the depth of the Burrow-like structures.[71] The changes on the layers are detailed enough to know that oxygenation-change events duration was sufficient to allowe the migration and establishment of trace-producing organisms, establishing an "equilibrium" with bottom-water oxygen conditions.[71] |
![]() Thalassinoides found on the Sachrang Formation | |
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Burrowing and track ichnofossils |
Burrow-like ichnofossils. Interpreted as the feeding burrow of a sediment-ingesting animal.[74] A more recent study has find that Scoloplos armiger and Heteromastus filiformis, occurring in the German Wadden Sea in the lower parts of tidal flats, make burrows that are homonymous with numerous trace fossils of the ichnogenus.[75] Chondrites burrows from Holzmaden are mostly filled with granular Calcite crystals, clay minerals, and rare framboidal Pyrite. The local dysoxic seawater is reflected on the palaeoredox conditions, relating framboidal pyrite and biogenic processes. Chondrites appear to be capable of colonizing environments characterized by oxygen levels well below levels needed for survival of other animals, being considered an "extremotolerant" ichnotaxon.[76] |
Illustration of Chondrites bollensis | |
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Burrowing and track ichnofossils. |
Burrow-like ichnofossils. It consists commonly on a subhorizontal branching burrow system consisting of radiating tunnels filled with fecal pellets.[77] It has been interpreted as a product made by an Endobenthic deposit-feeding animal, specially a Fodinichnia, burrows produced by benthonic subsurface food-mining activity, as is proved by the tunnels and pelletal infill.[77] The study of the Fecal Pellets has revelated that the maker of this ichnogenus was an epicontinental shelf setting non-selective deposit feeder, ingesting particles on the sediment surface without selection. A mode of feeding common on aquatic Benthos, reported on modern animals such as Spionid Polychaete worms, tropical Holothurians and Spatangoid urchins.[78] |
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Burrowing and track ichnofossils. |
Burrow-like ichnofossils. It is controversial, since is considered a strictly a junior synonym of .[80] |
![]() Example of Planolites fossil |
Invertebrata[]
Anthozoa[]
Genus | Species | Location | Material | Notes | Images |
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Polyps |
A stone cora of the family Caryophylliidae inside Hexacorallia. Related to shallow waters, this genus is the main coral found on the Sachrang Formation, resembling the modern Polycyathus muellerae. Its fossils are related with near-land facies, Coralline Islands and relatively small landmases shuch as the Bohemian Massif. |
![]() Thecocyathus mitrae from Banz |
Porifera[]
In the non-bituminous facies located on Obereggenen im Breisgau (Shore of the Black Forest High), especially the lower semicelatum subzone, pyritized individual needles of silica sponges (Demospongiae and Hexactinellida) are found, rarely on pelagic layers to very often on the low depth marine deposits.[5] They are usually associated with radiolarian stone cores. In Dusslingen and Reutlingen, these sponge needles could be barytized in phosphorites of the Haskerense subzone and are much more common here than in any other zone of the Lower Toarcian. These needles are absent in the bituminous horizons of the entire Lower Toarcian.[5] Increased amounts of Sponge needles (dominated by Hexactinellida) are also found on the arenaceous facies of the nearshore unit that is the Unken member, being the only section if its region hosts them, probably due to be an active and well oxygenated bottom.[20] The location of this member as a possible bay on the south of the vindelician land probably allow to the development of more pre-Toarcian AOE conditions, hence the presence of biota otherwise rare on bituminous layers.[20]
Annelida[]
Genus | Species | Location | Material | Notes | Images |
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Multiple Specimens. |
A sessile, marine annelid tube worm of the family Serpulidae. Its affinities with the genus Serpula are controversial, since the genus is known mostly since Cretaceus strata. Although there are other fossils assigned to the genus on same age deposits of France.[83] Presumably these specimens have fallen from their growth areas.[5] |
![]() Head of a modern Serpula vermicularis | |
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A polychaete of the family Dorvilleidae inside Eunicida. It was considered as part of the genus . Eunicidan species with prionognath jaws, absent on Bituminous layers |
Mollusca[]
Brachiopoda[]
Genus | Species | Location | Material | Notes | Images |
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Shells |
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Shells |
A Discinidae rhynchonellatan. This genus was found had a planktotrophic larval stage, that adapted while growing to the local redox boundary, when this fluctuated near the sediment–water interface and oxygen availability prevailed, allowing benthic colonization. Is found on associations with and .[84] |
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Lingula[57] |
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Shells |
A Lingulidae rhynchonellatan. Associations of bioturbation infauna are dominated on certain sections by /Lingula agrupations, developed under longer-term oxygenated conditions within the substrate and bottom waters.[84] |
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Rhynchonella[57] |
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Shells |
A rhynchonellatan. Found associated with Plicatula on long-term well-oxygenated conditions within the substrate and bottom waters.[84] |
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Shells |
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Shells |
Bivalvia[]
Genus | Species | Location | Material | Notes | Images |
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An scallop. |
![]() Colony of specimens | |
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An scallop. Found mostly on the "Dactylioceras-Monotis-Bank", a deposit derived from large scale tectonic events on the Bohemian coastline |
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An scallop. |
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Chlamys[57] |
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A scallop. |
![]() Single specimen |
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A scallop. The presence of this genus along endo- and epibenthic bivalves, which are absent farther up the section, suggest a delayed overstepping of anoxic bottom waters on the Altdorf High.[90] |
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A scallop. |
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A propeamussiid mud scallop. |
![]() Various specimens on the same rock | |
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A plicatulid mud scallop. |
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20.000 specimens/m2 |
A ostreoidan. It is the type fossil of the Sachrang Formation. Originally it was named "Posidonia bronni", thought to be a new genus, and the strata was denominated the Posidonia layers after it. Years later it turned out to be a junior synonym of Bositra, and thus, it was reassigned. However, the name of the layers was retained. The habitat and mode of life of Bositra has been debated for more than a century. There have been different interpretations, such as a pseudoplanktonic organism,[94] a benthic organism[95] related to open marine floor, where it was the main inhabitant of the basinal settings,[96][97] a free swimming mode of life filtering phytoplankton,[93] and a hybrid mode, where it has a life cycle with holopelagic reproduction controlled by the change on Oxygen levels,[98] and even a chemosymbiotic lifestile, related to the large crinoid rafts, being the main "Safe conduct" to evade anoxic events.[99] All the opinions along the years led to a large study in 1998, where the size/frequency distribution, the density of growth thanks to the lines related to the shell size and the position of the redox boundary by total organic carbon diagrams has revealed that Bositra probably had a benthic mode of life.[100] |
![]() Thousands of specimens in one matrix | |
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A "" ostreoidan. Another Genera mistaken with "Posidonia bronni". |
![]() Various specimens in one matrix | |
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A bakevelliid mud oyster. |
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A gryphaeid mud oyster. |
Various specimens | |
Gryphaea[57] |
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A gryphaeid mud oyster. |
![]() Various specimens |
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A gryphaeid mud oyster. |
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Pinna[57] |
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A Pinnoid oyster. |
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A Pteriidaeoid wing-oyster. |
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A Nuculid nut clam |
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A Pholadomyid clam |
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An Astartid clam |
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A clam |
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An inoceramid clam. Being the second most common genera of Bivalve on the Formation, it had been object to several studies to find its ecological niche, like Bositra. Several opinions include a pseudoplanktonic-only organism, able to live in open sea,[97] or a benthonic-only organism.[96] On the 1998 evaluation with Bositra, was found that probably has a benthic early life that translated to a faculatively pseudoplanktonic mode of adult life.[100] |
![]() Single specimen | |
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An inoceramid clam. |
Thousands of specimens on a single rock | |
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An inoceramid clam. |
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A Clam, type member of the family Solemyidae inside Solemyida. |
![]() Single specimen | |
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A cucullaeid clam. |
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A lucinid clam. |
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A clam. This Genus had a lecithotrophic and planktotrophic larval development.[84] |
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A clam. |
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A Limidae file clam. |
![]() Plagiostoma giganteum, specimen multiview | |
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A Limidoid file clam. |
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A Limidoid file clam. |
Gastropoda[]
Genus | Species | Location | Material | Notes | Images |
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A sea Snail. Is the oldest known holoplanktonic gastropod, thanks to a bilateral symmetrical shells as an adaption to active swimming. Also the most common of the sea snails of the Formation, it is also one of the most varied in size terms, with some of the biggest specimens of snail from the Lower Toarcian know.[105] It has been related to large floating driftwood as one of the primary settlers.[105] |
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Shells |
A sea Snail. Possible holoplanktonic gastropod.[105] |
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Shells |
A Pleurotomariidae sea Snail. |
![]() Specimen | |
Zygopleura[57] |
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Shells |
A Zygopleuridae sea Snail. |
![]() Specimens |
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Shells |
A Eucyclidae sea Snail. |
Specimens | |
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Shells |
A Eucyclidae sea Snail. |
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Shells |
A Procerithiidae sea Snail. |
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Shells |
A Naticidae moon Snail. |
![]() Extant Specimen | |
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Shells |
A Snail of uncertain placement. |
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A Pterotracheidae sea Slug. Among the oldest pelagic floating Slugs, Pterotrachea liassica had a more extended larval period than modern extant Pterotrachea coronata, because one additional whorl is present.[106] |
Modern Specimen |
Cephalopoda[]
Genus | Species | Location | Material | Notes | Images |
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A Nautilidae Nautilidan. Includes the largest specimen of Cenoceras known, with 80 cm width. Two specimens, identified as Nautilus sp. from Holzmaden where found encrusted with Serpulids and Bryozoans.[110] |
![]() Nautilidae shell from Banz, probably Cenoceras | |
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A Lytoceratidae Ammonite. Lytoceras can get quite big, with nearly 50 cm in diameter. |
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A Lytoceratidae Ammonite. |
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A Hammatoceratidae Ammonite. |
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A Hammatoceratidae Ammonite. |
![]() Hammatoceras | |
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A Hildoceratidae Ammonite. |
![]() Grammoceras specimen from Banz | |
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Shells |
A Hildoceratidae ammonite. The co-occurrence on Altdorf of boreal () and Tethyan faunal elements (Frechiella) is striking, suggesting clear connection with both regions.[114] |
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Shells |
A Hildoceratidae ammonite |
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Neolioceratoides[115] |
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A Hildoceratidae Ammonite. |
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Cleviceras[116] |
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Specimens |
A Hildoceratidae ammonite |
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A Hildoceratidae Ammonite. |
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A Hildoceratidae Ammonite. |
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A Hildoceratidae Ammonite. |
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A Hildoceratidae Ammonite. |
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A Hildoceratidae Ammonite. |
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A Hildoceratidae Ammonite. |
![]() Harpoceras specimen | |
Tiltoniceras[113] |
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A Hildoceratidae Ammonite. |
Tiltoniceras specimen |
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A Hildoceratidae Ammonite. |
Hildoceras specimen | ||
Mercaticeras[119] |
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Complete Shells |
A Hildoceratidae Ammonite. |
![]() Mercaticeras specimen |
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Shells |
A Hildoceratidae ammonite |
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A Hildoceratidae Ammonite. |
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A Hildoceratidae Ammonite. |
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Shells |
A Dactylioceratinae ammonite, inside Eoderoceratoidea. |
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A Dactylioceratidae Ammonite. |
![]() Dactyliocerascommune on Holzmaden | |
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A Dactylioceratidae Ammonite. |
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A Dactylioceratidae Ammonite. Is common on the bituminous marls (incorrectly designated as "Wilder Schiefer") of the Altdorf High. |
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A Dactylioceratidae Ammonite. |
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A Dactylioceratidae Ammonite. |
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Type Coeloceratidae Ammonite. |
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A Phylloceratidae Ammonite. The largest ammonite found in the Posidonienschiefer comes from the Ohmden quarry,and belongs to a Phylloceras heterophyllum with a diameter of 87 cm.[119] |
![]() Phylloceras restoration | |
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A Ammonite. |
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A Ammonite. |
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A Ammonite. |
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A Ammonite. |
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A Belemnite. Chitinobelus rostrum was composed of aragonite with organic material, while normal Belemnites had calcite. Has been suggested this rostrum was calcitic. |
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A Belemnite. |
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A Belemnite. |
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A Belemnite. Includes some of the Biggest Know Belemnites, with an estimated maximum up to 4.5 m long in life, although, most specimens would have been rather smaller. |
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A Belemnite. |
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A Belemnite. Includes really large specimens |
![]() Youngibelus Reconstruction | |
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A Belemnite. |
![]() Passaloteuthis Holzmaden specimen | |
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A Belemnite. |
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A Belemnite. |
![]() Salpingoteuthis specimen | |
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Incertade sedis Belemnites. |
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A (Vampyromorpha).[129] The Loligosepiidae is believed to be ancestral to the Recent vampire squid, Vampyroteuthis infernalis.[130] Hooklets in the food residues in the posterior mantle indicate that Loligosepia preyed upon belemnites.[128] |
Holzmaden specimen | |
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A (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. Gladii of Loligosepia can be distinguished from Jeletzkyteuthis by the transition lateral field/hyperbolar zone. A specimen of this genus found on Ohmden has appeared predating a .[132] The association of this 2 genera shows the predatory behaviour of this group when lived on Epicontinental seas, being rather different than extant Vampyromorphs.[132] |
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A (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. Gladius with weakly arcuated hyperbolar zones. |
![]() specimen | |
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A (Vampyromorpha). Related to the modern Vampyroteuthis infernalis. It is distinguished from Geoteuthis and Loligosepia by its median rib: this rib forms a narrow ridge between two narrow grooves. Probably hosted finds similar to modern Vampyroteuthis.[132] |
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A Plesioteuthididae (Vampyromorpha). was originally described as "Geoteuthis" sagittata. |
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Type member of the inside Vampyromorphida. The taxonomic position of Lioteuthis is uncertain, although the wings reaching the proximal gladius section and the smooth median field suggest affinity to the [135] |
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A Teudopseinae (Vampyromorphida). |
![]() Teudopsis Ohmden specimen | |
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A Coleoidean. It has been found adult individuals of Clarkeiteuthis which caught small teleost fish of the species Leptolepis bronni. Further indirect evidence for the hunting behaviour comes from their body orientation in the water during life.[138] |
Holzmaden specimen | |
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A Coleoidean. Has been confused with Acrocoelites tripartitus, hence the species name. |
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A Coleoidean. Some specimens belong to Clarkeiteuthis (=Phragmoteuthis) conocauda, but others are clearly different. |
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A Coleoidean. Sueviteuthis had at least six arms with rather simple hooks, similar to the present of the genus Phragmoteuthis. |
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A possible primigenial Cuttlefish. Is one of the most important fossils of Cephalopods on the Sachrang Formation, due to be one of the Earliest examples of Pigments found on any species, also one of the first historically.[142] The pigments are preserved on various specimens with Eumelanin related to its ink sacs and include even phosphatized musculature.[143] |
Arthropoda[]
Cycloidea[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
Juracyclus[144] |
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|
Incomplete carapace |
The First Cycloid Arthropod from the Jurassic, from the family inside Cycloidea.[144] Cycloids are a group of maxillopod arthropods that span between the Paleozoic until the latest Cretaceous, probably related to the crustaceans and probably detritivores.[144] |
Isopoda[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
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|
Single posterior portion |
An Isopod, member of the family , probably related with Cymothoida. Reported as being from the Late Toarcian Jurensismergel Formation, was recovered from a Deeper shelf environment composed by bituminous limestones, found only locally on the Posidonienschiefer, concretely the upper Bifrons substage. Middle Jurassic representatives of the genus have mouthparts that matches with extant parasitic genera.[146] |
Ostracoda[]
Genus | Species | Location | Material | Notes | Images |
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Cunchs |
A marine Ostracodan, incertade sedis inside Podocopida. |
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Cunchs |
A Marine Ostracodan of the family Protostomia. The specimens of this genus are rather fargmentary and of uncertain nature. |
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Cunchs |
A marine Ostracodan, member of the family inside Podocopida. Rather abundant on the Toarcian profiles on Europe, this genus has a Mussel-like shape, with a very clean and round morphology. |
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Cunchs |
A marine Ostracodan, member of the family Macrocyprididae inside Podocopida. |
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Cunchs |
A marine Ostracodan, member of the family Pontocyprididae inside Podocopida. |
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Cunchs |
A Marine Ostracodan of the family inside Podocopida. This genus is the main reported on the marine facies of the Dobbertin Clay Pit. |
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Cunchs |
A Marine Ostracodan of the family inside Podocopida. This genus is the main reported on the marine facies of the Dobbertin Clay Pit. |
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Cunchs |
A Marine Ostracodan of the family inside Podocopida. The genus is rare on the layers. |
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Cunchs |
A Marine Ostracodan of the family Protocytheridae inside Podocopida. A genus related with Fish fossils and anoxic bottoms. |
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Cunchs |
A Marine Ostracodan of the family inside Podocopida. |
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Cunchs |
A Marine Ostracodan of the family Cytheruridae inside Podocopida. Is rare and the specimens found are rather incomplete. |
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Cunchs |
A Marine Ostracodan of the family Polycopidae inside Cladocopina. Scarce but well preserved specimens. |
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Cunchs |
A Marine Ostracodan of the family Bythocytheridae inside Cladocopina. |
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Cunchs |
A Marine Ostracodan of the family Cytherellidae inside Platycopida. |
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Cunchs |
A Marine Ostracodan of the family Cytherellidae inside Platycopida. |
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Cunchs |
A Marine Ostracodan of the family Bairdiidae inside . Abundant and diverse, is found associated with Ammonite shells. |
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Cunchs |
A Marine Ostracodan of the family Bairdiidae inside . |
Malacostraca[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
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|
Single Specimen inside an Ammonite Shell. |
An hermit crab of the family Paguridae. This specimens where found inside an Ammonite shell, probably looking to evade anoxic conditions or predators. |
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Mecochirus[145] |
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Complete Specimens |
A Decapodan of the family Mecochiridae. |
|
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|
Isolated Chelae |
A Decapodan of the family Glypheidae. |
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[156][157] |
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|
An Astacidea Decapodan of the family . Reaching large sizes of almost half a meter (39–47 cm), Uncina Posidoniae is among the largest known Jurassic Crustaceans. Uncina posidoniae is also the largest representative of the genus Uncina.[157] This large crustacean has been found associated with Ammonite and Bivalve filled Bentos, where probably hunted different kinds of prey.[157] Its large claws would have been perfect to hunt small invertebrates and vertebrates.[157] |
![]() posidoniae specimen The Largest complete posidoniae specimen, with 44 cm long. |
|
|
Associated Partial & Complete Specimens |
A gregarious Polychelidan Lobster. Specimens of Tonneleryon schweigerti where recovered generally in cluster of several individuals, a characteristic unique to this species on the whole Polychelida group.[158] The specimens in these accumulations are of similar size, lacking characters of exuviae such as a split median line or disjunction of carapace and first pleonite. Due to that and the disposition of the specimens probably represent a mass-mortality assemblage and suggest this species was gregarious.[158] |
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A Coleiidae Decapodan. Was confussed with Proeryon hartmanni specimens. Although these finds point to some potential for exceptional preservation, the Pfronten locality was never systematically sampled.[159] It was assigned to the genus Gabaleryon the basis of its ovoid carapace, a transverse groove cutting the axial carina on pleonal terga and an uropodal exopod with curved diaeresis.[159] It shares similarities with Gabaleryon garassinoi and Gabaleryon moorei.[159] A species from Gomaringen is the first know with preserved ommatidia.[160] |
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|
A Coleiidae Decapodan. The second largest Decapodan from the formation, P. giganteus is a species reaching a larger size than most other polychelidans, with up to 15 cm. On the Sachrang Formation there is the most abundant variety of species from the genus, ranging from different sizes and morphologies, that indicate different habitat & feeding adaptations on the genus. Some like P. hartmani show less adaptations to hunt for small nectobenthic preys than other relatives, being abundant on Oyster-filled waters. There is a relatively abundance of the genus in deep-water settings from the Toarcian onward.[166] |
giganteus sp. specimen from Schandelah | ||
|
|
Partial Specimens. |
A Penaeidae Decapodan. |
Antrimpos specimen | |
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|
Partial Specimens. |
A Penaeidae Decapodan. |
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Acanthochirana[168] |
|
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|
An Aegeridae Decapodan. |
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|
Partial Specimens. |
An Erymidae Decapodan. |
specimen | |
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An Erymidae Decapodan. |
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Complete Specimens |
Type genus of the Erymidae Decapodan family. Originally, was named Glyphea amalthei,informally used by Quenstedt and housed on the Museum Naturkunde in Württemberg. A series of posterior revisions probe it was a different genus.[175] |
Eryma specimen | |
|
|
Single Chela |
An Erymidae Decapodan. It was erroneously reported from the Late Toarcian, as was found on bituminous limestone on the Unterer Stein it is clear its provenance from the Sachrang Formation.[174] |
||
Achelata[178] |
|
|
Single complete specimen in late larval stage |
The specimen reported represents the oldest fossil record of an achelatan lobster larva, and the first representative of achelatan lobsters in the Posidonia Shale. Shares similarities with the late Jurassic genus Cancrinos. It is also the first example among crustaceans, which possibly could have lived as part of the plankton.[178] |
California spiny lobsters are relatives of the Larval specimen from the Sachrang Formation |
Palinurina[167] |
|
|
Partial Specimens |
A Spiny Lobster of the family Palinuridae |
|
Stomatopoda[179] |
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|
Single Incomplete Specimen |
A Stomatopoda Malacostracan. Its affinities haven't been tested. |
![]() A female Odontodactylus scyllarus mantis shrimp, maybe related to the Posidonia Specimen |
Thoracica[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
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|
Numerous disarticulated individuals, associated with fossil wood.[180] |
A phosphatic-shelled Cirripede of the family Eolepadidae.[180] Toarcolepas is provisionally interpreted as the oldest epiplanktonic cirripede known, and is thought to have lived attached to floating driftwood.[180] |
![]() Modern genus Lepas is among the most related taxa to |
Arachnida[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
Liassoscorpionides[181] |
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|
Single Incomplete Specimen. |
The type genus of the family , probably related to . Being the only Jurassic scorpion known, there is no evidence that L. schmidti was aquatic (which was suggested in the past) and in the absence of further, better preserved material it should be excluded from future considerations of broad patterns of scorpion evolution.[182] |
Insecta[]
Insects are a common terrestrial animals that were probably washed into the sea due to monsoon conditions present on the Sachrang Formation.[183]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
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Incomplete Specimens |
Uncertain Affinity Insects, whose remains are scarce to allow an assignation to a concrete family. Also Includes insects with characters never seen in other families and genera. |
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(Extinct clade of Basal Insects) of the family . |
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Isolated Wings |
An of the family . |
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Specimens |
An Odonatopteran (ancient winged insects) from the family . |
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Isolated Wings |
An Indeterminate Family dragonfly. Dragonflies non assiganted to a concrete family due to the incomplete or fragmentary nature of its remains. |
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A dragonfly of the family . Heterophlebia is a relative abundant Genus, present on most of the pits of the Sachrang Formation, even on Holzmaden. Heterothemis is the second most distributed insect on the formation. |
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Isolated Wings |
A dragonfly of the family . |
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A dragonfly of the family . The Kerkhofen specimens of Sphenophlebia pommerana shows an almost complete dragonfly with head and abdomen as well the two pairs of wings that are partially on top of each other, consisting of fore and hind wings.[194] |
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Isolated Wings |
A dragonfly of the family . |
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Isolated Wings |
A dragonfly of the family . Campterophlebia is the largest Early Jurassic Insect Know, with a wings size up to 20 cm.[195] |
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A dragonfly of the family . |
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A cockroach of the family . |
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Isolated Wings |
A cockroach of the family Mesoblattinidae. |
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Isolated Wings |
A cockroach of the family . |
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Isolated Wings |
A cockroach of the family Blattulidae. |
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Isolated Wings |
A stick insect of the family . Schesslitziella is one of the described insects found more near the Bohemian Massif, where probably belong most of the terrestrial invertebrate fauna. |
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grasshoppers of uncertain Placement. |
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A grasshopper of the family Protogryllidae. |
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A grasshopper of the family Elcanidae. |
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Elcana[30] |
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A grasshopper of the family Elcanidae. |
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A short-horned grasshopper of the family Acrididae. |
![]() Extant member of Acrididae, Acridiopsis was probably similar | |
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Grasshoppers of the family Locustopsidae. |
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Isolated Wings |
A Planthopper of uncertain placement. |
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Isolated Wings |
A Planthopper of the family Archijassidae. |
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Planthoppers of the family Fulgoridiidae. |
![]() | |
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Isolated Wings |
A Froghopper of the family Procercopidae. |
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A Shore bug (Saldidae) Of Uncertain Placement. |
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A Shore bug (Saldidae) of the family . |
![]() Extant member of Saldidae, genera where probably similar | |
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Beetles of uncertain placement inside Coleoptera. Diverse beetle specimens that due to its preservation (incomplete) or its morphological traits are not assigned to any concrete family. The abundance of beetle Elytrons indicates the proximity of terrestrial habitats. |
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Syntomopterus[184] |
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of the family Trachypachidae. |
![]() Extant member of Trachypachidae, Posidonia genera where probably similar | |
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A of the family Hydrophilidae. |
![]() Extant member of Hydrophilidae, Posidonia genera where probably similar | |
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Beetles of the family Coptoclavidae. |
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A Beetle of the family . |
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Elytrons |
A Giant Beetle. It is among the largest found on all the Jurassic. |
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Isolated Wings |
Mayfly of the family . |
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Isolated Wings |
Hanginflies of the family Bittacidae. |
![]() Extant member of Bittacidae, Posidonia genera where probably similar | |
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Isolated Wings |
Scorpionflies of the family Orthophlebiidae. |
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Protorthophlebia[184] |
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Isolated Wings |
Scorpionflies of the family Protorthophlebiidae. |
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Isolated Wings |
Scorpionflies of the family Pseudopolycentropodidae. |
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Flies Of Uncertain Placement. Some of the Specimens where listed as Nymphs but are now tougth to be wingless Dipterans |
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Isolated Wings |
A Fly of the family . |
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Isolated Wings |
A of the family Anisopodidae. |
![]() Extant member of Anisopodidae, Posidonia genera where probably similar | |
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Isolated Wings |
A Snakefly of the family . |
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Isolated Wings |
A Snakefly of the family . |
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Isolated Wings |
A Fly of the family . |
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Isolated Wings |
A Fly of the family . |
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Isolated Wings |
A Fly of the family . It represents the oldest know genus of this primitive family. |
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Isolated Wings |
A primitive Crane fly of the family Tanyderidae. Extant members of the family are nectar feeder while extinct members cannot be determined precisely.[206] |
![]() Extant member of Tanyderidae, Posidonia genera where probably similar | |
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Isolated Wings | |||
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Isolated Wings |
A Crane fly of the family Limoniidae. |
![]() Extant member of Limoniidae, Posidonia genera where probably similar | |
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Isolated Wings |
A Phantom Crane fly of the family . |
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Isolated Wings |
An Amphiesmenopteran of the family Necrotauliidae. The ovipositor, like terminalia of female N. parvulus, indicate that these insects laid their eggs rather in soil than in water |
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Isolated Wings |
Lepidopterans probably related with the family . Compared with their record on Grimmen, on Lower Saxony Lepidopterans are rather scarce and bad preserved. |
![]() Extant member of , Posidonia genera where probably similar | |
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Isolated Wings |
A Lacewing of uncertain placement. |
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Isolated Wings |
A Lacewing of the family . |
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Actinophlebia[211] |
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Isolated Wings |
A Silky Lacewing of the family Psychopsidae. |
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Isolated Wings |
A Lacewing of the family . A large genus with wings around 50 mm. |
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Isolated Wings |
A Giant Lacewing (Kalligrammatidae) of the subfamily and . It is one of the oldest known representatives of the Giant pollinator lacewings. The genus Liassopsychops was previously referred to Psychopsidae. Another specimen related, Specimen Hi 21 is regarded here as Kalligrammatidae incertae sedis. This along the occurrence of two distantly-related genera of Kalligrammatidae in the lower Toarcian is unexpected.[212] The Toarcian Kalligrammatidae lived in warm and dry conditions, as proven by its recovery from the zone of Hondelange but absence on Grimmen and other northern locations. Both genera where among the largest insects found on all the formation.[212] |
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Isolated Wings |
A lance Lacewing of the family . |
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Isolated Wings |
Lance Lacewings of the family Osmylidae. Tetanoptilon is the largest non-Kalligrammatidae lacewing of the Jurassic, with a forewings length up to 470 mm and a wingspan estimated at 11 cm. Like the Kalligrammatids, this genus is limited to the southern zones, asbsent in northern deposits.[212] |
![]() Extant member of Osmylidae, Posidonia genera where probably similar | |
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Isolated Wings |
A lance Lacewing of the family Mesochrysopidae. |
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Isolated Wings |
A of the family Tettigarctidae. |
Extant member of Tettigarctidae, Posidonia genera where probably similar | |
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Isolated Wings |
Pentatomomorphans of the family . Are related with the family Lygaeoidea, being possible ancestral forms of this last one. |
![]() Extant member of Lygaeoidea, Posidonia genera where probably similar | |
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Isolated Wings |
A Sternorrhynchan of the family . It is curious for its peculiar venation on its wings. Has been proposed its own family, . |
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Isolated Wings |
A Coleorrhynchan of the family Progonocimicidae. |
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Isolated Wings |
A Pseudo-Wasp of the family Sepulcidae. |
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Isolated Wings |
A Wasp of the family . |
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Isolated Wings |
A Wood Wasp of the family . |
Echinodermata[]
Echinoderm debris is pretty abundant on the shale-free Unken and Salzburg members, including Crinoid skeleton elements, also that of the Ophiurida; the Echinoids take their place, where really blossomed at that time. That's why Pedicellaria are observed very often.[20]
Asterozoa[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
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Isolated Arm Plates |
An Ophiuridan of the family . |
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Isolated Arm Plates |
An Ophiuridan of the family . |
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|
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Isolated Arm Plates |
An Ophiuridan of the family . |
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Ophiomusa[220] |
|
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Isolated Arm Plates |
An Ophiuridan of the family . |
|
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|
Isolated Arm Plates |
An Ophiuridan of the family . |
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|
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Isolated Arm Plates |
An Ophiuridan of the family . |
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Isolated Arm Plates |
An Ophiuridan of the family . |
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|
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Isolated Arm Plates |
An Ophiuridan of the family . |
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Isolated Arm Plates |
An Ophiuridan of the family . |
||
|
|
Isolated Arm Plates |
An Ophiuridan of the family Ophiohelidae. |
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Isolated Arm Plates |
An Ophiuridan of the family . |
||
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|
Isolated Arm Plates |
An Ophiuridan of incertade sedis family on the order . |
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|
|
Isolated Arm Plates |
An Ophiuridan of incertade sedis family on the order Ophionereididae. |
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|
Isolated Arm Plates |
An Ophiuridan of the family Ophiactidae. Very rare on the layers. |
![]() Modern Specimen | |
|
|
Isolated Arm Plates |
An Ophiuridan of the family Ophiolepididae. |
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An Ophiuridan of the family Ophiolepididae. |
![]() Holotype specimens from the Sachrang Formation | |
|
|
Isolated Arm Plates |
An Ophiuridan of the family Ophiacanthida. Very Common, related to non anoxic water sedimentation. |
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|
An Ophiuridan of the family . The Mostly know Arterozoan from the formation, including a single articulated specimen. |
![]() Fossil Specimen | |
|
|
Isolated Arm Plates |
An Ophiuridan of the family Ophiuridae. Its relationships haven't been confirmed and it is based on very fragmentary remains. |
||
|
|
Single specimen |
An Asteroidean of the family Astropectinidae. Only a relatively complete specimen is known from Banz Abbey. |
![]() Pentasteria sp. from Banz |
Echinoidea[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
Cidaris[225] |
|
|
Specimens |
A sea urchin of the family Cidaridae. Common on several layers. Cidaris is genus that still alive today. A bottom dweller, is commonly found associated with Belemnnite fossils, probably due to eating its carcasses. |
![]() Modern Specimen |
|
|
Specimens |
A sea urchin of the family Pedinidae. It is the most common sea urchin in the formation, present on all the levels with specimens of various sizes. Vinculated to sea bottom sediments, before Gasteropods and Bivalves, Diademospsis was the third major colonizer of the bottom, in between anoxic changes. |
Specimens from Holzmaden | |
|
|
Specimens |
A sea urchin of the family Pedinidae |
||
|
|
Specimens |
A sea urchin of the family Diadematidae |
Specimens from Holzmaden | |
|
|
Specimens |
A sea urchin of the family Pseudodiadematidae |
||
|
|
Specimens |
A sea urchin of the family |
Holothuroidea[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
|
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Specimens |
A sea cucumber of the family Chiridotidae. It is the only major genus of Sea Cucumbers reported locally on the Posidonienschiefer. It was identified originally as Chirodonta mesoliasicus and Chirodonta heptalmorpha. Represents a possible cosmopolitan holothurian that occupied all three types of deep-sea ecosystems as an organic-enrichment opportunist. |
||
|
|
Specimens |
A sea cucumber of the family inside Apodida. The "fishhooks" from this genus are only rarely found in the tenuicostatum zone |
Specimen | |
|
|
Specimens |
A sea cucumber of the family . Occurs sporadically in non-bituminous sediments of the upper bifrons zone |
||
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|
Specimens |
A sea cucumber of the family inside Dendrochirotida. A Genus whose identification was originally complex. |
Crinoidea[]
Genus | Species | Location | Material | Notes | Images |
---|---|---|---|---|---|
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Isolated Stems |
A Crinoidean of the family . |
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Isolated Stems |
A Crinoidean of the family Isocrinida. |
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Isolated Stems |
Type genus of Crinoidean of the family Isocrinida. |
![]() Close view of one specimen ![]() Single specimen | |
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The largest known Crinoidean, from the family . Among the tallest animals of its period, Seirocrinus is also one of the most famous fossils from the Sachrang Formation. It consists of fossils of colonies along large wood trunks, with specimens up to 14 m long, with the largest specimen reaching 26 m long,[230] what makes it among the tallest know Mesozoic organisms, one of the largest invertebrates know on the fossil record and one of the tallest know animals. It was an open ocean organism that lived in rafting woods, probably filtering food and serving as a refuge for other animals, such as ammonites.[231] The crinoids had a large colonization process, based on the status of the fossil wood found.[232][233] The large rafts were the home for a high variety of marine organisms, such as Balanoideans, Ammonites and other. It has been estimated that without the presence of modern raft wood predators (that appeared on the Bathonian) those rafts can last up to 5 years, being that the main reason the crinoids were able to reach such huge sizes. The large rafts were also probably essential to distribute animals along the Early Jurassic Seas.[234] |
![]() Close view of one specimen ![]() Single specimen | |
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Type genus of Crinoidean from the family . Like Seirocrinus, Pentacrinites formed colonies on rafting wood, getting a different role than bigger crinoid and appearing on the first stages of the decomposition of the rafting wood. Was a smaller genus, with specimens of no more than 1 meter long, usually measuring 40–70 cm. |
Close view of one specimen ![]() Reconstruction |
Vertebrata[]
Fishes[]
Chondrichthyes[]
Genus | Species | Location | Material | Notes | Images |
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Synechodus[237] |
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Teeth |
Thype shark of the family Synechodontiformes. |
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Articulated vertebral column, girdles, both fin spines and clasper organ |
A member of the family Palaeospinacidae. |
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Anterior part of body with basicranium, palatoquadrates, Meckel's cartilage, ceratohyals, epihyals, teeth, traces of the branchial arches and the anterior finspine |
Type member of the family Palaeospinacidae. |
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Meckelian Cartilages, Jaws, teeth, Palatoquadrates, placoid scales and dearticualted parts of the labial, hyoid and branchial skeleton. |
A shark of the family Hybodontidae. The Type specimen belongs to a large hybodontid, with an estimated total length of up to 3 m.[240] It has a Meckelian Cartilage more robust than Hybodus hauffianus and an acrodontine dentition, as seen on Bdellodus.[240] Probably is related with "Hybodus" delabechei, very likely a junior synonym of Crassodus.[240] |
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Teeth. |
A shark of the family Hybodontidae. An aberrant hybodontid with crushing dentition. |
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Type genus of the family Hybodontidae. It is the most abundant shark on the layers of the Sachrang Formation, with some of the best preserved specimens of the genus known. It was probably an open ocean hunter, with small horns over the eyes. With a size around 2 m, it was also one of the largest representatives of the Chondrichthyes on the formation.[244][245] |
![]() Hybodus Holzmaden specimen, among the best preserved of the genus, with Belemmnites inside. Hybodus Another Holzmaden specimen | |
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Type genus of the family . |
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[246] |
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A shark of the family Hexanchiformes. It was identified originally as a member of the genus Palaeospinax. |
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A member of Callorhynchidae inside Chimaeriformes. Similar to Callorhinchus, among the oldest known of its type. It the first Modern lineage Chimaeras from the Toarcian. |
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Acanthorhina[248] |
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Head and postcranial remains |
A member of inside Chimaeriformes. An aberrant Chimaera with an extrange elongated nose and horns over the skull. |
Acanthorhina |
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A member of inside Chimaeriformes. An aberrant Chimaera with a second jaw-like structure on its head. |
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Isolated Fin Spine |
A member of inside Chimaeriformes. |
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Batoidea[252] |
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SMNS 52666, Incomplete Specimen |
A possible member of Rajidae inside Batoidea. It was originally identified as a member of . This Genus was found to be sister taxa to the extant genus Raja. It is probably the oldest described pelagic Rajiform. It wears an enlogated rostrum. |
Actinopterygii[]
Genus | Species | Location | Material | Notes | Images |
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Complete Specimen |
First ganoin-scaled (Halecomorphi) from the Posidonienschiefer. The type specimen measures 51 cm, and has elongated and serrated body scales before the dorsal fin and tiny ganoid scales after it.[253] |
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Nearly complete specimen with broken skull |
First ganoin-scaled (Halecomorphi) from the Posidonienschiefer. Elongated morphology, with a length of ~39 cm, covered by smooth, massive ganoin scales.[253] |
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A member of the family inside |
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Various Complete and nearly complete Specimens |
Type genus of the family Caturidae inside Amiiformes |
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A possible representative of the family Saurichthyidae. Is based on rather fragmentary specimens. |
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The youngest representative of the family Saurichthyidae, known for its large jaws, similar to modern Belonidae. |
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Type genus of the family inside Ptycholepiformes. It is one of the youngest representatives of its family. |
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A member of the family . Is among the most abundant fishes on the late liassic of Europe, present on the sub-Mediterranean boreal, with specimens of several sizes. |
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A member of the family Leptolepididae. The most common member of its family, Leptolepis is commonly associated with crustaceans and small marine invertebrates, probably the main constituents of its diet. One on the most predated vertebrates on the formation, with abundance of larger fishes and reptiles with specimens associated. |
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MB. f.7612, nearly complete specimen. |
A member of the family Leptolepididae. Was identified as , but this name is currently occupied by a Japanese fish genus of Early Cretaceous age.[260] It differs from Leptolepis coryphaenoides in the presence of a few autapomorphies and also in the retention of several primitive features not present on the last one.[259] Small genus, of about 14 cm length.[259] |
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A member of the family Pachycormidae. |
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A member of the family Pachycormidae. Large representative of the family, reaching sizes up to 2.3 m. |
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Type member of the family Pachycormidae. Large representative of its family, with a size up to 1.5 m. One specimen preserved the alimentary canal, with the stomach filled by numerous hooklets that can be referred to the coleoid cephalopod Phragmoteuthis, implying a diet of cephalopods for this genus.[265] |
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A large member of the family Pachycormidae. |
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Ohmdenia[268] |
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Single desarticulated Specimen |
A large member of the family Pachycormidae, with a length of up to 2.5–3 m and an estimated weight over 200 kg.[268][269] Previously considered a junior synonym of Pachycormus, although the cranial bones suggest a different genus. Among the largest fish found on the formation, it is a key fossil in the transition to large filter feeding fishes.[269] Being coeval with another basal Pachycormiformes show the specialization of the group during the late lower jurassic.[269] Ohmdenia is the sister taxa to group of suspension-feeding Giant Middle-Late Jurassic Fishes (Including the famous Leedsichthys), showing alterations on its dental structure, with jaw indicates a diet based on soft body prey.[269] Its evolutionary significance is comparable to that of the genus Aetiocetus for the modern Baleen whale.[269] |
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A deep-bodied neopterygian, the type genus of the family Dapediidae. Unpublished material indicates the presence of one or even two more still undescribed species of Dapedium in the Lower Toarcian.[271] A specimen of Lytoceras has a Dapedium sp. in the body chamber probably trapped when feeding on the carcass of the cephalopod, and may therefore indicate an opportunistic carrion feeder.[272] |
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A deep-bodied neopterygian of the family Dapediidae. |
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Lepidotes[274] |
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A common member of the Lepisosteiformes. |
![]() Lepidotes ![]() Lepidotes |
Strongylosteus[275][276] |
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A large member of the Chondrosteidae and the largest non-reptilian marine vertebrate of the Sachrang Formation, with a size between 3 and 4.5 m, and an estimated weight over 800 kg to 1 tonne.[275] Has been suggested as a junior synonym of Chrondrosteus, although there haven't been any new revision about the status of the genus. It is related to modern sturgeons, but with a different kind of mouth than common species, made for hunting prey in open waters, with a strong lower jaw, similar to modern beluga sturgeons.[277] |
Sarcopterygii[]
Genus | Species | Location | Material | Notes | Images |
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A large coelacanth of the family Mawsoniidae, related to the genera Axelrodichthys, Chinlea, Diplurus and the type, Mawsonia.[279] The largest specimen known from the Sachrang Formation is GPIT.OS.770 (Holotype), with a length over 1.6 m.[278] The specimen presents an ossified lung inside the abdominal cavity, and most of the body, being also one of the most complete coelacanths of the Jurassic found.[278][279] Some recent discoveries from the Middle Jurassic show specimens of up to 3.5 m long.[280] Trachymetopon precedes the presence of the family Mawsoniidae in Europe by about 120 Ma and the northernmost occurrence of a member of the group, implying an extensive geographical range during the Early Jurassic.[279] Due to the specimens being found on pelagic deposits suggest that probably was an open ocean swimmer.[278][279] |
Trachymetopon GPIT.OS.770 |
Amniota[]
Ichthyosauria[]
Inderminate specimens are known.[7]
Genus | Species | Location | Material | Notes | Images |
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Type genus of the family Temnodontosauridae. A large macroraptorial ichthyosaur, apex predator of its environment. It ranges between 9 and the 12 m, being one of the largest known ichthyosaurs, characterized by skulls and jaws over 1 m in length, with the largest being over 1.9 m long. It has been found with fragments of young icthyosaur in his stomach.[283] Of 39 specimens of Temnodontosaurus studied, a 21% (8 Specimens) show pathologies along their body, with several injuries post traumatic, probably done by other marine reptiles.[284] |
![]() Temnodontosaurus hunting Temnodontosaurus trigonodon specimen. | |
Leptopterygius?[285] |
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A possible member of the family Leptonectidae. Mostly of the specimens of this genus have been referred to Leptonectes or Temnodontosaurus, although some remains on the Posidonienschiefer are too complex to being clearly referred. |
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Type genus of the family Stenopterygiidae. A common Toarcian ichthyosaur, present in multiple layers. The rather exquisite level of preservation has led to know even the coloration, that exposes a clear countershading, with an upper part being more obscure than the lower, similar to modern killer whales, the Heaviside's dolphin or the Dall's porpoise. There is also evidence of changes in color with ontogenic changes, going from dark juveniles to countershaded adults. The skin was flexible & scaleless, as in dolphins.[290] The study of several specimens has revelated that Stenopterygius quadriscissus underwent a size-related trophic niche shift through ontogeny, shifting from a piscivorous diet to a teuthophagous diet, known thanks to exquisitely preserved stomach contents.[291] Stenopterygius was a relatively opportunistic feeder, as gut contents tend to track relative species abundance.[291] |
![]() Restoration ![]() Fossil | ||
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Type genus of the family Suevoleviathanidae. Includes specimens up to 4 m long. |
Suevoleviathan integer fossil | |
Hauffiopteryx[295] |
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Complete Specimens |
Small sized ichthyosaur, probably a member of Parvipelvia, sister group to Stenopterygius + Ophthalmosauridae.[297] A small- to mid-sized ichthyosaur, 2–3 m in length, with a relatively short and slender antorbital rostrum.[296] |
Hauffiopteryx typicus fossil |
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A large ichthyosaur of the family Leptonectidae with convergent evolution with modern swordfish. Like these fishes, Eurhinosaurus is believed to be a fast swimming predator, able to hunt fish schools on same way. Large specimens of up to 6 m are known. |
Complete specimen from the Sachrang Formation ![]() Eurhinosaurus restoration |
Plesiosauria[]
Genus | Species | Location | Material | Notes | Images |
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An indeterminate plesiosauroid plesiosaur. An impressively preserved immature specimen, different from Hydrorion brachypterygius and Seeleyosaurus guilelmiimperatoris, the most abundant plesiosaurs found locally, and its anatomical characters suggest it represents a new genus.[302] It fossilized buff-coloured material, identified as mainly composed of calcium phosphate and interpreted as phosphatised muscle tissues.[302] It also contains eumelanin and hence possibly corresponds to areas dark-coloured in life.[302] The stomach contains quartz grains that were most likely ingested during the animal's life, maybe used for food trituration. This gastroliths have importance for its exotic provenance compared with the surrounding lithology of the deposits (mostly shale), and as sandy turbidites have never been reported from the SW German Basin, the individual may have acquired the sand-sized grains many kilometres away from the burial site.[302] Strata containing fine sand ('Glaukonit und viel Feinsand') at (western side of the Black Forest between Freiburg and Basel), 200 km from Holzmaden suggest a nearshore deposit was allocated here, and that the black forest emerged at this time. Probably this young specimen reached that location in search for gastroliths.[302] |
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Complete Specimen. |
A basal Plesiosaur that has been linked with Cryptoclididae. It is one of the smallest from the Posidonia, with a complete skeleton measuring less than 2.5 m. It is considered a possible junior synonym of Seeleyosaurus.[304] |
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A Plesiosaur of the family Microcleididae. It was named originally "Plesiosaurus guilelmiimperatoris". It was a moderate‐sized plesiosauroid, measuring up to 3,5 m in length with a skull length of 170 mm.[306] |
![]() Seeleysaurus | |
Microcleidus[307] |
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Type member of the Plesiosaur family Microcleididae. Small Plesiosaur, with a length of less than three metres. It is characterised by a really elongated neck, and was probably an ichthyophagous form that occurred rarely in the Posidonienschiefer fauna. |
![]() Microcleidus |
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A junior synonym of M. brachypterygius.[302] |
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Nearly complete specimen. |
A Rhomaleosauridae Plesiosaur. Its detailed fossils have helped to study plesiosaur movement.[312] |
![]() Meyerasaurus | |
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Complete specimen. |
A basal member of Pliosauridae. A moderately sized (3.4 m) Pliosaurid, ecologically adapted to fish hunt, as has been observed due to comparing the long snouted skull with that of Peloneustes, Gharial Crocodiles or Dolphins.[313] |
![]() Hauffiosaurus |
Sphenodontia[]
Genus | Species | Location | Material | Notes | Images |
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An aquatic sphenodont of the family Pleurosauridae. Palaeopleurosaurus evidences that there was a slightly skeletal specialization for an aquatic lifestyle, achieved through the Jurassic gradually on pleurosaurs.[314] It has similarities with other marine reptiles, such with members of Sauropterygia the presence of a defined suture between the centrum and the neural arch, along with reduced sternum.[315] Probably had a semiaquatic style of life, although not as adapted as Pleurosaurus, as show limited morphological evidence of adaptation to a complete aquatic lifestyle, defined by no Osteosclerosis and the lack of Pachyostosis, except for a thicker shaft region in the humerus, that is as narrow as in terrestrial rhynchocephalians, such as the terrestrial Clevosaurus.[315] Palaeopleurosaurus probably was still able to walk on land, for example for Oviposition.[315] Recent studies suggest a shorter lifespan than modern Tuatara, based on irregular spacing of growth marks.[316] |
Testudinata[]
Genus | Species | Location | Material | Notes | Images |
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A marine turtle of the superfamily Eurysternidae inside Thalassochelydia. Is the main formally identified fossil as Turtle from the Sachrang Formation, representing a rather basal genus. The pleurals resemble those of the genus Plesiochelys.[317] Being found in the zone of Franconia that on the Toarcian was at -80 km from the shore can suggest that early marine turtles lived in the epicontinental waters of the European shallow seas before reach richer ecosystem diversity on the Late Jurassic.[317] This would explain the serious lack of Turtle fossils on the formation, as mostly of the deposits are located far from the coast.[317] |
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Thalassochelydia[318] |
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A marine turtle of the clade Thalassochelydia inside Angolachelonia. The only testudine identified from the northern realm of the Sachrang Formation.[318] |
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Possible unclassified testudine remains. Münster (1834) cited: "there were also rare things at the quarries of Altdorf, among other remains there were ones of a turtle on lias limestone"". The remains are not catalogued and some specimens are in Private Collections.[323] |
Crocodylomorpha[]
Genus | Species | Location | Material | Notes | Images |
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Plagiophthalmosuchus[324] |
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A Longirostrine Thalattosuchian, the most basal know. Was considered synonymous with Steneosaurus. Longirostrine specialist, probably active fish hunter. |
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A longirostrine thalattosuchian of the family Machimosauridae. Was considered synonymous with Steneosaurus until in 2020 this last was recovered as invalid.[327] It reached large sizes, with specimens exceeding 5 m, being a generalist predator.[324] |
Macrospondylus bollensis fossil with Skin Impressions | |
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A mesorostrine thalattosuchian of the family Teleosauridae. A marine crocodylomorph with a diet probably based on fish. Was considered synonymous with Steneosaurus until recently.[331] Due to this unusual placement of the external nares, Mystriosaurus was more terrestrial, or spent a greater amount of time on land, than other teleosauroids. This would explain its greater presence in zones of the formation more proximal to the emerged landmasses. Its morphology suggest it was a mesorostrine generalist.[331] |
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A longirostrine thalattosuchian of the family Teleosauridae. Platysuchus was slightly more robust than its contemporaneous relatives, being probably adapted to hunt more voluminous fish. A heavily armoured, semi-terrestrial longirostrine generalist form, indicated by the extensive and tightly packed rows of dorsal osteoderms.[324] |
![]() Platysuchus | |
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A thalattosuchian with a complex assignation, probably the basalmost metriorhynchoid. Pelagosaurus typus was a small-bodied thalattosuchian (∼1 m in length) considered to be an adept aquatic pursuit predator, with a long streamlined snout ideal for snapping at fast moving prey (one specimen was found with Leptolepis fishes inside) and large, anterolaterally placed orbits for increased visual acuity.[334] |
Pterosauria[]
Genus | Species | Location | Material | Notes | Images |
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A Novialoidean Pterosaur, type genus of the family Campylognathoidea. Mark Witton suggests the construction of Campylognathoides' extremely robust forelimbs, with proportionally long wing fingers, could be a specialization for a fast aerial lifestyle comparable to those of Falcons and mastiff bats, being more probably an insect & vertebrate hunter and living on nearshore environments.[338] |
Nearly complete Campylognathoides | ||
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Pelvis and several vertebrae. |
A Novialoidean Pterosaur, probably a member of the family Campylognathoidea. Has been assigned to the genus Campylognathoides, although it is clearly different than any other pterosaur from the Sachrang Formation.[39] The name "Schandelopterus" is invalid and lacks any study, assigned without species to refer to the specimen on private German Fossil Groups. The pelvis indicates a laterally, slightly upwardly directed orientation of the acetabula which does not support a bird-like bipedal locomotion of this pterosaur as has been suggested on the past.[39] |
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Parapsicephalus[339] |
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Skull |
A Rhamphorhynchinae Pterosaur. Has been assigned to the genus Dorygnathus. It has a really complete skull that can help to explain the status of the genus Parapsicephalus.[339] |
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A Rhamphorhynchinae Pterosaur. It is one of the best known Early Jurassic Pterosaurs.[340] Unlike Campylognthoides, Dorygnathus was an oceanic hunter, with teeth disposed to catch marine prey, such as Belemnittes and several species of fishes. Dorygnathus mistelgauensis is considered a junior synonym until more data can be recovered from the specimen, held on a private collection.[340] |
![]() Dorygnathus on terrestrial pose ![]() Nearly complete Dorygnathus | ||
"Ohmdenodraco"[340] |
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Femur and a broken tibia-fibula. |
A possible Rhamphorhynchinae Pterosaur. Like "Schandelopterus", "Ohmdenodraco" is an invalid name, used on private fossil groups to refer to SMNS 80439, assigned as "?Dorygnathus sp." originally, although characters on the tibia and femur are distinct to any Pterosaur found on the Posidonia.[340] |
Saurischia[]
Possible teeth from Dinosaurs are known from the Lias Clay pit of Unterstürmig (Referred as "Various Archosaur teeth").[7] "Dinosaur Fossils" are cited to be present on several levels on Schandelah.[39]
Genus | Species | Location | Material | Notes | Images |
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An indeterminate possible Theropod dinosaur, possibly Neotheropoda. Hasn't been revised since 1984. The Cervical Vertebrae was found on Schandelah and was described as having a similar appearance and size to those found on the Triassic genus Pterospondylus (What would make it a late Coelophysidae member, with a length of 1.8 m).[348] But can be alternatively from a Plesiosaur.[348] A series of teeth found on Altdoft can include theropod teeths, probably from smaller specimens (speculated less than 60 cm animals).[7] |
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Sauropodiformes[349] |
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3 cm long, tooth-studded fragment of a lower jaw |
An indeterminate possible Sauropodomorph dinosaur, possibly a member of Sauropodiformes inside Anchisauria (Resemble Yunnanosaurus teeth).[349] The Fossil was reported from the Lias Epsilon level, that on Oedhof is occupied by the Sachrang Formation.[349] It was found with abundant Plant debris and Belemnite remains.[349] Hasn't been revised since 1956 |
![]() The Oedhof jaw can belong to a creature similar to Yunnanosaurus |
Ohmdenosaurus[350] |
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Tibia and astragalus |
A Gravisaurian Sauropod. One of the few formally described from the Toarcian. Was confused as a Plesiosaur bone.[350] |
![]() Ohmdenosaurus |
Synapsida[]
Genus | Species | Location | Material | Notes | Images |
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Possible Cynodont Remains, Incertade sedis inside Cynodontia. The specimens were listed on several notes on the 1800s (As "mammal teeth"), although its existence has not been proved. Can be related to the Trithelodontidae or other late surviving Cynodonts by related characters, such as Irajatherium, but also to the Mammaliaformes. If its presence is proved, it would be the first Synapsid found on the Sachrang Formation. |
![]() Morganucodon, example of Lower Jurassic Synapsid of Europe. The supposed synapsid teeth of the Sachrang Formation (in case of be proven to exist) maybe where related to this genus |
References[]
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- ^ a b Martill, D. M. (1993). Soupy substrates: a medium for the exceptional preservation of ichthyosaurs of the Posidonia Shale (Lower Jurassic) of Germany. Kaupia, 2, 77-97.
- ^ Schmid–Röhl, A., & Röhl, H. J. (2003). Overgrowth on ammonite conchs: environmental implications for the Lower Toarcian Posidonia Shale. Palaeontology, 46(2), 339-352.
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- ^ a b c d e f g h i j k l m n o Prauss, M., Ligouis, B., & Luterbacher, H. (1991). Organic matter and palynomorphs in the 'Posidonienschiefer'(Toarcian, Lower Jurassic) of southern Germany. Geological Society, London, Special Publications, 58(1), 335-351.
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- ^ a b Wille, W. et al. (1979) Dinoflagellates from the Lias of Sw Germany. [ Dinoflagellaten aus dem Lias Sмdwestdeutschlands. ] Neues Jahrbuch fмr Geologie und Paleontologie. Abhandlungen Vol. 158 # 2 P. 221- 258
- ^ a b c d e f g h i Wille, W. (1982) Evolution and ecology of Upper Liassic dinoflagellates from southwest Germany. Neues Jahrbuch fмr Geologie und Paleontologie. Abhandlungen Vol. 164 # 1 P. 74- 82
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- ^ Feist‐Burkhardt, S. (1995). Weiachia fenestrata gen. et sp. nov., a new Phallocystean dinoflagellate cyst from the Lower Jurassic of Switzerland. Palynology, 19(1), 211-219.
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External links[]
- Posidonia Shale
- Jurassic Germany
- Fossils of Germany
- Prehistoric fauna by locality