Tritemnodon

Tritemnodon; Temporal range: 50.5–46.2 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ Early Eocene
	Restoration of Tritemnodon agilis
	Tritemnodon agilis skeleton
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Hyaenodonta
Genus:	†Tritemnodon; Matthew, 1906
Type species
	†Tritemnodon agilis; Marsh, 1872
Synonyms
	synonyms of speacies: T. agilis: Limnocyon agilis (Marsh, 1872) ; Sinopa agilis (Matthew, 1901) ; Sinopa brevicalcaratus (Cope, 1872) ; Sinopa gracilis (Wortman, 1902) ; Stypolophus agilis ; Stypolophus brevicolcarabus (Cope, 1872) ; Stypolophus brevicalcaratus (Cope, 1872) ; ;

Tritemnodon ("three cutting teeth") was a genus of hyaeanodontid which lived 50.5-46.2 million years ago.^[6]^[7] Fossils of Tritemnodon agilis have been found in Utah and Wyoming (Willwood Formation of Big Horn County and the Lower Bridger Formation of Uinta County), and it was the size of a wolf.

Phylogeny[]

The phylogenetic relationships of genus Tritemnodon are shown in the following cladogram.^[8]^[9]^[10]^[11]

†Hyaenodonta

†

	†

	†Hyaenodonta sp. (Quarry L-41, Fayum, Egypt)

	†Lahimia

	†

	†

†

†

	†

	†Pyrocyon

†Galecyon

	†

	†

†

†Tritemnodon

	†Tritemnodon agilis

	†Tritemnodon sp. (FMNH PM 55839)

†

†

†

†

†Teratodontidae

†

†

†Orienspterodon

†

	†Hemipsalodon

?	†

†Akhnatenavus clade

†

	†

	†

†

†
	†Pterodontina

	†

†Falcatodon

†Sectisodon

	†Exiguodon

	†Isohyaenodon zadoki

†Isohyaenodon
(†Isohyaenodontina)

†Isohyaenodon andrewsi

	†Sivapterodon

	†Hyainailouros bugtiensis

†Hyainailouros napakensis

†Hyainailouros

	†Hyainailouros sulzeri

	†Hyainailourinae sp. (Arrdrift)

†Simbakubwa

	†

	†Metapterodontini

	†Megistotherium

	†

†

†

Hyainailouridae

†Lahimia clade

†Arfia clade

†Galecyon clade

†Indohyaenodon clade

†Tritemnodon clade

References[]

^ Matthew, W. D. (1906.) "The Osteology of Sinopa, a Creodont Mammal of the Middle Eocene." Proc. U. S. Nat. Mus., Vol. XXX, pp. 203-233, pl. XVI
^ O. C. Marsh (1872.) "Preliminary description of new Tertiary mammals. Part II." American Journal of Science 4(21):202-224
^ Matthew, W. D. (1901.) "Additional observations on the Creodonta." Bulletin of the American Museum of Natural History, 14: 1-38
^ E. D. Cope (1872.) "Third account of new Vertebrata from the Bridger Eocene of Wyoming Valley." Proceedings of the American Philosophical Society (separate) 1-4
^ J. L. Wortman (1902.) "Studies of Eocene Mammalia in the Marsh Collection, Peabody Museum." The American Journal of Science, series 4 13:433-448
^ Gingerich, Philip D., and Harvey A. Deutsch (1989). "Systematics and evolution of early Eocene Hyaenodontidae (Mammalia, Creodonta) in the Clarks Fork Basin, Wyoming". Museum of Paleontology, University of Michigan. 27 (13): 327–391.CS1 maint: uses authors parameter (link)
^ Naoko Eg (2004.) "A new genus and species of hyaenodontid creodont from the Pondaung Formation (Eocene, Myanmar)" Journal of Vertebrate Paleontology 24(2): 502–506 doi:10.1671/2481
^ Borths, Matthew R; Stevens, Nancy J (2017). "Deciduous dentition and dental eruption of Hyainailouroidea (Hyaenodonta, "Creodonta," Placentalia, Mammalia)". Palaeontologia Electronica. 20 (3): 55A. doi:10.26879/776.
^ Matthew R. Borths; Nancy J. Stevens (2019). "Simbakubwa kutokaafrika, gen. et sp. nov. (Hyainailourinae, Hyaenodonta, 'Creodonta,' Mammalia), a gigantic carnivore from the earliest Miocene of Kenya". Journal of Vertebrate Paleontology. 39 (1): e1570222. doi:10.1080/02724634.2019.1570222. S2CID 145972918.
^ Floréal Solé; Bernard Marandat; Fabrice Lihoreau (2020). "The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian". Geodiversitas. 42 (13): 185–214. doi:10.5252/geodiversitas2020v42a13.
^ Solé, F.; Morlo, M.; Schaal, T.; Lehmann, T. (2021). "New hyaenodonts (Mammalia) from the late Ypresian locality of Prémontré (France) support a radiation of the hyaenodonts in Europe already at the end of the early Eocene". Geobios. 66–67: 119–141. doi:10.1016/j.geobios.2021.02.004. S2CID 234848856.

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[1] Matthew, W. D. (1906.) "The Osteology of Sinopa, a Creodont Mammal of the Middle Eocene." Proc. U. S. Nat. Mus., Vol. XXX, pp. 203-233, pl. XVI

[2] O. C. Marsh (1872.) "Preliminary description of new Tertiary mammals. Part II." American Journal of Science 4(21):202-224

[3] Matthew, W. D. (1901.) "Additional observations on the Creodonta." Bulletin of the American Museum of Natural History, 14: 1-38

[4] E. D. Cope (1872.) "Third account of new Vertebrata from the Bridger Eocene of Wyoming Valley." Proceedings of the American Philosophical Society (separate) 1-4

[5] J. L. Wortman (1902.) "Studies of Eocene Mammalia in the Marsh Collection, Peabody Museum." The American Journal of Science, series 4 13:433-448

[6] Gingerich, Philip D., and Harvey A. Deutsch (1989). "Systematics and evolution of early Eocene Hyaenodontidae (Mammalia, Creodonta) in the Clarks Fork Basin, Wyoming". Museum of Paleontology, University of Michigan. 27 (13): 327–391.CS1 maint: uses authors parameter (link)

[7] Naoko Eg (2004.) "A new genus and species of hyaenodontid creodont from the Pondaung Formation (Eocene, Myanmar)" Journal of Vertebrate Paleontology 24(2): 502–506 doi:10.1671/2481

[Matthew2017-8] Borths, Matthew R; Stevens, Nancy J (2017). "Deciduous dentition and dental eruption of Hyainailouroidea (Hyaenodonta, "Creodonta," Placentalia, Mammalia)". Palaeontologia Electronica. 20 (3): 55A. doi:10.26879/776.

[Simbakubwa-9] Matthew R. Borths; Nancy J. Stevens (2019). "Simbakubwa kutokaafrika, gen. et sp. nov. (Hyainailourinae, Hyaenodonta, 'Creodonta,' Mammalia), a gigantic carnivore from the earliest Miocene of Kenya". Journal of Vertebrate Paleontology. 39 (1): e1570222. doi:10.1080/02724634.2019.1570222. S2CID 145972918.

[Geodiversitas4213-10] Floréal Solé; Bernard Marandat; Fabrice Lihoreau (2020). "The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian". Geodiversitas. 42 (13): 185–214. doi:10.5252/geodiversitas2020v42a13.

[hyaenodontsGeobios-11] Solé, F.; Morlo, M.; Schaal, T.; Lehmann, T. (2021). "New hyaenodonts (Mammalia) from the late Ypresian locality of Prémontré (France) support a radiation of the hyaenodonts in Europe already at the end of the early Eocene". Geobios. 66–67: 119–141. doi:10.1016/j.geobios.2021.02.004. S2CID 234848856.

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Tritemnodon Temporal range: 50.5–46.2 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ Early Eocene

Restoration of Tritemnodon agilis

Tritemnodon agilis skeleton
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Hyaenodonta
Genus:	†Tritemnodon Matthew, 1906^[1]
Type species
†Tritemnodon agilis Marsh, 1872^[2]
Synonyms
synonyms of speacies: T. agilis: Limnocyon agilis (Marsh, 1872) Sinopa agilis (Matthew, 1901)^[3] Sinopa brevicalcaratus (Cope, 1872)^[4] Sinopa gracilis (Wortman, 1902)^[5] Stypolophus agilis Stypolophus brevicolcarabus (Cope, 1872) Stypolophus brevicalcaratus (Cope, 1872)